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  No. of taxa found: 3 142, specimens: 56 337.  |  Filter your search by clicking selected family or country.  |  Help page  |  Go to list of taxa by order.
OrderFamilySpeciesCountryIndividualsImage  County/Province | Collection notes
Coleoptera   Scarabaeidae  Ablabera flavoclypeata Wallengren,1881   South Africa 4Melolonthinae Afr.3 (4) types?
Ablabera pilosula Boheman,   South Africa 3Melolonthinae Afr.3 (3)
Ablabera pulicaria Fåhraeus,1857   South Africa 1Melolonthinae Afr.3 (1)
Ablabera spp.    South Africa 1Melolonthinae Afr.3 (1)
Acanthobodilus immundus Creutzer,1799   Afghanistan 3Aphodiinae Pal.8 (3)
Bulgaria 13Aphodiinae (9) Pal.5+10
Pal.29 (4)
Germany 1Aphodiinae (1) Pal.16
Greece 28Aphodiinae Pal.5 (28)
Hungary 5Aphodiinae (5) Pal.15+16
Italy 1Aphodiinae Pal.5 (1)
Romania 1Aphodiinae Pal.16 (1)
Russian Federation 1Aphodiinae Pal.22 (1)
Sweden 3  Öland
Aphodiinae Main (3)
Turkey 4 Alan Dufberg 48 (1)
Aphodiinae Pal.28 (1)
Aphodiinae Pal.30 (2)
Caucasus, Ryssland / Georgien / Azerbajdzjan / Armenien / Iran / Turkiet 1Aphodiinae Pal.22 (1)
Unknown 1Aphodiinae (1) Pal.16 (from Tala-Thal
Semirjeschensk)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 5Aphodiinae Pal.5 (5)
Uzbekistan 1Aphodiinae Pal.16 (1)
Acanthobodilus languidulus Schmidt,1916   Morocco 1Aphodiinae Pal.17 (1)
Acanthovalgus spp.    America borealis, United States/Canada 1Cetoniinae N.Am.4 (1)
Achranoxia pelzi (Semenov & Medvedev,1936)   Turkmenistan 1Melolonthinae Pal.1 (1) as ssp. of A. expallida
Acrossus atratus (Waterhouse,1875)   Spain 1Aphodiinae Pal.17 (1)
Acrossus bimaculatus (Laxman,1778)   Unknown 9Aphodiinae Pal.19 (9)
Acrossus carpetanus (Graëlls,1847)   Spain 2Aphodiinae Pal.19 (2)
Unknown 1Aphodiinae Pal.19 (1)
Acrossus depressus (Kugelann,1792)   Norway 11Aphodiinae Main (11)
Sweden 302  Sk Bl Ha Sm Öl Go Ög Vg Sö Up Vr Dr Gä Hs Hr Jä Ån Vb Nb Ly Pi Lu To
Alan Dufberg 47 (16)
Aphodiinae Main (286)
Switzerland 2Aphodiinae Pal.4 (1)
Pal.28 (1)
Acrossus histrio (Balthasar,1932)   China 3Aphodiinae (3) Pal.17+19 (one paratype)
Acrossus klickai (Balthasar,1932)   China 1Aphodiinae (1) Pal.17
Acrossus laticollis (Baudi,1870)   Italy 4Aphodiinae Pal.11 (4)
Acrossus luridus (Fabricius,1775)   Sweden 124  Skåne Halland Småland Öland
Alan Dufberg 47 (17)
Aphodiinae Main (107)
Turkey 7Aphodiinae (4) Pal.5+19
Pal.25 (1)
Pal.28 (2)
Acrossus rubripennis (Horn,1870)   Canada 2Ontario
Aphodiinae N.Am.1 (2)
Canada 1Quebec
Aphodiinae N.Am.1 (1)
United States 1Virginia
Aphodiinae N.Am.1 (1)
United States 3West Virginia
Aphodiinae N.Am.1 (3)
Acrossus rufipes (Linnaeus,1758)   Denmark 4Aphodiinae Main (4)
Sweden 785  Sk Bl Ha Sm Öl Go GS Ög Vg Bo Ds Nä Sö Up Vs Vr Dr Gä Hs Hr Jä Ån Vb Nb Ås Ly Pi Lu
Alan Dufberg 47 (12)
Aphodiinae Main (773)
Acrossus semiopacus (Reitter,1887)   China 1Aphodiinae Pal.28 (1)
Acrossus siculus buturensis (Dellacasa,1983)   Italy 1Aphodiinae Pal.18 (1)
Acrossus siculus siculus (Harold,1862)   Italy 1Aphodiinae Pal.18 (1)
Actinobolus radians Westwood,1841   Unknown 1Dynastinae (1) S.Am.21
Adebrattia depressa Bordat & Howden,1995   Malaysia 1Borneo
Type Coleoptera 3036 (1) Holotype 03041:1 of Adebrattia depressa Bordat & Howden 1995
  Adebrattia depressa Bordat & Howden 1995

Adebrattia spp.    Malaysia 1Borneo
Aphodiinae Or.6 (1)
Adoretosoma atritarse Fairmaire,1891   India 1Rutelinae Or.7 (1)
Adoretosoma chromaticum (Fairmaire,1886)   Laos 1Rutelinae Or.7 (1)
Adoretosoma spp.    China 1Rutelinae Pal.8 (1)
Adoretus afghanus Machatschke,1958   Afghanistan 3Rutelinae Pal.3 (3)
Adoretus andrewesi Ohaus,1914   India 1Rutelinae Or.8 (1)
Adoretus bicaudatus Arrow,1914   Sri Lanka 1Rutelinae Or.5 (1)
Adoretus clypeatus Burmeister,1844   Egypt 1Rutelinae Pal.8 (1)
Adoretus compressus (Weber,1801)   Indonesia 1Java
Rutelinae Or.8 (1)
Thailand 8Rutelinae Or.5 (8)
Adoretus cribratus White,1844   India 1Rutelinae Or.8 (1)
Adoretus cribrosus Harold,1869   South Africa 4Rutelinae Afr.1 (4)
Adoretus cupreus Arrow,1899   South Africa 3Rutelinae Afr.1 (3)
Adoretus feminalis Arrow,1914   Sri Lanka 22Rutelinae Or.5 (22)
Adoretus flaveolus Fåhraeus,1857   South Africa 2Rutelinae Afr.1 (2)
Adoretus fraudulentus Peringuey,1902   South Africa 1Rutelinae Afr.1 (1)
Adoretus gazella Frey,1971   Sudan 1Rutelinae Afr.2 (1) Paratype
Adoretus hanströmi Machatschke,1970   South Africa 6Rutelinae (6) Types 853:1-6 Holotype Paratype 00853:1-6 of Adoretus hanströmi Machatschke 1970
  Adoretus hanströmi Machatschke 1970

Adoretus ictericus Burmeister,1844   South Africa 3Rutelinae Afr.2 (3)
Adoretus imitator Benderitter,1927   Namibia 2Rutelinae Afr.1 (2)
Adoretus iranicus Petrovitz,1958   Afghanistan 1Rutelinae Pal.3 (1)
Adoretus ladakanus nuristanicus Balthasar,1936   Afghanistan 1Rutelinae Pal.3 (1)
Adoretus nigrifrons (Steven,1809)   Afghanistan 3Rutelinae Pal.3 (3)
Russian Federation 4Rutelinae Pal.8 (4)
Adoretus picticollis Fåhraeus,1857   South Africa 2Rutelinae Afr.2 (2)
Adoretus pruinosus Ballion,1871   Afghanistan 1Rutelinae Pal.3 (1)
Adoretus spp.    South Africa 2Rutelinae Afr.2 (2)
Adoretus suturalis Arrow,1914   Sri Lanka 7Rutelinae Or.5 (7)
Adoretus tessulatus Burmeister,1855   South Africa 4Rutelinae Afr.2 (4)
Adoryphorus couloni (Burmeister,1847)   Australia 1Dynastinae Pac.2 (1)
Unknown 2Dynastinae Pac.2 (2)
Aegialia arenaria (Fabricius,1787)   Denmark 48Aphodiinae Main (48)
Germany 6Aphodiinae (6) Pal.13+16
Ireland 3Aphodiinae (3) Pal.16
Netherlands 1Aphodiinae (1) Pal.16
Sweden 279  Skåne Blekinge Halland Öland Gotland Gotska Sandön Västergötland
Alan Dufberg 47 (16)
Aphodiinae Main (263)
United Kingdom 3England
Aphodiinae Pal.13 (3)
Aegialia blanchardi Horn,1887   Canada 5British Columbia
Aphodiinae N.Am.4 (3)
N.Am.7 (2)
Canada 1Yukon
Aphodiinae N.Am.7 (1)
United States 6Washington
Aphodiinae N.Am.3 (6)
Aegialia comis (Lewis,1895)   Japan 2Aphodiinae (2) Pal.16
Aegialia conferta Horn,1871   Canada 1Ontario
Aphodiinae N.Am.3 (1)
United States 1Arizona
Aphodiinae N.Am.3 (1)
United States 1Washington
Aphodiinae (1) N.Am.3 (as A.nigrella Brown)

United States 2California
Aphodiinae N.Am.3 (2)
Aegialia convexa Fall,1932   United States 3California
Aphodiinae N.Am.3 (3)
Aegialia crassa LeConte,1857   Canada 1British Columbia
Aphodiinae (1) N.Am.4
Aegialia cylindrica (Eschscholtz,1822) (Psammoporus)   United States 8Alaska
Aphodiinae N.Am.3 (8)
Aegialia latispina LeConte,1878   United States 1Washington
Aphodiinae N.Am.3 (1)
United States 5Arizona
Aphodiinae N.Am.3 (5)
Aegialia nitida Waterhouse,1975   Japan 1Aphodiinae (1) Pal.13
Aegialia opifex Horn,1887   Canada 6Ontario
Aphodiinae N.Am.3 (6)
Aegialia rufa (Fabricius,1792)   Denmark 3Aphodiinae Main (3)
Germany 1Aphodiinae Pal.13 (1)
Sweden 1 Skåne
Aphodiinae Main (1)
Aegidium spp.    Unknown 1Orphninae Afr.1 (1)
Aegopsis curvicornis Burmeister,1847   Unknown 2Dynastinae S.Am.17 (2)
Aethiessa floralis (Fabricius,1787)   Algeria 3Cetoniinae Pal.17 (3)
Italy 2Cetoniinae Pal.17 (2)
Tunisia 6Cetoniinae (6) Pal.9+17+25
Aethiessa mesopotamica Burmeister,1842   Afghanistan 1Cetoniinae (1) Pal.10 (as A. bagdadensis Burmeister)

Aethiessa squamosa (Gory & Percheron,1833)   Algeria 1Cetoniinae Pal.17 (1) as floralis var. squamosa Gory & Pecheron
Tunisia 1Cetoniinae (1) Pal.17 (as floralis var. squamosa Gory & Pecheron)

Afrodiastictus ceratus (Clement,1969)   Sierra Leone 2Aphodiinae Afr.15 (2)
Afrodrepanus impressicollis Boheman,1857   South Africa 3Scarabaeinae Afr.58 (3)
Afrodrepanus marshalli (Boucomont,1921)   Kenya 1Scarabaeinae Afr.58 (1)
Aganocrossus amoenus (Boheman,1857)   Ghana 21Aphodiinae Afr.17 (21)
Namibia 4Aphodiinae (4) Afr.22
South Africa 10Aphodiinae Afr.22 (8)
Afr.32 (2)
Sri Lanka 21Aphodiinae (21) Or.1 (as A.khutianus Bl.)+6
Aganocrossus amoenus ab.fuscus (Petrovitz)   Gambia 47Aphodiinae Afr.4 (47)
Senegal 25Aphodiinae Afr.4 (25)
Sierra Leone 1Aphodiinae (1) Afr.15
Aganocrossus ciprianii (Balthasar,1939)   Zambia 1Aphodiinae Afr.30 (1)
Aganocrossus clementianus (Endrödi,1964)   Gambia 1Aphodiinae (1) Afr.20
Sierra Leone 2Aphodiinae (2) Afr.15
Aganocrossus fugitivus (Péringuey,1901)   South Africa 36Aphodiinae Afr.32 (24)
Afr.34 (12)
Aganocrossus hutteri (Petrovitz,1973)   Burkina Faso 1Aphodiinae Afr.30 (1)
Aganocrossus krikkeni (Bordat,1992)   Kenya 1Aphodiinae Afr.4 (1)
Aganocrossus meticulosus (Paulian,1942)   Kenya 11Aphodiinae (11) Afr.4+24
Aganocrossus pallidicornis (Walker,1858)   India 1Aphodiinae Or.7 (1)
Aganocrossus pseudourostigma (Balthasar,1939)   Ethiopia 53Aphodiinae Afr.4 (51)
Afr.27 (2)
Kenya 30Aphodiinae Afr.27 (30)
Tanzania 21Aphodiinae Afr.4 (21)
Aganocrossus urostigma (Harold,1862)   Ethiopia 1Aphodiinae Afr.27 (1)
Japan 1Aphodiinae Pal.28 (1)
Lesotho 44Aphodiinae Afr.21 (44)
Malaysia 6Aphodiinae Or.3 (6)
Nepal 12Aphodiinae Or.3 (12)
South Africa 22Aphodiinae (22) Afr.21+25
Sri Lanka 10Aphodiinae Or.1 (10)
Tanzania 2Aphodiinae Afr.26 (2)
Thailand 7Aphodiinae Or.3 (7)
Aganocrossus vestitus (Boheman,1857)   Ethiopia 47Aphodiinae Afr.29 (47)
Kenya 1Aphodiinae Afr.4 (1)
South Africa 1Aphodiinae Afr.34 (1)
Aganocrossus zebrinus (Petrovitz,1958)   South Africa 3Aphodiinae (3) Afr.16 (incl. 3 paratypes)

Agaocephala cornigera Serville,1825   Brazil 1Dynastinae S.Am.17 (1)
Unknown 3Dynastinae S.Am.17 (3)
Agaocephala inermicollis Arrow,1914   Brazil 1Dynastinae S.Am.18 (1)
Unknown 2Dynastinae S.Am.18 (2)
Agenius clavus Schaum,1844   South Africa 1Cetoniinae Afr.20 (1)
Agenius jansoni Peringuey,1908   Lesotho 1Cetoniinae Afr.20 (1)
Agestrata orichalca (Linnaeus,1789)   China 1Cetoniinae Pal.27 (1)
Agestrata spp.    Unknown 1Sydasien
Cetoniinae Or.4 (1)
Agoliinus aleutus Eschscholtz,1822   Canada 6British Columbia
Aphodiinae N.Am.1 (6)
Agoliinus lapponum Gyllenhal,1808   Faroe Islands 8Aphodiinae Main (8)
Norway 41Aphodiinae Main (41)
Sweden 358  Vs Dr Gä Hs Me Hr Jä Ån Nb Ås Ly Pi Lu To
Alan Dufberg 48 (27)
Aphodiinae Main (330)
Type Coleoptera (1) Lectotype 06298:1 of Aphodius rhenonum Zetterstedt 1828
  Aphodius rhenonum Zetterstedt 1828

Agoliinus piceus Gyllenhal,1808   Norway 22Aphodiinae Main (22)
Sweden 354  Sö Vr Dr Hs Me Hr Jä Vb Nb Ly Pi Lu To
Alan Dufberg 48 (33)
Aphodiinae Main (321)
Agrilinus ater (DeGeer,1774)   Sweden 559  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up Vr Dr Gä Hs Jä Vb
Alan Dufberg 48 (26)
Aphodiinae Main (533)
Agrilinus breviusculus Motschulsky,1866   Japan 2Aphodiinae Pal.17 (1)
Pal.28 (1)
Agrilinus lindbergi (Petrovitz,1959)   Afghanistan 6Type Coleoptera (6) Holotype Paratype 00928:1-6 of Aphodius lindbergi Petrovitz 1961
  Aphodius lindbergi Petrovitz 1961

Agrilinus nemoralis Erichson,1848   Sweden 174  Sk Ha Sm Öl Ög Up Vs Vr Dr Gä Jä To Nb
Alan Dufberg 48 (23)
Aphodiinae Main (150)
Type Landin (1) Holotype 03482:1 of Aphodius ardoei Landin 1954
  Aphodius ardoei Landin 1954

Agrilinus ruricola Melsheimer,1844   United States 4Aphodiinae N.Am.6 (4)
Aidophus flaveolus (Harold,1867)   Argentina 1Aphodiinae S.Am.2 (1)
Aidophus parcus (Horn,1887)   United States 1Florida
Aphodiinae N.Am.8 (1)
Alissonotum africanum Endrödi,1974   Cameroon 1Dynastinae Afr.5 (1)
Unknown 1Dynastinae Afr.5 (1)
Alissonotum cribratellum (Fairmaire,1893)   Laos 1Dynastinae Or.10 (1)
Thailand 5Dynastinae Or.10 (5)
Alissonotum piceum (Fabricius,1775)   Sri Lanka 2Dynastinae Or.3 (2)
Alissonotum piceum krombeini (Fabricius,1775)   Sri Lanka 4Dynastinae Or.9 (4)
Allidiostoma strobeli (Steinheil,1872)   Argentina 4Allidiostomatinae S.Am.1 (4)
Allogymnopleurus chloris (Klug,1855 )   Unknown 4Scarabaeinae Afr.29 (4)
Allogymnopleurus consoicus (Peringuey,1901)   South Africa 1Scarabaeinae (1) Afr.30
Allogymnopleurus thalassinus var. subcupratus (Fåhraeus,,1857)   South Africa 4Scarabaeinae (4) Afr.29
Allogymnopleurus umbrinus Gerstaecker,1871   Tanzania 3Scarabaeinae Afr.57 (3)
Allomyrina dichotoma (Linnaeus,1771)   Japan 8Dynastinae (8) Pal.4+5
Taiwan 1Dynastinae Or.12 (1)
Unknown 3Dynastinae Pal.4 (3)
Allomyrina spp.    Thailand 2Dynastinae Or.11 (2)
Alocoderus hydrochaeris (Fabricius,1798)   Egypt 5Aphodiinae (5) Pal.2+5+18
France 2Aphodiinae Pal.18 (2)
Greece 1Aphodiinae Pal.18 (1)
Italy 1Aphodiinae Pal.10 (1)
Morocco 4Aphodiinae (4) Pal.5+18
Portugal 10Madeira
Aphodiinae Pal.3 (10)
Russian Federation 1Aphodiinae (1) Pal.17 (as ssp. tataricus Harold)

Spain 4Aphodiinae (4) Pal.2+5
Spain 24Canary Islands
Aphodiinae (19) Pal.3+10+15
Pal.27 (3)
Pal.28 (1)
Pal.30 (1)
Spain 3Mallorca
Aphodiinae Pal.24 (3)
Tunisia 11Aphodiinae (11) Pal.5+16
Alocoderus procerus (Harold,1867)   South Africa 2Aphodiinae (2) Afr.23
Unknown 2Aphodiinae Afr.26 (2)
Alocoderus turbatus (Baudi,1870)   Cyprus 1Aphodiinae Pal.11 (1) as Aphodius lineimargo Reitter
Amazula suavis Burmeister,1847   Namibia 1Type Schein (1) Lectotype 00002:1 of Amazula suavis hobohmi Schein 1960
  Amazula suavis hobohmi Schein 1960

Ammoecioides davisianus Bordat,1999   South Africa 1Aphodiinae Afr.31 (1)
Ammoecius brevis Erichson,1848   Finland 1Aphodiinae Main (1) as Aphodius b.
France 2Aphodiinae Pal.15 (1)
Pal.18 (1)
Russian Federation 2Aphodiinae Pal.5 (2) as A. scroris
Sweden 199  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Dr Gä Jä
Alan Dufberg 47 (18) as Aphodius b.
Aphodiinae Main (181) as Aphodius b.
United States 1Connecticut
Aphodiinae (1) N.Am.
Unknown 1Aphodiinae Pal.18 (1)
Ammoecius dentatus Schmidt,1908   Morocco 1Aphodiinae Pal.17 (1)
Ammoecius elevatus (Olivier,1789)   France 1Aphodiinae (1) Pal.17+18
Morocco 1Aphodiinae Pal.17 (1)
Ammoecius franzi Petrovitz,1964   Morocco 1Aphodiinae Pal.17 (1)
Ammoecius lugubris Boheman,1857   South Africa 13Aphodiinae (13) Afr.23+24
Ammoecius lusitanicus Erichson,1848   Morocco 4Aphodiinae Pal.16 (4)
Ammoecius mimus Peringuey,1901   South Africa 6Aphodiinae (6) Afr.23+24+25
Ammoecius muchei Petrovitz,1962   Turkey 7Aphodiinae Pal.11 (6)
Pal.25 (1) uncertain id.
Ammoecius rugifrons Aubé,1850   Tunisia 1Aphodiinae Pal.10 (1)
Ammoecius terminatus Harold,1869   South Africa 7Aphodiinae Afr.34 (7)
Amphimallon arianae (Fairmaire,1879)   Greece 1Melolonthinae Pal.4 (1)
Amphimallon assimile (Herbst,1790)   Austria 2Melolonthinae Pal.9 (2)
Germany 1Melolonthinae Pal.11 (1)
Amphimallon atrum (Herbst,1790)   France 3Melolonthinae Pal.9 (3)
Germany 1Melolonthinae Pal.4 (1)
Amphimallon caucasicum (Gyllenhal,1817)   Bulgaria 5Melolonthinae Pal.9 (5)
Greece 2Melolonthinae Pal.9 (2)
Turkey 14Melolonthinae Pal.14 (8)
Pal.16 (4)
Pal.18 (2)
Amphimallon cf. solstitiale Medvedev,1951   Unknown 3Melolonthinae Pal.4 (3)
Amphimallon fallenii (Gyllenhal,1817)   Denmark 1Melolonthinae Main (1)
Sweden 88  Skåne Blekinge Halland Småland Öland
Alan Dufberg 48 (9)
Melolonthinae Main (79)
Amphimallon fissiceps (Fairmaire,1860)   Morocco 1Melolonthinae Pal.3 (1)
Amphimallon fuscum (Scopoli,1786)   Italy 1Melolonthinae Pal.3 (1)
Amphimallon majale (Razoumowski,1789)   Austria 1Melolonthinae Pal.3 (1)
Germany 1Melolonthinae Pal.11 (1)
Spain 1Melolonthinae Pal.3 (1)
Switzerland 1Melolonthinae Pal.3 (1)
Amphimallon nigrum (Waltl,1835)   Spain 2Melolonthinae Pal.4 (2)
Amphimallon ochraceum (Knoch,1801)   Albania 1Melolonthinae Pal.13 (1)
France 2Melolonthinae Pal.4 (2)
Germany 1Melolonthinae Pal.9 (1)
Amphimallon parvulus Rosenhauer,1856   Spain 1Melolonthinae Pal.4 (1)
Amphimallon pini (Olivier,1789)   France 85Melolonthinae Pal.4 (85)
Amphimallon pseudomajale Sabatinelli,1977   Italy 1Melolonthinae Pal.9 (1)
Amphimallon pygiale Mulsant,1846   Spain 1Melolonthinae Pal.4 (1)
Amphimallon ruficorne (Fabricius,1773)   Germany 2Melolonthinae Pal.4 (2)
Amphimallon seidlitzi (Brenske,1891)   Spain 5Melolonthinae (5) Pal.4+10
Amphimallon solstitiale Medvedev,1951   Afghanistan 2Melolonthinae Pal.10 (2)
Austria 1Melolonthinae Pal.9 (1)
Denmark 3Melolonthinae (3)
Russian Federation 3Melolonthinae Pal.4 (3)
Sweden 348  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up Vs Vr Dr Gä
Alan Dufberg 48 (6)
Melolonthinae Main (342) incl. larvae
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 4Melolonthinae Pal.9 (4)
Amphimallon solstitiale mesasiaticum Medvedev,1951   Uzbekistan 1Melolonthinae Pal.13 (1) (Not in Pal.13)
Amphimallon solstitiale sibiricum Medvedev,1951   Russian Federation 1Melolonthinae Pal.13 (1) (Not in Pal.13)
Amphimallon vivesi Baraud,1967   Unknown 1Melolonthinae Pal.4 (1)
Anachalcos aurescens Bates,1888   Guinea-Bissau 1Scarabaeinae Afr.19 (1)
Sierra Leone 1Scarabaeinae Afr.2 (1)
Tanzania 1Scarabaeinae Afr.46 (1)
Anachalcos convexus (Boheman,1857)   Ethiopia 1Scarabaeinae Afr.2 (1)
Guinea-Bissau 5Scarabaeinae Afr.2 (5)
Kenya 6Scarabaeinae (6) Afr.19
Namibia 1Scarabaeinae (1) Afr.32
Sierra Leone 1Scarabaeinae Afr.2 (1)
South Africa 1Scarabaeinae Afr.2 (1)
Anachalcos cupreus Fabricius,1775   Guinea 2Scarabaeinae Afr.46 (2)
Senegal 1Scarabaeinae Afr.46 (1)
South Africa 1Scarabaeinae Afr.46 (1)
Tanzania 3Scarabaeinae Afr.46 (3)
Tanzania 1Scarabaeinae Afr.46 (1)
Unknown 3Scarabaeinae Afr.46 (3)
Anachalcos procerus Gerstaecker,1874   Malawi 1Scarabaeinae Afr.19 (1)
Tanzania 1Scarabaeinae Afr.47 (1)
Togo 1Scarabaeinae Afr.47 (1)
Ancistrosoma blanchardi Sallé,1886   Colombia 2Melolonthinae S.Am.1 (2)
Ancistrosoma farinosum Sallé,1849   Unknown 1Melolonthinae S.Am.1 (1)
Venezuela 1Melolonthinae S.Am.1 (1)
Ancognatha scarabaeoides Erichson,1847   Bolivia 1Dynastinae (1) S.Am.30
Colombia 2Dynastinae S.Am.19 (2)
Ecuador 1Dynastinae (1) S.Am.35
Unknown 2Dynastinae S.Am.19 (2)
Ancognatha ustulata Burmeister,1847   Venezuela 1Dynastinae S.Am.19 (1)
Ancognatha vulgaris Arrow,1911   Bolivia 1Dynastinae (1) S.Am.30
Colombia 1Dynastinae S.Am.19 (1)
Unknown 1Dynastinae S.Am.19 (1)
Anisocanthon sericinus (Harold,1868)   Argentina 1Scarabaeinae S.Am.6 (1)
Anisochelus neglectus Burmeister,1844   South Africa 18Hopliini Afr.12 (18)
Anisonyx alticola Andreae,1968   South Africa 40Hopliini (40) Afr.18+19
Anisonyx badiitibialis Schein,1959   South Africa 4Type Schein (4) Holotype Paratype 02000:1-4 of Anisonyx badiitibialis Schein 1959
  Anisonyx badiitibialis Schein 1959

Anisonyx brincki Schein,1959   South Africa 26Type Schein (26) Holotype Paratype 02001:1-26 of Anisonyx brincki Schein 1959
  Anisonyx brincki Schein 1959

Anisonyx brincki rudebecki Schein,1959   South Africa 3Type Schein (3) Holotype Paratype 02002:1-3 of Anisonyx brincki rudebecki Schein 1959
  Anisonyx brincki rudebecki Schein 1959

Anisonyx crinitum Latreille   South Africa 1Hopliini Afr.20 (1)
Unknown 2Hopliini Afr.20 (2)
Anisonyx ditus Peringuey,1902   South Africa 2Hopliini Afr.2 (2)
Anisonyx ditus ssp.pseudoditus Peringuey,1902   South Africa 3Hopliini (3) Types 2003:2-4
Anisonyx hessei Schein,1959   South Africa 2Type Schein (2) Paratype 01999:2-3 of Anisonyx hessei Schein 1959
  Anisonyx hessei Schein 1959

Anisonyx hilaris Peringuey,1902   South Africa 2Hopliini Afr.1 (2)
Anisonyx ignitus Castelnau,1840   South Africa 2Hopliini Afr.1 (2)
Anisonyx lepidotus Wiedemann,1821   South Africa 2Hopliini Afr.1 (2)
Anisonyx longipes Linnaeus,1764   South Africa 3Hopliini Afr.2 (3)
Anisonyx lynx Fabricius,1776   South Africa 4Hopliini Afr.1 (4)
Anisonyx nasuus Wiedemann,1821   South Africa 2Hopliini Afr.2 (2)
Anisonyx nigerrimus Schein,1959   South Africa 2Type Schein (2) Paratype 02004:2-3 of Anisonyx nigerrimus Schein 1959
  Anisonyx nigerrimus Schein 1959

Anisonyx pauperatus Peringuey,1902   South Africa 2Hopliini Afr.1 (2)
Anisonyx proletarius Peringuey,1902   South Africa 2Hopliini Afr.2 (2)
Anisonyx pseudomilitaris Schein,1959   South Africa 2Hopliini Afr.1 (2)
Anisonyx sesuto Peringuey,1908   South Africa 55Hopliini Afr.1 (55)
Anisonyx ursus Fabricius,1776   South Africa 10Hopliini (10) Afr.2+19+20
Unknown 1Hopliini Afr.20 (1)
Anisoplia agnata Reitter,1889   Russian Federation 1Rutelinae Pal.8 (1)
Anisoplia agricola (Poda,1761)   Bosnia and Herzegovina 3Bosnia
Rutelinae Pal.7 (3) as A.cyantigera Scopoli
Germany 2Rutelinae Pal.7 (2)
Germany 1Rutelinae Pal.9 (1) as A.cyantigera Scopoli
Greece 6Rutelinae Pal.3 (6)
Hungary 1Rutelinae (1) Pal.7 (as A.cyantigera Scopoli)
Russian Federation 1Rutelinae Pal.7 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 9Rutelinae Pal.3 (9)
Unknown 4Rutelinae Pal.7 (4)
Anisoplia andreae Baraud,1991   Greece 1Rutelinae Pal.10 (1)
Anisoplia angorensis Petrovitz,1971   Turkey 1Rutelinae Pal.12 (1)
Anisoplia austriaca (Herbst,1783)   Austria 7Rutelinae (7) Pal.3+8
Bulgaria 2Rutelinae Pal.11 (2)
Germany 1Rutelinae Pal.9 (1)
Turkey 2Rutelinae Pal.12 (1)
Pal.13 (1)
Anisoplia austriaca hordearia Petrovitz,1959   Greece 3Rutelinae Pal.3 (3)
Anisoplia austriaca major Reitter,1889   Armenia 1Rutelinae Pal.10 (1)
Anisoplia baetica Erichson,1847   Spain 4Rutelinae Pal.7 (4)
Anisoplia bodenmeyeri?    Iran 1Rutelinae Pal.9 (1)
Anisoplia bromicola (Germar,1817)   Italy 1Rutelinae Pal.7 (1)
Anisoplia bulgarica Apfelbeck,1909   Italy 1Rutelinae Pal.8 (1)
Anisoplia bureschi  Sacharieva-Stoilova,1958   Greece 1Rutelinae Pal.10 (1)
Anisoplia campicola Menetries,1832   Kyrgyzstan 1Rutelinae Pal.10 (1)
Anisoplia deserticola Fischer,1824   Russian Federation 2Rutelinae Pal.7 (2)
Anisoplia dispar Erichson,1847   Greece 1Rutelinae Pal.8 (1)
Anisoplia erichsoni Reitter,1889   Bosnia and Herzegovina 2Bosnia
Rutelinae Pal.7 (2)
Anisoplia faldermanni Reitter,1883   Iran 1Rutelinae Pal.9 (1)
Anisoplia farraria Erichson,1847   Russian Federation 8Rutelinae Pal.8 (8)
Anisoplia flavipennis Brullé,1832   Greece 11Rutelinae (11) Pal.3+8
Turkey 1Rutelinae Pal.12 (1)
Anisoplia hebes Reitter,1903   Greece 1Rutelinae Pal.8 (1)
Anisoplia horticola (Linnaeus,1758)   Austria 6Rutelinae Pal.1 (6)
Germany 2Rutelinae Pal.1 (2)
Romania 1Rutelinae Pal.1 (1)
Spain 5Rutelinae Pal.1 (5)
Switzerland 2Rutelinae Pal.1 (2)
Anisoplia lata Erichson,1847   Austria 11Rutelinae (11) Pal.2+3
Greece 2Rutelinae Pal.13 (2)
Hungary 7Rutelinae Pal.8 (7)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 5Rutelinae Pal.2 (5)
Anisoplia lata lamiensis (Apfelbeck,1909)   Greece 6Rutelinae Pal.3 (6)
Anisoplia monticola (Erichson,1847)   Italy 6Rutelinae Pal.2 (6)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 7Rutelinae Pal.2 (7)
Anisoplia pubipennis Blanchard,1851   Greece 13Rutelinae (13) Pal.2+3
Tunisia 1Rutelinae Pal.13 (1) as Anisoplia pallidipennis
Anisoplia reitteriana Semenov,1903   Armenia 1Rutelinae Pal.10 (1)
Anisoplia remota Reitter,1889   Spain 1Rutelinae Pal.7 (1)
Anisoplia sabatinellii Baraud,1991   Italy 1Rutelinae Pal.10 (1)
Anisoplia signata Faldermann,1835   Russian Federation 3Rutelinae Pal.8 (3)
Anisoplia spp.    Croatia 1Rutelinae Pal.13 (1)
Unknown 1Rutelinae Pal.12 (1) Female
nec segetum
Anisoplia tempestiva Erichson,1847   Bosnia and Herzegovina 2Bosnia
Rutelinae Pal.8 (2)
France 7Rutelinae Pal.8 (7)
Hungary 1Rutelinae Pal.8 (1)
Italy 1Rutelinae Pal.8 (1)
Romania 1Rutelinae Pal.13 (1)
Anisoplia tenebralis Burmeister,1844   Turkey 2Rutelinae Pal.3 (2)
Anisoplia thessalica Reitter,1889   Greece 1Rutelinae Pal.1 (1)
Anisoplia villosa (Goeze,1777)   France 1Rutelinae Pal.7 (1)
Greece 1Rutelinae Pal.7 (1)
Romania 1Rutelinae Pal.13 (1)
Spain 1Rutelinae Pal.1 (1)
Anisoplia zwickii Fischer von Waldheim,1824   Russian Federation 2Rutelinae Pal.8 (2)
Anochilia incilis Janson,1885   Madagascar 1Cetoniinae (1) Afr.17
Anochilia nitida Pouillaude,1915   Madagascar 1Cetoniinae (1) Afr.17
Anomala aeneiventris Fairmaire,1883   Borneo, Indonesia / Malaysia 2Rutelinae Or.7 (2) as Anomala atriplicis F.
Anomala affinis Ganglbauer,1882   Syria 1Rutelinae (1) Pal 5 (as Euchlora a.)

Turkey 1Rutelinae (1) Pal 1 (as Euchlora a.)

Unknown 2Rutelinae (2) Pal 5 (as Euchlora a.)

Anomala albopilosa (Hope,1839)   Japan 1Rutelinae (1) Pal 4 (as Euchlora a.)

Anomala anoguttata Burmeister,1844   Unknown 1Rutelinae Or.? (1)
Anomala antiqua (Gyllenhal,1817)   Australia 1Rutelinae Pac.1 (1)
Unknown 2Rutelinae (2) Pac.1
Anomala aulax Wiedemann   Viet Nam 1Rutelinae Or.8 (1)
Anomala ausonia Erichson,1847   Italy 1Rutelinae (1) Pal.5 (as Euchlora a.)

Spain 4Rutelinae (4) Pal 1 (as Euchlora a.)

Tunisia 1Rutelinae Pal.10 (1)
Anomala calcarata Arrow,1899   South Africa 1Rutelinae Afr.1 (1)
Anomala callifera Ohaus,1937   Myanmar 2Rutelinae (2) Or.6 (incl. types 3247:2-3)

Anomala calligrapha Bates,1888   Costa Rica 1Rutelinae S.Am.7 (1)
Anomala chalybaea Burmeister,1844   Philippines 2Rutelinae Or.6 (2)
Anomala chamaeleon Fairmaire,1887   Korea, Republic of 1Rutelinae Pal.5 (1)
Mongolia 2Rutelinae Pal.5 (2)
Anomala chloromela Arrow,1911   Sri Lanka 6Rutelinae Or.4 (6)
Anomala chloronota Arrow,1911   Sri Lanka 4Rutelinae Or.4 (4)
Anomala cincta Say,1835   Colombia 3Rutelinae S.Am.1 (3)
Anomala conformis Walker,1859   Sri Lanka 2Rutelinae Or.2 (2)
Anomala corpulenta Motschulsky,1854   China 3Rutelinae Pal.5 (3)
Korea, Republic of 2Rutelinae Pal.5 (2)
Anomala cuprea (Hope,1839)   Japan 9Rutelinae (9) Pal.4+5+10
Anomala cupricollis Chevrolat,   Mexico 1Rutelinae N.Am.2 (1)
Anomala cupripes (Hope,1839)   Indonesia 1Sumatra
Rutelinae Or.? (1)
Korea, Republic of 1Rutelinae Pal.4 (1) as Euchlora c.
Malaysia 2Rutelinae Or.4 (2)
Taiwan 2Rutelinae Pal.4 (2) as Euchlora c.
Thailand 8Rutelinae Or.5 (8)
Anomala daimiana Harold,1877   Japan 4Rutelinae Pal.4 (4)
Anomala denticrus (Ohaus,1910)   Borneo, Indonesia / Malaysia 2Rutelinae (2) Or.6 (as Euchlora d.)

Anomala denuda Arrow,1899   Cameroon 4Rutelinae Afr.2 (4)
Gabon 3Rutelinae Afr.2 (3)
South Africa 1Rutelinae Afr.2 (1)
Tanzania 3Rutelinae Afr.3 (3)
Anomala devota (Rossi,1790)   France 1Rutelinae Pal.4 (1)
Italy 1Rutelinae Pal.11 (1)
Spain 4Rutelinae Pal.1 (2) as Euchlora d.
Pal.4 (2)
Unknown 3Rutelinae (3) Pal.4+9
Anomala digressa Casey,   United States 1Arizona
Rutelinae N.Am.2 (1)
Anomala dimidiata (Hope,1831)   India 1Rutelinae Or.7 (1)
Anomala distanti Arrow,1899   Namibia 2Rutelinae Afr.1 (2)
South Africa 3Rutelinae Afr.1 (3)
Anomala distinguenda Blnchard,1850   Kenya 2Rutelinae Afr.3 (2)
Senegal 1Rutelinae Afr.2 (1)
Anomala dita Peringuey,1902   South Africa 8Rutelinae Afr.1 (8)
Anomala dorsalis (Fabricius,1775)   Unknown 2Rutelinae Or.7 (2)
Anomala dubia (Scopoli,1763)   Austria 11Rutelinae Pal.5 (11)
Austria 1Rutelinae (1) Pal 1 (as Euchlora d.)

Denmark 12Rutelinae Main (12)
Finland 3Rutelinae Main (3)
Germany 2Rutelinae (2) Pal.9+10
Germany 1Rutelinae (1) Pal 1 (as Euchlora d.)

Italy 1Rutelinae Pal.5 (1)
Netherlands 2Rutelinae Pal.10 (2)
Sweden 377  Skåne Blekinge Halland Öland Östergötland Västergötland Uppland Ångermanland
Alan Dufberg 48 (12)
Rutelinae Main (365)
Switzerland 2Rutelinae Pal.10 (2)
Switzerland 3Rutelinae (3) Pal 1 (as A.aenea DeGeer)

Anomala dussumieri Blanchard,1850   Sri Lanka 6Rutelinae Or.3 (6)
Anomala errans (Fabricius,1775)   Russian Federation 1Rutelinae Pal.4 (1)
Anomala exoleta Faldermann,1835   China 3Rutelinae Pal.4 (3)
Mongolia 1Rutelinae Pal.4 (1)
South Africa 7Rutelinae Afr.1 (7)
Anomala geniculata (Motschulsky,1866)   Japan 2Rutelinae Pal.11 (2)
Anomala gracilenta Reitter,1903   China 1Rutelinae Pal.10 (1)
Anomala grandis Hope,1839   India 1Rutelinae Or.? (1)
Indonesia 1Java
Rutelinae Or.? (1)
Anomala gravida Arrow,1911   Sri Lanka 15Rutelinae Or.3 (15)
Anomala immarginata Ohaus,1902   Cameroon 1Type Ohaus (1) Paratype 03249:2 of Anomala immarginata Ohaus 1902
  Anomala immarginata Ohaus 1902

Anomala immatura Boheman,1860   Unknown 2Rutelinae Afr.2 (2)
Anomala inconstans Burmeister,1844   Guatemala 1Rutelinae S.Am.1 (1)
Anomala leonfairmairei Zorn,2005   China 1Rutelinae (1) Pal.4 (as Anomala micans Fairmaire)
Anomala lucens Ballion,1871   China 2Rutelinae Pal.4 (2)
India 2Rutelinae Pal.4 (2)
Japan 1Rutelinae Pal.4 (1)
Anomala lucicola (Fabricius,1798)   United States 8Connecticut
Rutelinae N.Am.1 (8)
Anomala luculenta Erichson,1847   Bulgaria 2Rutelinae Pal.11 (2)
Japan 4Rutelinae Pal.4 (4)
Korea, Republic of 1Rutelinae Pal.4 (1)
Anomala malaisei Ohaus,1937   Myanmar 2Rutelinae (2) Or.6 (incl. types 3248:2-3)

Anomala marginipennis Arrow,1912   India 1Rutelinae Or.8 (1)
Anomala megalops Bates,   Mexico 1Rutelinae N.Am.2 (1)
Anomala mongolica Faldermann,1835   Korea, Republic of 1Rutelinae Pal.4 (1)
Anomala mychus ssp.fumosus Germ.   Brazil 3Rutelinae S.Am.1 (3)
Anomala occulicollis Arrow,1911   India 4Rutelinae Or.7 (4)
Anomala octiecostata Burmeister,1844   Japan 2Rutelinae Pal.10 (2)
Anomala ogloblini Medverev,1949   Russian Federation 1Rutelinae Pal.10 (1)
Anomala osakana Sawada,1942   Japan 2Rutelinae Pal.10 (2)
Anomala osmanlis Blanchard,1851   Syria 1Rutelinae Pal.2 (1)
Turkey 6Rutelinae Pal.2 (2)
Pal.13 (4)
Anomala pellucida Arrow,1911   Sri Lanka 1Rutelinae Or.4 (1)
Anomala praticola? (Fabricius,1781)   Romania 3Rutelinae Pal.3 (3)
Anomala propinqua Arrow,1912   Nepal 1Rutelinae Or.8 (1)
Anomala puncticollis? Harold,1877   Japan 1Rutelinae Pal.10 (1)
Anomala quadripunctata (Olivier,1789)   Spain 1Rutelinae Pal.4 (1)
Anomala rufiventris Redtenbacher,1844   India 2Rutelinae Or.7 (2)
Anomala rufocuprea Motschulsky,1860   Japan 7Rutelinae Pal.4 (7)
Anomala semicastanea Fairmaire,1888   China 1Rutelinae Pal.4 (1)
Anomala seminigra Lansb.,1883   Indonesia 1Nias
Rutelinae (1) Or.6 (as Euchlora s.)

Anomala semipurpurea Burmeister,1855   Indonesia 1Sumatra
Rutelinae Or.? (1)
Anomala separata Brancsik,1897   South Africa 4Rutelinae Afr.1 (4)
Anomala sieversi Heyden,1887   Japan 1Rutelinae Pal.4 (1)
Anomala smaragdina Eschscholtz,1822   Philippines 1Rutelinae Or.? (1)
Unknown 1Rutelinae Or.? (1)
Anomala solida Erichson,1847   Bosnia and Herzegovina 1Rutelinae Pal.11 (1)
Bulgaria 4Rutelinae Pal.1 (4) as Euchlora s.
China 2Rutelinae Pal.4 (2)
Greece 2Rutelinae (2) Pal 1 (as Euchlora s.)

Turkey 1Rutelinae (1) Pal 1 (as Euchlora s.)

Anomala sp.cf.fracta Walker   Sri Lanka 9Rutelinae Or.4 (9)
Anomala spiloptera Burmeister,1855   China 1Rutelinae Pal.4 (1)
Anomala spp.    China 1Rutelinae (1) Pal 1
Sri Lanka 5Rutelinae Or.4 (5)
Sri Lanka 9Rutelinae Or.4 (9)
Anomala superflua Arrow,1911   Sri Lanka 15Rutelinae Or.3 (15)
Anomala testacea Hope,1831   India 1Rutelinae Or.8 (1)
Anomala testaceipennis Blanchard,1856   Paraguay 2Rutelinae S.Am.1 (2)
Anomala triangularis Schönfeldt,1890   Japan 1Rutelinae Pal.10 (1)
Anomala truncata H.Bates,1890   China 1Rutelinae (1) Or.6 (as Euchlora t.)

Indonesia 1Java
Rutelinae (1) Or.6 (as Euchlora t.)

Malaysia 1Rutelinae (1) Or.6 (as Euchlora t.)

Anomala undulata Melsheimer,1844   United States 1Pennsylvania
Rutelinae N.Am.1 (1)
Anomala valida Burmeister,1844   Ecuador 2Rutelinae S.Am.1 (2)
Anomala varians? (Olivier,1789)   Japan 1Rutelinae Pal.5 (1)
Anomala varicolor Arrow,1911   Malaysia 6Rutelinae Or.4 (6)
Sri Lanka 24Rutelinae Or.4 (24)
Anomala variegata Hope,1831   India 1Rutelinae Or.8 (1)
Anomala viridana Kolbe,1886   Japan 1Rutelinae (1) Pal 1
Anomala viridis (Fabricius,1775)   Unknown 1Rutelinae Or.? (1)
Anomala viridis? (Fabricius,1775)   China 1Rutelinae Or.7 (1) as Euchlora v.
Indonesia 1Java
Rutelinae Or.6 (1) as Euchlora v.
Malaysia 3Rutelinae Or.6+8 (3) as Euchlora v.
Sri Lanka 1Rutelinae Or.6 (1) as Euchlora v.
Unknown 4Rutelinae Or.6+7 (4) as Euchlora v.
Anomala vitis (Fabricius,1775)   Bosnia and Herzegovina 1Bosnia
Rutelinae Pal.4 (1) as Euchlora v.
Germany 1Rutelinae Pal.9 (1)
Greece 2Rutelinae (2) Pal 1+4 (as Euchlora v.)

Hungary 1Rutelinae (1) Pal 1 (as Euchlora v.)

Italy 4Rutelinae Pal.4 (2) as Euchlora v.
Pal.13 (2)
Turkey 16Rutelinae Pal.9 (3)
Pal.12 (11)
Pal.13 (2)
Anomala vittata Gebler,1841   China 2Xinjiang
Rutelinae Pal.4 (2)
Anomala walkeri Arrow,1911   Sri Lanka 134Rutelinae (134) Or.1+2
Anomalomorpha giveni Carne,1957   Australia 2Dynastinae Pac.2 (2)
Anomiopus spp.    Peru 10Scarabaeinae S.Am.2 (10)
Anomius annamariae Baraud,1982   Spain 1Aphodiinae Pal.17 (1)
Unknown 1Aphodiinae Pal.28 (1)
Anomius castaneus Illiger,1803   Tunisia 2Aphodiinae Pal.21 (2)
Anomius peyerimhoffi Thery,1925   Morocco 1Aphodiinae Pal.15 (1)
Anoplocheilus figuratus Boheman,1857   South Africa 3Cetoniinae (3) Afr.14
Anoplognathus abnormis MacLeay,1873   Australia 1Rutelinae (1) Pac.2
Anoplognathus aeneus Waterhouse,1868   Australia 1Rutelinae (1) Pac.2
Anoplognathus boisduvalii Boisduval,1835   Australia 2Rutelinae (2) Pac.1
Anoplognathus brunnipennis (Gyllenhal,1817)   Australia 1Rutelinae (1) Pac.2
Anoplognathus chloropyrus (Drapiez,1819)   Unknown 1Rutelinae (1) Pac.2
Anoplognathus concolor Burmeister,1855   Australia 1Rutelinae (1) Pac.2
Anoplognathus olivieri (Dalman,1817)   Australia 8Rutelinae (8) Pac.1
Australia 1Tasmania
Rutelinae (1) Pac.1
Anoplognathus porosus (Dalman,1817)   Australia 4Rutelinae (4) Pac.1
New Zealand 1Rutelinae (1) Pac.1
Anoplognathus punctulatus Olliff,1890   Australia 1Rutelinae (1) Pac.2
Anoplognathus rugosus Kirby,1818   Australia 5Rutelinae (5) Pac.1
Anoplognathus spp.    Australia 3Rutelinae Pac.2 (3)
Anoplognathus velutinus Boisduval,1835   Australia 2Rutelinae (2) Pac.2
Unknown 1Rutelinae Pac.2 (1)
Anoplognathus viridiaeneus (Donovan,1805)   Australia 2Rutelinae (2) Pac.2
Anoplognathus viriditarsis Leach,1815   Australia 10Rutelinae (10) Pac.2
Australia 3Rutelinae (3) Pac.2
Anoplostethus opalinus Brullé,1837   Australia 1Rutelinae Pac.3 (1)
Anoxia australis (Gyllenhal,1817)   France 2Melolonthinae Pal.7 (2)
Italy 1Melolonthinae Pal.1 (1)
Spain 14Melolonthinae (14) Pal.1+7
Anoxia cypria Zurcher,1911   Cyprus 2Melolonthinae Pal.18 (2) as Anoxia mavromoustakisi
Anoxia emarginata Coquerel,1860   Morocco 1Melolonthinae Afr.2 (1)
Anoxia maldesi Baraud,1980   Tunisia 1Melolonthinae Pal.12 (1)
Anoxia matutinalis matutinalis Laporte,1832   Italy 3Melolonthinae Pal.12 (3)
Anoxia matutinalis sardoa Laporte,1832   Italy 4Melolonthinae Pal.1 (4)
Anoxia naviauxi Baraud,1990   Turkey 3Melolonthinae Pal.15 (1)
Pal.16 (2)
Anoxia orientalis (Krynicky,1832)   Bulgaria 12Melolonthinae (12) Pal.5+6
Greece 1Melolonthinae Pal.2 (1)
Hungary 3Melolonthinae Pal.17 (2)
Pal.18 (1)
Italy 1Melolonthinae Pal.2 (1)
Romania 8Melolonthinae Pal.9 (8)
Russian Federation 1Melolonthinae Pal.2 (1)
Turkey 16Melolonthinae (12) Pal.2+6+7
Pal.15 (1)
Pal.16 (2)
Pal.18 (1)
Anoxia pasiphae Reitter,1890   Greece 2Melolonthinae Pal.7 (2)
Anoxia pilosa (Fabricius,1792)   Austria 1Melolonthinae Pal.1 (1)
Germany 2Melolonthinae Pal.9 (1)
Pal.17 (1)
Hungary 2Melolonthinae Pal.1 (2)
Italy 2Melolonthinae Pal.18 (2)
Turkey 1Melolonthinae Pal.18 (1)
Unknown 6Melolonthinae Pal.17 (6)
Anoxia rattoi Escalera,1906   Morocco 1Melolonthinae Pal.17 (1)
Anoxia scutellaris rumelica Apfelbeck,1899   Bulgaria 10Melolonthinae (7) Pal.1+7
Pal.15 (1)
Pal.16 (2) as Anoxia rumelica
Anoxia spp.    Italy 2Melolonthinae Pal.8 (2)
Anoxia villosa (Fabricius,1781)   France 4Melolonthinae Pal.15 (4)
Germany 5Melolonthinae (3) Pal.1+11
Pal.17 (2)
Greece 1Melolonthinae Pal.7 (1)
Spain 2Melolonthinae Pal.1 (2)
Turkey 6Melolonthinae Pal.7 (6)
Unknown 3Melolonthinae Pal.1 (1)
Pal.17 (2)
Anthoplia floricola (Fabricius,1787)   Algeria 1Rutelinae Pal.3 (1)
Bosnia and Herzegovina 1Bosnia
Rutelinae Pal.8 (1)
Morocco 2Rutelinae Pal.8 (2)
Spain 8Rutelinae Pal.8 (8)
Spain 2Rutelinae (2) Pal.2 (as Anisoplia nigripennis Olivier)

Anthracophora crucifera (Olivier,1789)   India 5Cetoniinae Or.4 (5)
Sri Lanka 1Cetoniinae Or.3 (1) as ssp. ceylonensis Kraatz
Unknown 2Cetoniinae (2) Or.5
Anthracophora rusticola Burmeister,1842   China 3Cetoniinae Pal.17 (3)
Anticheira campisilvatica Ohaus,1905)   Trinidad and Tobago 3Trinidad
Rutelinae (3) S.Am.4
Anticheira virens (Drury,1773)   Colombia 2Rutelinae (2) S.Am.4
Unknown 1Rutelinae (1) S.Am.4
Aphodius abdominalis Bonelli,1812   Austria 28Aphodiinae (28) Pal.2+19 (as A. mixtus Villa)+11
France 1Aphodiinae Pal.16 (1)
Germany 1Aphodiinae Pal.19 (1) as A. mixtus Villa
Switzerland 22Aphodiinae Pal.7 (22)
Unknown 10Aphodiinae (10) Pal.2+19 (as A. mixtus Villa)

Aphodius abscissus Petrovitz,1958   Nigeria 1Aphodiinae Afr.16 (1) incl. 1 paratype
Aphodius adustus Klug,1855   Botswana 2Aphodiinae Afr.9 (2) as ab. porectus Schmidt
Namibia 2Aphodiinae Afr.23 (2)
Sierra Leone 2Aphodiinae Afr.15 (2) as ab. porectus Schmidt
South Africa 7Aphodiinae Afr.23 (3)
Afr.27 (4)
Tanzania 1Aphodiinae Afr.9 (1) as ab. porectus Schmidt
Aphodius affinis Panzer,1823   Unknown 3Aphodiinae Pal.20 (3)
Aphodius aleutus Eschscholtz,1822   United States 7Alaska
Aphodiinae N.Am.4 (7)
United States 2Arizona
Aphodiinae (2) N.Am.2 (as A.plutonicus Fall)

United States 1Oregon
Aphodiinae N.Am.1 (1)
United States 21Washington
Aphodiinae N.Am.1 (21)
Aphodius alpinus (Scopoli,1763)   Austria 20Aphodiinae (20) Pal.5+22
France 2Aphodiinae Pal.22 (2)
Hungary 2Aphodiinae Pal.22 (2)
Switzerland 32Aphodiinae (32) Pal.1+5+7+13
Unknown 14Aphodiinae Pal.22 (14)
Aphodius amblyodon Daniel,1900   Italy 3Aphodiinae Pal.11 (3)
Aphodius analis (Fabricius,1787)   China 3Aphodiinae (3) Pal.14
Aphodius anthracinus LeConte,1858   United States 2New Mexico
Aphodiinae N.Am.5 (2)
Aphodius ardens Harold,1866   Tanzania 2Aphodiinae (2) Afr.26
Aphodius arenarius (Olivier,1789)   Austria 1Aphodiinae (1) Pal.15 (as A. putridus (Geoffroy
1785))
Germany 1Aphodiinae Pal.18 (1)
Romania 2Aphodiinae (2) Pal.6 (as A. rhododactylus Marsh.)

Sweden 116  Skåne Blekinge Halland Småland Öland Västergötland
Alan Dufberg 47 (1)
Aphodiinae Main (115)
Syria 1Aphodiinae Pal.18 (1)
Unknown 4Aphodiinae Pal.18 (4)
Aphodius armatulus Fairmaire,1893   Somalia 1Aphodiinae (1) 1 syntype Afr.20
Aphodius asahinae ?   Japan 1Aphodiinae Pal.24 (1)
Aphodius asphaltinus Kolenati,1846   Turkey 3Aphodiinae Pal.11 (3)
Aphodius ater (DeGeer,1774)   Denmark 29Aphodiinae (29)
Finland 1Aphodiinae (1)
Germany 1Aphodiinae Pal.18 (1)
Norway 5Aphodiinae (5)
Russian Federation 1Aphodiinae Pal.6 (1)
Switzerland 1Aphodiinae Pal.18 (1)
United Kingdom 1Scotland
Aphodiinae Pal.10 (1)
Unknown 3Aphodiinae Pal.22 (3)
Aphodius bayeri Endrödi,1956   Gambia 1Aphodiinae (1) Afr.20
Aphodius biangulatus Fairmaire,1893   Senegal 2Aphodiinae (2) Afr.24
Sudan 1Aphodiinae (1) Afr.24
Aphodius bibatillatus Petrovitz,1958   South Africa 2Aphodiinae (2) Afr.19
Aphodius bicoloratus Schmidt,1908   Ghana 1Aphodiinae Afr.18 (1)
Aphodius biseriatus Schmidt,1908   Sri Lanka 1Aphodiinae Or.1 (1)
Aphodius bonnairei (Reitter,1892)   France 10Aphodiinae Pal.21 (10)
Aphodius bonvouloiri Harold,1860   Spain 2Aphodiinae (2) Pal.17
Aphodius bovis Endrödi,1960   Kenya 1Aphodiinae Afr.8 (1)
Aphodius brachysomus Solsky,1874   Russian Federation 1Aphodiinae Pal.15 (1)
Aphodius brasiliensis Castelnau,1840   Colombia 1Aphodiinae S.Am.1 (1)
Ecuador 4Aphodiinae S.Am.1 (4)
Aphodius brincki Landin,1959   Tanzania 2Aphodiinae Afr.20 (1)
Type Coleoptera (1) Paratype 03490:2 of Aphodius brincki Landin 1959
  Aphodius brincki Landin 1959

Aphodius brunneus Thunberg,1818   South Africa 2Aphodiinae (2) Afr.18
Aphodius brusewitzi (Landin,1974)   Ethiopia 1Type Coleoptera (1) Holotype 03487:1 of Aphodius brusewitzi Landin 1974
  Aphodius brusewitzi Landin 1974

Aphodius bucharicus Petrovitz,1961   Turkey 1Aphodiinae Pal.11 (1)
Turkmenistan 3Aphodiinae (3) Pal.11+16 (incl. 2 paratypes)

Aphodius burgeoni Paulian,1939   Congo 1Aphodiinae Afr.26 (1) syntype
Aphodius buxtoni Paulian,1939   Sudan 1Aphodiinae (1) Afr.24
Aphodius caffer Wiedemann,1823   Ethiopia 3Aphodiinae (3) Afr.9+19
South Africa 2Aphodiinae (2) Afr.19
Aphodius caffer ab.parasorex Landin   Ethiopia 4Aphodiinae Afr.9 (4)
South Africa 2Aphodiinae (2) Afr.19
Aphodius calcaratoides Paulian,1942   Zambeze river 1Aphodiinae (1) 1 paratype Afr.25
Aphodius calcaratus Boheman,1857   Namibia 10Aphodiinae (10) Afr.22+27
Aphodius calidus Harold,1871   Gambia 6Aphodiinae Afr.3 (6)
Aphodius capicola Peringuey,1901   South Africa 1Aphodiinae (1) Afr.24
Aphodius capitulatus Clouet,1898   Kenya 1Aphodiinae (1) Afr.20 (inc. 1 syntype)

Aphodius caspius Menetries,1832   Kyrgyzstan 6Aphodiinae Pal.21 (6)
Russian Federation 1Aphodiinae Pal.11 (1)
Aphodius cf. aleutus Eschscholtz,1822   Canada 1Labrador
Aphodiinae N.Am.6 (1)
Aphodius circumcinctus Schmidt,1840   Austria 1Aphodiinae Pal.20 (1)
Hungary 1Aphodiinae Pal.20 (1)
Aphodius circumductus Solsky,1876   Turkmenistan 1Aphodiinae Pal.11 (1)
Aphodius clavatus (Olivier,)   Spain 29Aphodiinae Pal.4 (29)
Aphodius clementi Peyerimhoff,1929   Chad 1Aphodiinae (1) Afr.24
Aphodius cognatus Fairmaire,1860   Afghanistan 29Aphodiinae (29) Pal.8+140 (as A.klugi Schnidt)

Egypt 1Aphodiinae (1) Afr.10 (as A.klugi Schnidt)

Morocco 1Aphodiinae Pal.11 (1)
Tunisia 3Aphodiinae Pal.11 (3)
Aphodius coloradensis Horn,1870   United States 2Arizona
Aphodiinae N.Am.5 (2)
Aphodius confinis Schmidt,1911   Ethiopia 1Aphodiinae (1) Afr.19
Aphodius congregatus Mannerheim,1853   Canada 2British Columbia
Aphodiinae N.Am.4 (2)
United States 6Alaska
Aphodiinae N.Am.4 (6)
Aphodius coniungatus (Panzer,1795)   Hungary 1Aphodiinae Pal.21 (1)
Italy 2Aphodiinae Pal.5 (2)
Spain 5Aphodiinae Pal.21 (5)
Caucasus, Ryssland / Georgien / Azerbajdzjan / Armenien / Iran / Turkiet 1Aphodiinae Pal.21 (1)
Unknown 1Aphodiinae Pal.21 (1)
Aphodius conspersus Horn,1887   United States 1California
Aphodiinae N.Am.2 (1)
Aphodius conspurcatus (Linnaeus,1758)   Denmark 6Aphodiinae (6)
Finland 1Aphodiinae (1)
Aphodius constans Duftschmid,1805   Italy 3Aphodiinae Pal.11 (3)
United Kingdom 1England
Aphodiinae (1) Pal.16
Unknown 1Aphodiinae Pal.22 (1)
Aphodius contaminatus (Herbst,1783)   Denmark 6Aphodiinae (6)
France 9Aphodiinae Pal.20 (9)
Greece 5Aphodiinae Pal.7 (5)
Italy 1Aphodiinae Pal.3 (1)
Italy 1Siciliy
Aphodiinae Pal.24 (1)
Sweden 337  Skåne Blekinge Småland Öland Gotland Östergötland Västergötland
Alan Dufberg 47 (26)
Aphodiinae Main (311)
United Kingdom 1Scotland
Aphodiinae Pal.3 (1)
Unknown 7Aphodiinae Pal.20 (7)
Aphodius contractus Klug,1845   Greece 2Aphodiinae Pal.4 (2)
Tunisia 1Aphodiinae (1) Pal.14
Aphodius convexus Erichson,1848   Switzerland 1Aphodiinae Pal.7 (1)
Aphodius copulatus Schmidt,1916   Tanzania 1Aphodiinae Afr.5 (1)
Aphodius corniculatus Boucomont,1933   Zambia 1Aphodiinae (1) Afr.24
Aphodius cornutus Wiedeman,1823   South Africa 9Aphodiinae (9) Afr.27
Aphodius corvinus (Erichson,1848)   Unknown 1Aphodiinae Pal.22 (1)
Aphodius cribrarius Brullé,1832   Greece 4Aphodiinae Pal.6 (4)
Italy 1Aphodiinae (1) Pal.16
Aphodius cruentus Klug,1855   Congo 11Aphodiinae (11) Afr.21
Tanzania 1Aphodiinae (1) Afr.21
Aphodius damarinus Kolbe,1908   Namibia 6Aphodiinae (6) Afr.21
South Africa 1Aphodiinae (1) Afr.21
Aphodius dentiger LeConte,1858   United States 1Texas
Aphodiinae N.Am.1 (1)
Aphodius depressus (Kugelann,1792)   Armenia 5Aphodiinae Pal.25 (5)
Austria 26Aphodiinae (23) Pal.2+4+10+16
Pal.27 (3)
Denmark 10Aphodiinae (10)
Germany 3Aphodiinae Pal.19 (3)
Italy 1Aphodiinae (1) Pal.16
Poland 1Aphodiinae Pal.16 (1)
Russian Federation 2Siberia
Aphodiinae Pal.19 (2)
Spain 1Aphodiinae Pal.4 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Aphodiinae Pal.10 (2)
Unknown 10Aphodiinae Pal.19 (10)
Aphodius diecki Harold,1870   Tunisia 4Aphodiinae Pal.6 (4)
Aphodius dimidiatus Roth,1851   Cameroon 1Aphodiinae Afr.9 (1)
Ethiopia 3Aphodiinae (3) Afr.9+19
Gambia 1Aphodiinae Afr.9 (1)
South Africa 2Aphodiinae (2) Afr.19
Zimbabwe 1Aphodiinae (1) Afr.19
Aphodius dionysii Clement,1958   Madagascar 1Aphodiinae (1) 1 paratype Afr.20
Aphodius distinctus (O.F.Müller,1776)   Austria 4Aphodiinae Pal.3 (4)
Finland 1Aphodiinae Main (1)
France 5Aphodiinae Pal.20 (5)
Spain 2Aphodiinae Pal.20 (2)
United States 1Aphodiinae (1) N.Am.
United States 1New York
Aphodiinae N.Am.5 (1)
United States 3Connecticut
Aphodiinae N.Am.5 (3)
Unknown 19Aphodiinae Pal.20 (19)
Aphodius ducorpsi Paulian,1942   Benin 1Aphodiinae (1) Afr.19 (incl. 1 paratype)
Aphodius ealaensis Endrödi, 1956   Congo, the democratic republic of the 1Aphodiinae Afr.26 (1) type of A. formosus var. ealaensis
Aphodius edgari Solsky,1876   Uzbekistan 1Aphodiinae (1) Pal.17
Aphodius elegans Allibert,1847   China 8Aphodiinae (6) Pal.11+21
Pal.28 (2)
Aphodius elisabethae Endrödi,1956   Congo 1Aphodiinae (1) Afr.16
Aphodius ellenbergeri Paulian,1942   South Africa 1Aphodiinae (1) 1 paratype Afr.21
Aphodius elongatulus Fabricius,1801   Unknown 2Aphodiinae Pal.22 (2) orig. name A. chinensis Harold
1861
Aphodius erraticus (Linnaeus,1758)   Afghanistan 3Aphodiinae (3) Pal.7+8
Armenia 9Aphodiinae Pal.25 (9)
Canada 1Ontario
Aphodiinae N.Am.1 (1)
Denmark 6Aphodiinae Main (6)
France 4Aphodiinae Pal.4 (4)
Germany 10Aphodiinae (10) Pal.14
Greece 14Aphodiinae (14) Pal.4+6
Italy 23Aphodiinae (23) Pal.3+4+10+14
Russian Federation 8Aphodiinae (6) Pal.4+14
Pal.24 (2)
Tunisia 9Aphodiinae Pal.10 (9)
Turkey 1Aphodiinae (1) Pal.14
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 4Aphodiinae (4) Pal.4+10
Uzbekistan 1Aphodiinae Pal.4 (1)
Aphodius esimoides Reitter,1893   Morocco 2Aphodiinae Pal.11 (2)
Tunisia 1Aphodiinae Pal.11 (1)
Unknown 1Aphodiinae Pal.18 (1)
Aphodius explanatus LeConte,1878   United States 13Alaska
Aphodiinae N.Am.4 (13)
Aphodius fasciatus (A.G. Olivier,1789)   France 5Aphodiinae Pal.23 (5) as A. uliginosus (Hardy
1847)
Uzbekistan 1Aphodiinae Pal.23 (1) as A. uliginosus (Hardy
1847)
Aphodius fastigiatus Schmidt,1908   Botswana 2Aphodiinae Afr.10 (2)
Tanzania 1Aphodiinae Afr.10 (1)
Aphodius feculentus Fairmaire,1892   Somalia 1Aphodiinae (1) Afr.24
Aphodius fimetarius (Linnaeus,1758)   Afghanistan 1Aphodiinae (1) Pal.7.
Armenia 2Aphodiinae Pal.25 (2)
Austria 11Aphodiinae (11) Pal.4+9
Bulgaria 1Aphodiinae Pal.29 (1)
Canada 6Ontario
Aphodiinae N.Am.1 (6)
China 1Aphodiinae Pal.22 (1)
Estonia 1Aphodiinae (1) Pal.15
Finland 2Aphodiinae (2)
France 9Aphodiinae Pal.4 (9)
Germany 20Aphodiinae (20) Pal.2+4+16+22
Greece 8Aphodiinae Pal.4 (7)
Pal.30 (1)
India 2Aphodiinae Pal.22 (2)
Italy 19Aphodiinae (19) Pal.3+4+9
Latvia 1Aphodiinae Pal.18 (1)
Norway 4Aphodiinae Main (4) Not checked
likely A. pedellus
Portugal 12Madeira
Aphodiinae Pal.4 (11)
Pal.27 (1)
Russian Federation 1Aphodiinae Pal.4 (1)
Spain 15Aphodiinae (15) Pal.2+4+9
Switzerland 12Aphodiinae Pal.2 (12)
Tunisia 1Aphodiinae Pal.4 (1)
Turkey 5Aphodiinae Pal.4 (4)
Pal.27 (1)
United Kingdom 1England
Aphodiinae Pal.2 (1)
United States 2Oregon
Aphodiinae N.Am.1 (2)
United States 2New York
Aphodiinae N.Am.5 (2)
United States 3Illinois
Aphodiinae N.Am.5 (3)
United States 3Washington
Aphodiinae N.Am.1 (3)
United States 2Connecticut
Aphodiinae N.Am.5 (2)
United States 3California
Aphodiinae N.Am.1 (3)
United States 1Arizona
Aphodiinae N.Am.1 (1)
United States 4Massachusetts
Aphodiinae N.Am.5 (4)
Unknown 7Aphodiinae Pal.22 (7)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Aphodiinae Pal.9 (2)
Uzbekistan 1Aphodiinae Pal.4 (1)
Aphodius finitimus Schmidt,1916   Tunisia 1Aphodiinae Pal.10 (1)
Aphodius flagrans Erichson,1843   South Africa 1Aphodiinae (1) Afr.21
Aphodius flavescens Balthasar,1961   Gambia 2Aphodiinae Afr.10 (2)
Aphodius flavus Endrödi,1955   Burundi 2Aphodiinae Afr.26 (2) paratypes
Gambia 4Aphodiinae Afr.5 (4)
Gambia 80Aphodiinae Afr.5 (80)
Sierra Leone 3Aphodiinae (3) Afr.15
Aphodius foetens (Fabricius,1787)   Austria 5Aphodiinae Pal.22 (5)
Denmark 4Aphodiinae (4)
France 1Aphodiinae (1) Pal.16
Germany 3Aphodiinae (3) Pal.15+22
Spain 8Aphodiinae (8) Pal.2+5
Sweden 338  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up Vr Gä Me Ån
Alan Dufberg 47 (13)
Aphodiinae Main (325)
Unknown 11Aphodiinae Pal.22 (11)
Aphodius foetidus (Herbst,1783)   Cyprus 1Aphodiinae Pal.4 (1)
France 8Aphodiinae Pal.22 (8)
Greece 1Crete
Aphodiinae Pal.30 (1)
Greece 8Aphodiinae Pal.11 (7)
Pal.30 (1)
Italy 4Aphodiinae Pal.3 (4)
Russian Federation 1Aphodiinae Pal.22 (1)
Saint Helena, Ascension and Tristan da Cunha 1Saint Helena
Aphodiinae Pal.18 (1)
Spain 5Mallorca
Aphodiinae Pal.24 (5)
Spain 25Aphodiinae (25) Pal.4+18 (as A. scybalarius)
Sweden 11 Skåne Gotska Sandön Västergötland Uppland
Aphodiinae Main (11)
Turkey 1Aphodiinae Pal.4 (1)
Unknown 1Aphodiinae Pal.22 (1)
Aphodius fossor (Linnaeus,1758)   Algeria 1Aphodiinae Pal.4 (1)
Austria 1Aphodiinae (1) Pal.14
Denmark 33Aphodiinae (33)
Germany 22Aphodiinae (22) Pal.13+14+16
Italy 4Aphodiinae (4) Pal.4+10+16
Sweden 425  Sk Bl Ha Sm Öl Go Ög Vg Bo Nä Sö Up Vr Dr Gä Hs
Alan Dufberg 47 (11)
Aphodiinae Main (414)
Turkey 1Aphodiinae Pal.4 (1)
United States 4Connecticut
Aphodiinae N.Am.4 (4)
United States 6New York
Aphodiinae N.Am.4 (6)
Unknown 6Aphodiinae (6) Pal.14
Turkestan, Kazakhstan / Uzbekistan / Kyrgyzstan / Tajikistan / Turkmenistan / Uzbekistan 1Aphodiinae (1) Pal.14
Aphodius foveiventris Raffray,1877   Gambia 23Aphodiinae Afr.8 (23)
Ghana 4Aphodiinae (4) Afr.18
Senegal 1Aphodiinae Afr.8 (1)
Aphodius frater Mulsant & Rey,1870   Georgia 1Aphodiinae Pal.18 (1)
Russian Federation 3Aphodiinae (3) Pal.15
Turkestan, Kazakhstan / Uzbekistan / Kyrgyzstan / Tajikistan / Turkmenistan / Uzbekistan 1Aphodiinae Pal.18 (1)
Aphodius freudei Endrödi,1964   Namibia 1Aphodiinae (1) 1 paratype Afr.23
Aphodius fucosus Schmidt,1922   United States 2California
Aphodiinae N.Am.5 (2)
Aphodius fumosulus Endrödi,1957   South Africa 11Aphodiinae Afr.8 (11)
Aphodius funestus Endrödi,1955   Kenya 1Aphodiinae Afr.2 (1)
Rwanda 1Aphodiinae Afr.26 (1) Paratype
Aphodius gallei Maf.   Costa Rica 3Aphodiinae S.Am.1 (3)
Aphodius gibbus Germar,1817   Austria 3Aphodiinae Pal.21 (1)
Pal.27 (2)
Switzerland 6Aphodiinae Pal.11 (6)
Aphodius gnomus Kozhantschikov,1916   Ethiopia 1Aphodiinae (1) Afr.19
Aphodius gnu Endrödi,1964   Congo 2Aphodiinae Afr.20 (2) Paratypes
Aphodius gorillae Bordat,1989   Kenya 6Aphodiinae Afr.5 (6)
Aphodius gracilis Boheman,1857   South Africa 3Aphodiinae (3) Afr.20+25
Aphodius granarius (Linnaeus,1767)   Afghanistan 1Aphodiinae Pal.8 (1)
Andorra 1Aphodiinae Pal.6 (1)
Canada 1British Columbia
Aphodiinae N.Am.4 (1)
China 1Aphodiinae (1) Pal.14
Egypt 1Aphodiinae Afr.9 (1)
France 2Aphodiinae (2) Pal.6+23
Germany 5Aphodiinae Pal.23 (5)
Italy 1Sicily
Aphodiinae Pal.24 (1)
Italy 7Aphodiinae (7) Pal.3+6+10
Morocco 4Aphodiinae (4) Pal.6+23
Netherlands 1Aphodiinae Pal.16 (1)
Oman 1Aphodiinae (1) Pal.7.
Portugal 10Madeira
Aphodiinae Pal.3 (10)
Russian Federation 1Aphodiinae Pal.23 (1)
Spain 7Mallorca
Aphodiinae Pal.24 (7)
Spain 32Aphodiinae (32) Pal.2+6+16+23
Syria 1Aphodiinae (1) Pal.15
Tunisia 6Aphodiinae Pal.6 (6)
Turkey 1Aphodiinae Pal.23 (1)
Ukraine 6Aphodiinae (6) Pal.15+23
United Kingdom 2England
Aphodiinae Pal.1 (2)
United States 10Connecticut
Aphodiinae N.Am.5 (10)
United States 6New York
Aphodiinae N.Am.5 (6)
United States 1California
Aphodiinae N.Am.2 (1)
Unknown 17Aphodiinae Pal.23 (17)
Aphodius gregarius Harold,1871   Afghanistan 3Aphodiinae (3) Pal.13
China 1Aphodiinae (1) Pal.13
Aphodius haafi Petrovitz,1961   Sri Lanka 8Aphodiinae Or.1 (8)
Aphodius haematicus Boheman,1857   Zambeze river 1Aphodiinae Afr.20 (1)
Aphodius haemorrhoidalis (Linnaeus,1758)   Canada 3Ontario
Aphodiinae N.Am.1 (3)
Finland 2Aphodiinae (2)
France 1Aphodiinae Pal.4 (1)
Germany 1Aphodiinae (1) Pal.13
Greece 1Aphodiinae Pal.4 (1)
Italy 26Aphodiinae (26) Pal.3+4+10
Norway 1Aphodiinae (1)
Poland 1Aphodiinae Pal.16 (1)
Russian Federation 1Aphodiinae (1) Pal.15
Spain 1Aphodiinae Pal.4 (1)
Sweden 403  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up Vr Dr Hs Vb Nb
Alan Dufberg 47 (20)
Aphodiinae Main (383)
Switzerland 1Aphodiinae Pal.16 (1)
Tunisia 1Aphodiinae Pal.4 (1)
United States 2Oregon
Aphodiinae N.Am.1 (2)
United States 7Connecticut
Aphodiinae (7) N.Am.4+5
United States 2California
Aphodiinae N.Am.1 (2)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 3Aphodiinae Pal.4 (3)
Unknown 1Aphodiinae Pal.24 (1)
Aphodius hamatus Say,1823   Canada 4Manitoba
Aphodiinae N.Am.4 (4)
Aphodius haroldianus Balthasar,1932   Japan 1Aphodiinae Pal.24 (1)
Aphodius hepaticolor Quedenfeldt,1884   Ethiopia 1Aphodiinae (1) Afr16
Senegal 21Aphodiinae Afr.8 (21)
Aphodius heynei Schmidt,1911   Tanzania 5Aphodiinae Afr.4 (5)
Tanzania 1Aphodiinae Afr.4 (1)
Aphodius hirticeps Peringuey,1901   South Africa 13Aphodiinae Afr.23 (13)
Aphodius howitti Hope,1847   Unknown 1Aphodiinae Pal.18 (1)
Aphodius humilis Roth,1851   Ghana 47Aphodiinae Afr.18 (47)
Kenya 1Aphodiinae (1) Afr.27
South Africa 1Aphodiinae Afr.27 (1)
Sri Lanka 2Aphodiinae Or.1 (2)
Tanzania 1Aphodiinae (1) Afr.21
Aphodius hyalinus Petrovitz,1961   Afghanistan 1Type Coleoptera (1) Holotype 00926:1 of Aphodius hyalinus Petrovitz 1961
  Aphodius hyalinus Petrovitz 1961

Aphodius ibericus Harold,1874   Morocco 1Aphodiinae (1) Pal.17
Aphodius immaturus Mulsant,1842   Spain 17Aphodiinae Pal.6 (17) as A. sericatus M.&R.
Aphodius infuscatopennis Schmidt,1909   Chile 1Aphodiinae S.Am.1 (1)
Aphodius inouei Nomura,1942   Japan 1Aphodiinae Pal.24 (1)
Aphodius insularis Petrovitz,1961   Sri Lanka 8Aphodiinae Or.1 (8)
Thailand 12Aphodiinae Or.3 (12)
Aphodius irregularis Westwood,1839   Nepal 1Aphodiinae Or.3 (1)
Aphodius isaburoi Nakane,1956   Japan 4Aphodiinae (4) Pal.17+24
Aphodius janssensi Paulian,1939   Congo 2Aphodiinae (2) Afr.19 ( 1 spec. labeled as Type)

Aphodius jeannelianus Balthasar,1945   Kenya 2Aphodiinae (2) Afr.20+24 (incl. paratype of Podotenoides jeanneli Paulian
1942)

Aphodius johnsoni Baraud,1976   Italy 2Aphodiinae Pal.11 (2)
Aphodius jugalis Péringuey,1901   Cameroon 1Aphodiinae Afr.10 (1)
Aphodius kachovskyi Koshantschikoff,1917   Ethiopia 168Aphodiinae (166) Afr.7+8
Afr.27 (2)
Aphodius kaffa Petrovitz,1964   Ethiopia 5Aphodiinae Afr.8 (5)
Aphodius kalungwensis (Paulian,1954)   Congo 2Aphodiinae (2) 2 paratypes Afr.19
Aphodius kanemicus Endrödi,1976   Tanzania 1Aphodiinae (1) Afr.16
Aphodius kindianus Petrovitz,1958   Guinea 1Aphodiinae (1) Afr.16 (incl. 1 paratype)

Aphodius klugi Schmidt,1910   Kazakhstan 1Aphodiinae Pal.18 (1)
Tunisia 1Aphodiinae Pal.18 (1)
Aphodius koshantschikovi Jacobson,1911   Cyprus 5Aphodiinae Pal.8 (5)
Aphodius lapponum Gyllenhal,1808   Iceland 69Aphodiinae (69)
Aphodius latecinctus Fairmaire,1903   Unknown 2Aphodiinae (2 ) Afr.24 (paratypes?)

Aphodius latus coryndoni Paulian,1942   Kenya 3Aphodiinae (3) Afr.1+19
Aphodius lentus Horn,1870   United States 1Indiana
Aphodiinae N.Am.5 (1)
Aphodius leo Paulian,1942   Kenya 1Aphodiinae (1) Afr.19 (incl. 1 paratype)

Aphodius leucopterus Klug,1845   Egypt 1Aphodiinae Afr.10 (1)
Aphodius lewisi Waterhouse,1875   Sri Lanka 4Aphodiinae Or.2 (4)
Aphodius libanonensis Petrovitz,1958   Turkey 1Aphodiinae Pal.11 (1)
Aphodius liguricus Daniel,1902.   Italy 1Aphodiinae Pal.11 (1)
Aphodius lividus (Olivier,1789)   Afghanistan 1Aphodiinae (1) Pal.7.
Australia 13Western Australia
Aphodiinae Pac.1 (13)
Bolivia 1Aphodiinae S.Am.1 (1)
Cyprus 1Aphodiinae Pal.16 (1)
Egypt 2Aphodiinae Afr.3 (2)
Ethiopia 1Aphodiinae Afr.3 (1)
Gambia 10Aphodiinae Afr.3 (10)
Guatemala 2Aphodiinae S.Am.1 (2)
India 2Aphodiinae Pal.23 (2)
Italy 2Aphodiinae Pal.10 (2)
Kazakhstan 1Aphodiinae Pal.23 (1)
Kenya 1Aphodiinae Afr.3 (1)
Kyrgyzstan 1Aphodiinae (1) Pal.16
Lesotho 1Aphodiinae (1) Afr.23
Libya 2Aphodiinae (2) Pal.15
Mexico 1Aphodiinae N.Am.1 (1)
Namibia 4Aphodiinae (4) Afr.23
Oman 4Aphodiinae (4) Pal.7.
Peru 1Aphodiinae S.Am.1 (1)
Poland 2Aphodiinae (2) Pal.16 (locality: Rapa)
Portugal 22Madeira
Aphodiinae Pal.4 (22)
Senegal 4Aphodiinae Afr.3 (4)
Sierra Leone 17Aphodiinae (17) Afr.15
South Africa 42Aphodiinae (42) Afr.23+24+25
Spain 19Aphodiinae Pal.6 (19)
Sudan 2Aphodiinae (2) Afr.26
Tunisia 1Aphodiinae Pal.6 (1)
United States 1Hawaii
Aphodiinae N.Am.1 (1)
United States 1Texas
Aphodiinae N.Am.1 (1)
United States 6California
Aphodiinae N.Am.1 (6)
Unknown 16Aphodiinae (16) Pal.16+23
Uzbekistan 4Aphodiinae (4) Pal.15+16+23
Zimbabwe 1Aphodiinae (1) Afr.23
Aphodius longispina Küster,1854   Italy 3Aphodiinae Pal.3 (3)
Spain 1Aphodiinae Pal.22 (1)
Aphodius longus Schmidt,1912   Kenya 1Aphodiinae (1) Afr.21
Aphodius lucidulus Boheman,1857   Lesotho 1Aphodiinae Afr.21 (1)
South Africa 3Aphodiinae Afr.25 (3)
Aphodius lugens Creutzer,1799   Greece 7Aphodiinae Pal.5 (7)
Russian Federation 1Aphodiinae Pal.23 (1)
Slovenia 1Aphodiinae (1) Pal.15
Spain 2Aphodiinae (2) Pal.5+23
Syria 1Aphodiinae Pal.23 (1)
Tunisia 5Aphodiinae Pal.11 (5)
Turkey 6Aphodiinae (4) Pal.13+23
Pal.27 (2)
Aphodius lunifer Solsky,1876   Turkestan, Kazakhstan / Uzbekistan / Kyrgyzstan / Tajikistan / Turkmenistan / Uzbekistan 2Aphodiinae (2) Pal.18
Uzbekistan 1Aphodiinae (1) Pal.17
Aphodius luridus (Fabricius,1775)   Armenia 10Aphodiinae Pal.25 (10)
Denmark 1Aphodiinae (1)
France 8Aphodiinae Pal.5 (8)
Germany 4Aphodiinae (4) Pal.15+19
Greece 1Aphodiinae Pal.19 (1)
Italy 1Aphodiinae Pal.19 (1)
Romania 5Aphodiinae (5) Pal.5+19
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 13Aphodiinae Pal.10 (13)
Unknown 24Aphodiinae Pal.19 (24)
Uzbekistan 5Aphodiinae (5) Pal.5+16+19
Aphodius lusingae (Paulian,1954)   Congo 1Aphodiinae (1) 1 paratype Afr.21
Aphodius lutulentus Haldeman,1843   United States 1South Carolina
Aphodiinae N.Am.5 (1)
United States 4Aphodiinae N.Am.5 (4)
Aphodius maculatus Sturm,1800   Austria 1Aphodiinae Pal.6 (1)
Aphodius madara Nakane,1960   Japan 2Aphodiinae (2) Pal.17
Aphodius magnicornis Bordat,1992   Kenya 4Aphodiinae Afr.1 (4)
Aphodius magnificus Schmidt,1909   Tanzania 1Aphodiinae (1) Afr.24
Aphodius mara  Krikken & Kaas,1984   Kenya 2Aphodiinae (2) 2 paratypes Afr.20
Aphodius marshalli Paulian,1942   South Africa 1Aphodiinae (1) 1 paratype Afr.24
Aphodius medioxinus Peringuey,1901   Namibia 1Aphodiinae (1) Afr.23
Aphodius menetriesi Menetries,1849   Afghanistan 1Aphodiinae Pal.8 (1)
Aphodius merdarius (Fabricius,1775)   Denmark 1Aphodiinae (1)
Estonia 5Aphodiinae (5) Pal.15
Finland 1Aphodiinae Main (1)
Germany 5Aphodiinae (5) Pal.13+21
Greece 1Aphodiinae Pal.6 (1)
Italy 2Aphodiinae (2) Pal.6+24
Sweden 171  Sk Bl Ha Sm Öl Go Ög Vg Bo Ds Up Dr Me Jä Ån
Alan Dufberg 47 (1)
Aphodiinae Main (170)
Unknown 10Aphodiinae Pal.21 (10)
Aphodius meruanus Balthasar,1967   Kenya 2Aphodiinae Afr.5 (2)
Aphodius militaris LeConte,1858   United States 1California
Aphodiinae N.Am.5 (1)
Aphodius mime Paulian,1942   Kenya 1Aphodiinae (1) Afr.19
Aphodius minusculus Fairmaire,1893   Kenya 1Aphodiinae (1) Afr.20
Aphodius mizo Nakane,1967   Japan 3Aphodiinae (3) Pal.17
Aphodius montana Clement,1958   Guinea 1Aphodiinae (1) 1 paratype? Afr.23
Aphodius montanus Erichson,1848   Italy 1Aphodiinae Pal.18 (1)
Aphodius montivagus Erichson,1848   Italy 19Aphodiinae Pal.7 (19)
Aphodius montuosus Schmidt,1911   Ethiopia 1Aphodiinae (1) Afr.19
Kenya 1Aphodiinae (1) Afr.27
South Africa 1Aphodiinae (1) Afr.19
Aphodius moquinus Fall,1907   United States 1Arizona
Aphodiinae N.Am.1 (1)
Aphodius mortuorum Balthasar,1932   China 1Aphodiinae (1) Pal.17
Aphodius murati Paulian,1939   Senegal 2Aphodiinae (2) Afr.24
Aphodius nemoralis Erichson,1848   Austria 1Aphodiinae Pal.22 (1)
Finland 1Aphodiinae Main (1)
Germany 1Aphodiinae Pal.6 (1)
Aphodius neohaematicus Landin,1967   Congo 10Aphodiinae (5) Afr.20+25
Type Landin (5) Paratype 03488:2-6 of Aphodius neohaematicus Landin 1967
  Aphodius neohaematicus Landin 1967

Gambia 9Aphodiinae Afr.10 (9)
Ghana 1Aphodiinae Afr.17 (1)
Madagascar 1Aphodiinae (1) Afr.20 (incl. type 3488:6)

Mozambique 1Aphodiinae (1) Afr.20
Aphodius neotomae Fall,1907   United States 2California
Aphodiinae N.Am.5 (2)
Aphodius nevadensis Horn,1870   United States 2Nevada
Aphodiinae N.Am.5 (2)
Aphodius nigrivittis Solsky,1876   Afghanistan 9Aphodiinae Pal.8 (9)
Kazakhstan 4Aphodiinae Pal.20 (4)
Aphodius nigrovirgatus Schmidt,1911   China 1Aphodiinae Pal.19 (1)
Aphodius novus Schmidt,1911   Ghana 1Aphodiinae Afr.18 (1)
Tanzania 1Aphodiinae Afr.10 (1)
Aphodius obliteratus (Panzer,1823)   Austria 1Aphodiinae (1) Pal.15
Bosnia and Herzegovina 1Bosnia
Aphodiinae (1) Pal.15
Greece 1Aphodiinae Pal.16 (1)
Italy 6Aphodiinae (6) Pal.11+15+17
Spain 1Aphodiinae Pal.20 (1)
Ukraine 1Aphodiinae Pal.16 (1)
United Kingdom 2England
Aphodiinae (2) Pal.15
Caucasus, Ryssland / Georgien / Azerbajdzjan / Armenien / Iran / Turkiet 1Aphodiinae Pal.20 (1)
Aphodius oblongus Say,1823   United States 2Arizona
Aphodiinae N.Am.5 (2)
United States 2New York
Aphodiinae N.Am.5 (2)
Aphodius obscurus (Fabricius,1792)   Armenia 1Aphodiinae Pal.25 (1)
Austria 4Aphodiinae (4) Pal.1+6
France 1Aphodiinae Pal.21 (1)
Greece 2Aphodiinae (2) Pal.2+21
Greece 31Aphodiinae (31) Pal.5 (as A.flavipennis Mill.)

Spain 1Aphodiinae Pal.2 (1)
Switzerland 4Aphodiinae Pal.2 (4)
Turkey 2Aphodiinae Pal.21 (1)
Pal.25 (1)
Unknown 15Aphodiinae Pal.21 (15)
Aphodius obscurus latinus Dellacasa,1983   Italy 1Aphodiinae (1) Pal.17
Aphodius okadoi ?   Japan 1Aphodiinae Pal.24 (1)
Aphodius opacus LeConte,1872   Canada 14British Columbia
Aphodiinae N.Am.1 (14)
Aphodius orinus Koshantschikov,1912   India 1Aphodiinae Or.2 (1)
Aphodius pallicolor Fairmaire,1886   Unknown 1Aphodiinae (1 ) Afr.20
Aphodius pallidigonis Waterhouse,1875   Japan 3Aphodiinae (3) Pal.17+24
Aphodius partitus Endrödi,1956   Sierra Leone 4Aphodiinae (4) Afr.15
Aphodius pavo Balthasar,1939   Tanzania 3Aphodiinae Afr.8 (3)
Aphodius pectoralis LeConte,1857   United States 1Oregon
Aphodiinae N.Am.1 (1)
Aphodius pedellus (DeGeer,1774)   Denmark 32Aphodiinae Main (32) Not checked
could be A. fimetarius
Spain 1Aphodiinae (1) Pal.17
Sweden 946  Sk Bl Ha Sm Öl Go GS Ög Vg Bo Ds Nä Sö Up Vs Vr Dr Gä Hs Me Hr Jä Ån Vb Nb Ås Ly Pi Lu To
Alan Dufberg 47 (13) as A. fimetarius (L.)
Aphodiinae Main (933) as A. fimetarius (L.)
Aphodius perblandus Balthasar,1933   China 1Aphodiinae (1) Pal.17
Aphodius peruanus Erichson,1833   Bolivia 9Aphodiinae Pal.20 (5)
S.Am.2 (4)
Aphodius phaeopterus LeConte,1878   Canada 1British Columbia
Aphodiinae N.Am.1 (1)
United States 3Oregon
Aphodiinae N.Am.5 (3)
Aphodius phalacronotoides Balthasar,1960   Gambia 92Aphodiinae Afr.10 (92)
Aphodius piceus Gyllenhal,1808   Unknown 3Aphodiinae Pal.22 (3)
Aphodius pictipennis Fairmaire,1897   Madagascar 1Aphodiinae (1) Afr.24
Aphodius pilosellus Schmidt,1907   Kenya 1Aphodiinae (1) Afr.20
Aphodius plagiatus/niger    Sweden 16 Skåne Öland Gotland
Aphodiinae Main (16)
Aphodius plasoni Käufel,1914   China 1Aphodiinae (1) Pal.17
Aphodius porcus (Fabricius,1792)   Denmark 2Aphodiinae (2)
Germany 3Aphodiinae (3) Pal.15
Sweden 40  Skåne Blekinge Öland
Alan Dufberg 47 (6)
Aphodiinae Main (34)
Unknown 5Aphodiinae Pal.21 (5)
Aphodius praecox Erichson,1848   Austria 5Aphodiinae Pal.19 (5)
Unknown 3Aphodiinae (3) Pal.2+19
Aphodius principalis Harold,1861   South Africa 2Aphodiinae (2) Afr.19
Aphodius prodromus (Brahm,1790)   Austria 1Aphodiinae Pal.20 (1)
Finland 4Aphodiinae Main (4)
France 2Aphodiinae Pal.20 (2)
Germany 10Aphodiinae (10) Pal.16+20
Italy 1Aphodiinae Pal.7 (1)
Italy 1Sicily
Aphodiinae Pal.24 (1)
Kazakhstan 3Aphodiinae Pal.20 (3)
Switzerland 2Aphodiinae Pal.7 (2)
Tanzania 1Aphodiinae Afr.9 (1)
Turkey 1Aphodiinae Pal.7 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 4Aphodiinae Pal.10 (4)
Unknown 15Aphodiinae Pal.20 (15)
Aphodius productus Boheman,1857   South Africa 1Aphodiinae (1) Afr.21
Aphodius pruinosus Reitter,1892   Kazakhstan 3Aphodiinae Pal.18 (3)
Turkmenistan 1Aphodiinae Pal.18 (1)
Turkmenistan 1Aphodiinae Pal.11 (1)
Aphodius przewalskyi Reitter,1887   China 53Aphodiinae Pal.13 (53)
Aphodius pseudocalcaratus Paulian,1952   South Africa 2Aphodiinae (2) Afr.16 (incl. 2 paratypes)


Aphodius pseudocoenosus     Greece 1Aphodiinae (1) Pal.17
Aphodius pseudohumilis Bordat,1989   South Africa 16Aphodiinae Afr.8 (16)
Aphodius puncticeps Paulian,1942   Kenya 1Aphodiinae (1 ) Afr.20
Aphodius punctipennis Erichson,1848   Afghanistan 59Aphodiinae (59) Pal.7+8+13
China 1Aphodiinae (1) Pal.13
Kyrgyzstan 2Aphodiinae Pal.22 (2)
Turkey 2Aphodiinae Pal.6 (2)
Aphodius pusillus (Herbst,1789)   Armenia 13Aphodiinae Pal.25 (13)
Finland 5Aphodiinae Main (5)
France 1Aphodiinae Pal.6 (1)
Germany 1Aphodiinae Pal.21 (1)
Kazakhstan 1Aphodiinae Pal.21 (1)
Norway 1Aphodiinae (1)
Romania 9Aphodiinae (9) Pal.2+6
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Aphodiinae Pal.10 (1)
Unknown 13Aphodiinae Pal.21 (13)
Aphodius pusio (Kolbe,1908)   Namibia 1Aphodiinae (1) Afr.18
Aphodius quadriguttatus (Herbst,1793)   Armenia 1Aphodiinae Pal.25 (1)
Bulgaria 2Aphodiinae Pal.6 (1)
Pal.29 (1)
Cyprus 2Aphodiinae Pal.6 (2)
Germany 1Aphodiinae Pal.19 (1)
Greece 4Aphodiinae (4) Pal.6+16
Spain 2Aphodiinae Pal.6 (2)
Sweden 221  Skåne
Alan Dufberg 47 (14)
Aphodiinae Main (207)
Turkey 5Aphodiinae (5) Pal.6+19
Unknown 4Aphodiinae Pal.19 (4)
Uzbekistan 3Aphodiinae (3) Pal.15+19
Aphodius quadrinaevulus Reitter,1892   Armenia 1Aphodiinae Pal.25 (1)
Greece 1Aphodiinae Pal.30 (1)
Turkey 1Aphodiinae Pal.30 (1)
Aphodius ragusae Reitter,1892   Italy 3Aphodiinae (3) Pal.18
Aphodius rangoonensis Petrovitz,1970   Nepal 6Aphodiinae Or.3 (6)
Aphodius reichei Harold,1859   China 5Aphodiinae Pal.19 (5)
Myanmar 1Aphodiinae Pal.19 (1)
Thailand 2Aphodiinae Or.2 (2)
Unknown 1Aphodiinae Pal.19 (1)
Aphodius rhinocerus mirus Schmidt,1916   Ethiopia 6Aphodiinae Afr.9 (6)
Aphodius rotschildi Schmidt,1911   Ethiopia 1Aphodiinae Afr.27 (1)
Rwanda 2Aphodiinae Afr.26 (2) Paratypes of Blackburnius africanus Endrödi
Aphodius ruandanus Endrödi,1955   Rwanda 2Aphodiinae (2) Afr.26 (incl. 2 paratypes)

Aphodius ruficlarus Fall,1932   United States 1Arizona
Aphodiinae N.Am.1 (1)
Aphodius rufipes (Linnaeus,1758)   Austria 6Aphodiinae (6) Pal.2+5+10+15
Finland 2Aphodiinae (2)
Germany 6Aphodiinae (6) Pal.13+19
Norway 7Aphodiinae (7)
Romania 1Aphodiinae Pal.19 (1)
Spain 2Aphodiinae Pal.5 (2)
Switzerland 4Aphodiinae (3) Pal.5+18
Pal.24 (1)
Tunisia 1Aphodiinae Pal.5 (1)
Unknown 3Aphodiinae Pal.19 (3)
Aphodius rufus (Moll,1788)   France 1Aphodiinae (1) Pal.16 (as A. scybalarius F.)

Germany 8Aphodiinae Pal.23 (8)
Greece 1Aphodiinae Pal.23 (1)
Hungary 1Aphodiinae Pal.23 (1)
Italy 2Aphodiinae Pal.23 (2)
Spain 10Aphodiinae (10) Pal.2 (as A. scybalarius F.)

Switzerland 3Aphodiinae Pal.18 (3)
Aphodius ruricola Melsheimer,1844   United States 2Connecticut
Aphodiinae N.Am.6 (2)
Aphodius rusicola Melsheimer,1845   Canada 3Newfoundland
Aphodiinae N.Am.4 (3)
Aphodius sareptanus Balthasar,1935   Russian Federation 7Aphodiinae Pal.6 (7)
Aphodius satellitius (Herbst,1789)   Armenia 1Aphodiinae Pal.25 (1)
Austria 1Aphodiinae Pal.19 (1)
Cyprus 1Aphodiinae Pal.6 (1)
France 1Aphodiinae Pal.19 (1)
Greece 1Aphodiinae Pal.19 (1)
Italy 2Aphodiinae (2) Pal.18
Unknown 6Aphodiinae Pal.19 (6)
Aphodius satyrus Reitter,1892   Austria 1Aphodiinae Pal.10 (1)
Italy 1Aphodiinae Pal.11 (1)
Unknown 1Aphodiinae Pal.22 (1)
Aphodius schaumi Harold,1859   Kenya 5Aphodiinae Afr.6 (5)
Tanzania 174Aphodiinae (174) Afr.6+7
Aphodius schlumbergeri samniticus Daniel,1902   Italy 4Aphodiinae (4) Pal.18
Aphodius schoutedeni Boucomont,1928   Angola 1Aphodiinae (1) Afr.24
Congo 4Aphodiinae (4) Afr.25 (incl. 4 syntypes)

Aphodius schusteri Balthasar,1935   Ghana 7Aphodiinae (7) Afr.18
British Africa 1Aphodiinae (1) Afr.19
Aphodius scrofa (Fabricius,1787)   Germany 2Aphodiinae (2) Pal.16+21
Greece 1Aphodiinae Pal.6 (1)
Italy 3Aphodiinae (3) Pal.10+15
Sweden 152  Skåne Öland
Alan Dufberg 47 (23)
Aphodiinae Main (129)
Unknown 6Aphodiinae Pal.21 (6)
Aphodius scrutator (Herbst,1789)   Armenia 5Aphodiinae Pal.25 (5)
Austria 11Aphodiinae (11) Pal.14
Bosnia and Herzegovina 1Bosnia
Aphodiinae (1) Pal.16
Italy 3Aphodiinae (3) Pal.14
Ukraine 1Aphodiinae Pal.4 (1)
Unknown 3Aphodiinae (3) Pal.14
Aphodius senegalensis Klug,1835   Benin 1Aphodiinae (1) Afr.19
Gambia 17Aphodiinae Afr.1 (17)
Nigeria 1Aphodiinae Afr.28 (1)
Senegal 12Aphodiinae Afr.1 (12)
Aphodius septemmaculatus (Fabricius,1781)   South Africa 2Aphodiinae (2) Afr.20
Aphodius sexmaculosus Schmidt,1911   Ghana 2Aphodiinae (2) Afr.18
Guinea 2Aphodiinae (2) Afr.23
Aphodius shantungensis Balthasar,1941   Thailand 5Aphodiinae Or.2 (5)
Aphodius sigwalti Bordat,1989   Gambia 1Aphodiinae Afr.5 (1)
Aphodius simoni Petrovitz,1959   Botswana 3Aphodiinae Afr.8 (3)
Aphodius solitarius Petrovitz,1961   Afghanistan 1Type Coleoptera (1) Holotype 00927:1 of Aphodius solitarius Petrovitz 1961
  Aphodius solitarius Petrovitz 1961

Aphodius sordidus (Fabricius,1775)   Denmark 4Aphodiinae (4)
Estonia 4Aphodiinae (4) Pal.15
Germany 1Aphodiinae Pal.23 (1)
United Kingdom 8Scotland
Aphodiinae Pal.9 (8)
Aphodius soudanensis Petrovitz,1958   Sudan 1Aphodiinae (1) Afr.19
Aphodius sp.1    South Africa 3Aphodiinae Afr.27 (3) as A. rudebecki Landin
manuscript name
Aphodius sp.2    South Africa 1Aphodiinae Afr.27 (1) as A. addonis Landin
manuscript name
Aphodius sp.3    South Africa 1Aphodiinae Afr.27 (1) as A. skukuzaensis Landin
manuscript name
Aphodius sp.4    South Africa 2Aphodiinae Afr.27 (2) as A. colasi Landin
manuscript name
Aphodius sparsus LeConte,1878   United States 2California
Aphodiinae N.Am.5 (2)
Aphodius sphacelatus (Panzer,1798)   Andorra 6Aphodiinae Pal.10 (6)
Bosnia and Herzegovina 1Aphodiinae Pal.20 (1)
Bulgaria 2Aphodiinae Pal.6 (2)
France 12Aphodiinae Pal.10 (12)
Italy 4Aphodiinae Pal.6 (4)
Spain 1Aphodiinae Pal.6 (1)
United Kingdom 1England
Aphodiinae Pal.18 (1)
Aphodius spp.    Botswana 4Aphodiinae Afr.8 (4)
Congo 1Aphodiinae (1) Afr.26
Ethiopia 1Aphodiinae Afr.10 (1)
Faroe Islands 1Aphodiinae Main (1)
Kenya 5Aphodiinae (5) Afr.27
Kenya 1Aphodiinae Afr.4 (1)
Kenya 1Aphodiinae Afr.8 (1)
Kenya 1Aphodiinae Afr.10 (1)
Mexico 2Aphodiinae N.Am.2 (2)
Nigeria 1Aphodiinae Afr.28 (1)
Senegal 1Aphodiinae Afr.10 (1)
South Africa 2Aphodiinae (2) Afr.27
Aphodius sticticus (Panzer,1798)   Armenia 6Aphodiinae Pal.25 (6)
Denmark 1Aphodiinae (1)
France 1Aphodiinae (1) Pal.13
Hungary 4Aphodiinae Pal.20 (4)
Italy 5Aphodiinae Pal.10 (5)
Romania 18Aphodiinae (18) Pal.3+6 (as A.equestris Panz.)

Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Aphodiinae Pal.10 (1)
Unknown 4Aphodiinae Pal.20 (4)
Aphodius strigilatus Roth,1851   Ethiopia 7Aphodiinae Afr.4 (5)
Afr.27 (2)
Aphodius sturmi Harold,1870   Bulgaria 4Aphodiinae (4) Pal.5+10
Germany 1Aphodiinae Pal.22 (1)
Greece 33Aphodiinae Pal.5 (33)
Italy 35Aphodiinae (35) Pal.5+10
Spain 19Aphodiinae Pal.5 (19)
Tunisia 14Aphodiinae Pal.5 (14)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 7Aphodiinae Pal.5 (7)
Aphodius suarius Faldermann,1835   Bulgaria 1Aphodiinae Pal.6 (1)
Greece 4Aphodiinae Pal.6 (4)
Turkey 2Aphodiinae Pal.27 (2)
Aphodius sublineatus ?   Japan 1Aphodiinae Pal.24 (1) = sublimbatus?
Aphodius subopacus Petrovitz,1964   Gambia 7Aphodiinae Afr.9 (7)
Aphodius subterraneus (Linnaeus,1758)   Afghanistan 1Aphodiinae (1) Pal.14
Austria 5Aphodiinae (5) Pal.x
Denmark 1Aphodiinae (1)
France 11Aphodiinae (11) Pal.14
Germany 5Aphodiinae (5) Pal.14+16
Italy 5Aphodiinae Pal.4 (5)
Kyrgyzstan 2Aphodiinae (2) Pal.14
Russian Federation 7Aphodiinae (7) Pal.14
Sweden 59 Skåne Halland Öland Östergötland Närke Södermanland Uppland Dalarna
Aphodiinae Main (59)
Turkey 3Aphodiinae Pal.4 (3)
Aphodius subteter Petrovitz,1962   Tanzania 2Aphodiinae Afr.8 (2)
Aphodius sulcatus F. (?)   Unknown 1Aphodiinae Pal.21 (1)
Aphodius syriacus Mulsant & Rey,1870   Syria 3Aphodiinae Pal.19 (3)
Aphodius szekeeyi Endrödi,1960   Congo 1Aphodiinae (1) Afr.19 (incl. 1 paratype)

Aphodius testaceicolor Balthasar,1937   Mozambique 1Aphodiinae Afr.20 (1)
Aphodius tetricus Harold,1871   Ethiopia 1Aphodiinae Afr.3 (1)
Aphodius thermicola Sturm,1800   Greece 1Aphodiinae Pal.6 (1)
Unknown 1Illyr.
Aphodiinae Pal.21 (1)
Unknown 7Aphodiinae Pal.21 (7)
Aphodius thomomysi Brown,1928   United States 1Arizona
Aphodiinae N.Am.2 (1)
Aphodius tomentosus (Müller,1776)   Denmark 1Aphodiinae (1)
Finland 1Aphodiinae (1)
Sweden 11 Skåne
Aphodiinae Main (11)
Aphodius tragicus (Schmidt,1916)   Ethiopia 14Aphodiinae Afr.5 (12)
Afr.27 (2)
Aphodius translusidus Petrovitz,1961   Afghanistan 2Aphodiinae Pal.8 (2)
Aphodius vandykei Barret,1931   United States 2California
Aphodiinae N.Am.2 (2)
Aphodius varians Duftschmid,1805   Austria 1Aphodiinae Pal.23 (1)
Croatia 1Aphodiinae Pal.23 (1)
France 1Aphodiinae Pal.16 (1)
Greece 20Aphodiinae (20) Pal.5+16
Hungary 1Aphodiinae (1) Pal.15
Italy 3Aphodiinae Pal.5 (3)
Kazakhstan 1Aphodiinae Pal.23 (1)
Russian Federation 2Aphodiinae (2) Pal.6+23
Spain 1Aphodiinae Pal.5 (1)
Unknown 11Aphodiinae (11) Pal.16+23
Aphodius variolosa Clement,1958   Guinea 1Aphodiinae Afr.23 (1) Paratype
Aphodius venaloides Endrödi,1960   Kenya 1Aphodiinae (1) Afr.24
Rwanda 3Aphodiinae (3) Afr.26
Aphodius vitellinus Klug,1845   Tunisia 1Aphodiinae Pal.11 (1)
Aphodius volkonskyi Paulian,1942   Unknown 1Aphodiinae (1) 1 cotype Afr.21
Aphodius weidholzi Petrovitz,1964   Gambia 4Aphodiinae Afr.9 (4)
Senegal 14Aphodiinae Afr.9 (14)
Aphodius wichei Petrovitz,1961   Sri Lanka 7Aphodiinae Or.1 (7)
Aphodius zborniki Balthasar,1935   Congo 4Aphodiinae (4) Afr.25
Aphodius zenkeri Germar,1813   Denmark 2Aphodiinae (2)
France 10Aphodiinae Pal.19 (10)
Germany 2Aphodiinae Pal.19 (2)
Italy 1Aphodiinae (1) Pal.18
Sweden 157  Skåne Blekinge Gotland
Alan Dufberg 47 (12)
Aphodiinae Main (145)
United Kingdom 4England
Aphodiinae (4) Pal.15
Aphodobius transvaalicus (Petrovitz,1964)   Namibia 6Aphodiinae Afr.35 (6) as Nobius
South Africa 1Aphodiinae Afr.35 (1) as Nobius
Aphodoharmogaster costipennis (Boheman,1857)   South Africa 2Aphodiinae Afr.31 (2)
Aphonus castaneus (Melsheimer,1846)   United States 5Connecticut
Dynastinae (5) N.Am.2+6
Unknown 2Dynastinae N.Am.2 (2)
Aphonus densicaudus Casey,1915   United States 1Pennsylvania
Dynastinae N.Am.2 (1)
Aplidia fissa Burmeister,1855   Turkey 1Melolonthinae Pal.18 (1)
Aplidia graeca (Kraatz,1882)   Greece 3Melolonthinae Pal.8 (3)
Aplidia lateriventris Reitter,1902   Turkey 2Melolonthinae Pal.18 (2)
Aplidia lodosi Baraud,1988   Turkey 1Melolonthinae Pal.14 (1) as Haplidia l.
Aplidia pruinosa Baudi,1870   Cyprus 1Melolonthinae Pal.12 (1)
Aplidia spp.    Cyprus 1Melolonthinae Pal.9 (1)
Greece 3Melolonthinae Pal.9 (3)
Italy 1Melolonthinae Pal.9 (1)
Turkey 1Melolonthinae Pal.14 (1) Female
Unknown 2Melolonthinae Pal.5 (2)
Aplidia tarsensis Kraatz,1882   Turkey 4Melolonthinae Pal.16 (4) as Haplidia t.
Aplidia transversa (Fabricius,1801)   Albania 1Melolonthinae Pal.18 (1)
Bosnia and Herzegovina 1Melolonthinae Pal.13 (1)
Germany 1Melolonthinae Pal.6 (1)
Greece 3Melolonthinae (3) Pal5+.8
Italy 2Melolonthinae Pal.5 (1)
Pal.13 (1)
Turkey 22Melolonthinae Pal.14 (19) as Haplidia t.
Pal.18 (3) as Haplidia t.
Unknown 7Melolonthinae Pal.13 (7)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 19Melolonthinae Pal.8 (19)
Aplidia transversa peloponnisica (Fabricius,1801)   Greece 1Melolonthinae Pal.12 (1)
Aplidia transversa transversa (Fabricius,1801)   Turkey 2Melolonthinae Pal.14 (2) as Haplidia t.
Aplidia vagepunctata (Kraatz,1882)   Turkey 6Melolonthinae Pal.14 (6) as Haplidia v.
Apogonia aequabilis Karsch,1882   Sri Lanka 27Melolonthinae Or.1 (27)
Apogonia bicavata Arrow,1913   Japan 2Melolonthinae Pal.12 (2)
Apogonia comosa Karsch,1882   Sri Lanka 153Melolonthinae (153) Or.1+2+3
Apogonia gracilis Arrow,1916   Sri Lanka 1Melolonthinae Or.1 (1)
Apogonia laevissima Frey,1973   Sri Lanka 2Melolonthinae Or.1 (2)
Apogonia rauca (Fabricius,1781)   Sri Lanka 20Melolonthinae Or.1 (20)
Apogonia solida (Walker,1859)   Sri Lanka 14Melolonthinae Or.1 (14)
Apogonia spp.    China 1Melolonthinae Or.10 (1)
Indonesia 11Java
Melolonthinae Or.9 (11)
Malaysia 3Melolonthinae (3) Or.9+10
Singapore 1Melolonthinae Or.9 (1)
Thailand 1Melolonthinae Or.9 (1)
Borneo, Indonesia / Malaysia 6Melolonthinae (6) Or.9+10
Archophileurus foveicollis (Burmeister,1847)   Brazil 1Dynastinae (1) S.Am.21
Unknown 1Dynastinae (1) S.Am.21
Archophileurus kolbeanus (Ohaus,1910)   Unknown 1Dynastinae (1) S.Am.21
Archophileurus ovis (Burmeister,1847)   Panama 1Dynastinae (1) S.Am.21
Archophileurus vervex (Burmeister,1847)   Argentina 2Dynastinae (2) S.Am.21
Uruguay 1Dynastinae (1) S.Am.21
Areoda banksi Laporte,1840   Brazil 1Rutelinae S.Am.1 (1)
Unknown 1Rutelinae S.Am.1 (1)
Areoda leachi MacLeay,1819   Argentina 1Rutelinae S.Am.1 (1)
Brazil 3Rutelinae S.Am.1 (3)
Aspidolea bleuzeni Dechambre,1992   Guyana 1Dynastinae (1) S.Am.32
Aspidolea cognata Höhne,1922   Colombia 4Dynastinae S.Am.19 (4)
Aspidolea fuliginea (Burmeister,1847)   Ecuador 2Dynastinae (2) S.Am.34
Astaena brasiliensis Blanchard,1850   Brazil 1Sericini S.Am.1 (1)
Ataeniopsis haroldi (Steinheil,1872)   Bolivia 1Aphodiinae S.Am.1 (1) as Ataenius h.
Ataenius abditus (Haldeman,1848)   United States 1New York
Aphodiinae N.Am.5 (1)
Ataenius australasiae Boheman,1858   Malaysia 8Borneo
Aphodiinae Or.7 (8)
Thailand 2Aphodiinae Or.2 (2)
Borneo, Indonesia / Malaysia 21Aphodiinae Or.3 (21)
Ataenius brevicollis (Wollaston,1854)   Portugal 7Madeira
Aphodiinae (7) Pal.12
Spain 2Canary Islands
Aphodiinae Pal.3 (2)
Ataenius caesoides Fairmaire,1899   Madagascar 1Aphodiinae (1) Afr.24
Ataenius capitosus Harold,1867   Peru 1Aphodiinae S.Am.1 (1) as Phalangochaeta c.
Ataenius carinator Harold,1874   Mexico 1Aphodiinae N.Am.2 (1)
Peru 1Aphodiinae S.Am.1 (1)
Ataenius cognatus (LeConte,1858)   Mexico 3Aphodiinae N.Am.5 (3)
United States 8Connecticut
Aphodiinae N.Am.5 (8)
United States 2New York
Aphodiinae N.Am.5 (2)
Unknown 2Aphodiinae N.Am.5 (2)
Ataenius confertus Fall,1909   United States 4Arizona
Aphodiinae N.Am.8 (4)
Ataenius coriarius Schmidt,1912   Senegal 1Aphodiinae (1) Afr.13
Ataenius deserti Blackburn,1894   Australia 1Queensland
Aphodiinae Pac.1 (1)
Ataenius gracilis (Melsheimer,1846)   Chile 1Aphodiinae S.Am.1 (1)
Costa Rica 1Aphodiinae S.Am.2 (1)
Mexico 1Aphodiinae N.Am.2 (1)
Peru 1Aphodiinae S.Am.1 (1)
South Africa 1Aphodiinae (1) Afr16
United States 2Georgia
Aphodiinae N.Am.2 (2)
United States 1West Virginia
Aphodiinae N.Am.2 (1)
United States 3Texas
Aphodiinae N.Am.2 (3)
United States 2New York
Aphodiinae N.Am.5 (2)
United States 1Louisiana
Aphodiinae N.Am.2 (1)
United States 1Illinois
Aphodiinae N.Am.5 (1)
United States 1Florida
Aphodiinae N.Am.2 (1)
Ataenius granocostatus  Schmidt,1912   Ghana 5Aphodiinae (5) Afr.17
Ataenius hirsutus Horn,1871   Mexico 6Aphodiinae N.Am.2 (6)
Ataenius horticola Harold,1869   Greece 1Aphodiinae (1) Pal.12
Ataenius imbricatus (Melsheimer,1845)   Costa Rica 2Aphodiinae S.Am.2 (2)
Ataenius insolitus Schmidt,1909   Gambia 6Aphodiinae (6) Afr.13
Guinea 1Aphodiinae Afr.16 (1)
Guinea-Bissau 1Aphodiinae (1) Afr14
Senegal 2Aphodiinae (2) Afr.13
Ataenius integer Harold,1869   Guatemala 1Aphodiinae S.Am.1 (1)
Peru 1Aphodiinae S.Am.1 (1)
Ataenius kratochvili Balthasar,1942   Cameroon 2Aphodiinae Afr.13 (2)
Sierra Leone 77Aphodiinae (77) Afr.13+14
Tanzania 1Aphodiinae Afr.13 (1)
Ataenius kratochvili-group    South Africa 1Aphodiinae Afr.31 (1)
Ataenius lamotteiroyi Bordat,1992   Sierra Leone 53Aphodiinae Afr.14 (53)
Ataenius languidus Schmidt,1910   Nicaragua 1Aphodiinae S.Am.2 (1)
Ataenius liogaster Bates,1867   Guatemala 1Aphodiinae S.Am.1 (1)
Nicaragua 3Aphodiinae S.Am.1 (3)
United States 2Hawaii
Aphodiinae N.Am.5 (2)
Ataenius microtrichopterus Lea,1923   Australia 1Queensland
Aphodiinae Pac.1 (1)
Ataenius monticola Paulian,1940   Cameroon 1Aphodiinae (1) 1 paratype Afr.24
Ataenius nocturnus (Nomura,1943)   United States 4Arizona
Aphodiinae N.Am.2 (4)
Ataenius nugator Harold,1880   Ecuador 3Aphodiinae S.Am.1 (3)
Mexico 2Aphodiinae N.Am.3 (2)
Peru 1Aphodiinae S.Am.1 (1)
Ataenius nugator-group    Canada 2Ontario
Aphodiinae N.Am.2 (2)
Canada 2Quebec
Aphodiinae N.Am.2 (2)
Mexico 2Aphodiinae N.Am.5 (2)
United States 17Arizona
Aphodiinae N.Am.2 (17)
United States 2West Virginia
Aphodiinae N.Am.2 (2)
United States 1Texas
Aphodiinae N.Am.2 (1)
United States 3Louisiana
Aphodiinae N.Am.2 (3)
United States 2Florida
Aphodiinae N.Am.2 (2)
United States 8California
Aphodiinae N.Am.2 (8)
United States 2Illinois
Aphodiinae N.Am.2 (2)
Ataenius orbicularis Schmidt,1914   Malaysia 4Aphodiinae Or.2 (4)
Thailand 11Aphodiinae Or.2 (11)
Ataenius peregrinator Harold,1877   Malaysia 1Aphodiinae Or.2 (1)
Mexico 2Aphodiinae N.Am.2 (2)
Ataenius picinus Harold,1867   Costa Rica 1Aphodiinae S.Am.1 (1)
Ataenius platensis (Blanchard,1843)   Mexico 1Aphodiinae N.Am.3 (1)
Peru 1Aphodiinae S.Am.1 (1)
Ataenius puncticollis Leconte,1858   Mexico 3Aphodiinae N.Am.4 (3)
Ataenius punctipennis Harold,1868   Colombia 1Aphodiinae S.Am.2 (1)
Ataenius scabrelloides Petrovitz,1962   United States 1Florida
Aphodiinae N.Am.8 (1)
Ataenius scalptifrons Bates,1887   Mexico 1Aphodiinae N.Am.2 (1)
Ataenius sparsicollis Blackburn,1904   Australia 1New South Wales
Aphodiinae Pac.1 (1)
Ataenius spp.    South Africa 2Aphodiinae Afr.31 (2)
Ataenius stephani Cartwright,1974   Mexico 2Aphodiinae N.Am.4 (2)
Ataenius stercorator (Fabricius,1775)   Peru 3Aphodiinae S.Am.1 (3)
Ataenius strigatus (Say,1823)   United States 1Illinois
Aphodiinae N.Am.8 (1)
Ataenius strigicaudus Bates,1887   Argentina 1Aphodiinae S.Am.2 (1)
Costa Rica 3Aphodiinae S.Am.2 (3)
Ataenius texanus Harold,1874   Guatemala 1Aphodiinae S.Am.1 (1)
Mexico 8Aphodiinae (8) N.Am.3+5
Ateuchetus cicatricosus (Lucas,1849)   Morocco 1Scarabaeinae Pal.33 (1) as Scarabaeus c.
Spain 3Scarabaeinae Pal.20 (3) as Scarabaeus c.
Tunisia 1Scarabaeinae Pal.50 (1) as Scarabaeus variolosus
Ateuchus chrysopyge (Bates,1887)   Guatemala 3Scarabaeinae S.Am.2 (3)
Ateuchus guatemalensis (Bates,1887)   Honduras 1Scarabaeinae S.Am.2 (1)
Ateuchus histeroides Weber,1801   United States 1Mississippi
Scarabaeinae N.Am.1 (1)
Unknown 2Scarabaeinae N.Am.2 (2)
America borealis, United States/Canada 1Scarabaeinae N.Am.2 (1)
Ateuchus rodriguezi (Preudhome de Borre,1886)   Nicaragua 1Scarabaeinae S.Am.7 (1)
Ateuchus semicribratum Harold,1868   Brazil 1Scarabaeinae S.Am.8 (1)
Ateuchus simplex LePeletier & Serville,1828   French Guiana 1Scarabaeinae S.Am.8 (1)
Ateuchus smaragdulus Olivier,   Amer. Merid. 1Scarabaeinae S.Am.2 (1)
Ateuchus spp.    Brazil 4Scarabaeinae S.Am.8 (4)
Honduras 4Scarabaeinae S.Am.5 (4)
Peru 1Scarabaeinae S.Am.2 (1)
America borealis, United States/Canada 2Scarabaeinae N.Am.2 (2)
Atrichelaphinis tigrina (Olivier,1789)   South Africa 7Cetoniinae (7) Afr.5+14+25
Unknown 4Cetoniinae (4) Afr.25
Augoderia nitidula Burmeister,1847   Brazil 3Dynastinae S.Am.17 (3)
Augosoma centaurus (Fabricius,1775)   Cameroon 3Dynastinae (3) Afr.10+11
Congo 2Dynastinae (2) Afr.10+11
Guinea 1Dynastinae Afr.10 (1)
Uganda 1Dynastinae Afr.20 (1)
Unknown 9Dynastinae (9) Afr.9+10+11
Aulacopalpus ciliatus (Solier,1851)   Chile 1Rutelinae (1) S.Am.4
Aulonocnemis crassecostata Fairmaire,1886   Kenya 2Aphodiinae (2) Afr.16
Australaphodius frenchi Blackburn,1892   South Africa 27Aphodiinae Afr.23 (6) As Aphodius f.
Afr.32 (21)
Autoserica bomuana Brenske,1899   Eritrea 2Sericini Afr.1 (2)
Autoserica caifesis (Brenske,1897)   Afghanistan 2Sericini Pal.1 (2)
Autoserica spp.    Cameroon 10Sericini Afr.1 (10)
Philippines 3Sericini Or.2 (3)
Tanzania 1Sericini Afr.1 (1)
Biphyllocera kirbyana White,1841   Australia 1Western Australia
Melolonthinae Pac.1 (1)
Bizanus caliginosus Peringuey,1902   South Africa 2Hopliini (2) Afr.12
Blackburneus ineptus (Petrovitz,1964)   Kenya 1Aphodiinae Afr.8 (1) as Aphodius
Blackburneus jugalis (Péringuey,1901)   Congo 1Aphodiinae Afr.20 (1) 1 paratype
as Aphodius biroi Endrödi 1957 (Blackburneus)
Blackburneus microreticulatus (Landin,1967)   South Africa 1Type Landin (1) as Aphodius Holotype 03489:1 of Aphodius microreticulatus Landin 1967
  Aphodius microreticulatus Landin 1967

Blackburneus stercorosus (Melsheimer,1845)   United States 1Connecticut
Aphodiinae N.Am.5 (1) as Aphodius
United States 4New York
Aphodiinae N.Am.5 (4) as Aphodius
United States 1Virginia
Aphodiinae N.Am.2 (1) as Aphodius
United States 1Tennessee
Aphodiinae N.Am.8 (1)
Blackburneus tachyoryctis (Endrödi,1960)   Tanzania 4Aphodiinae Afr.20 (4) Paratypes
as Aphodius
Blitopertha lineata (Fabricius,1798)   Algeria 2Rutelinae Pal.6 (2) as Phyllopertha l.
Greece 1Rutelinae Pal.6 (1) as Phyllopertha l.
Blitopertha lineolata (Fischer von Waldheim,1824)   Bulgaria 2Rutelinae Pal.1 (2)
Cyprus 2Rutelinae Pal.1 (2)
Greece 21Rutelinae Pal.1 (19)
Pal.13 (2)
Turkey 7Rutelinae Pal.1 (1)
Pal.12 (3)
Pal.12 (1) as ab. senticola Rtt.
Pal.13 (2)
Blitopertha nigripennis (Reitter,1888)   Armenia 3Rutelinae Pal.13 (3) as B.majuscula
Iran 2Rutelinae Pal.13 (2)
Israel 1Rutelinae Pal.10 (1) as B. abdita Petr.
Turkey 27Rutelinae Pal.9 (8) as B. majuscula Medvedev
Pal.12 (14) as B. majuscula Medvedev
Pal.12 (4) as B. abdita Petr.
Pal.13 (1) as B. majuscula
Bodiloides ictericus (Leicharting,1781)   Bulgaria 2Aphodiinae Pal.7 (2)
Denmark 4Aphodiinae (4)
Germany 1Aphodiinae (1) Pal.15
Greece 20Aphodiinae (20) Pal.7+23
Morocco 31Aphodiinae (31) Pal.7+16+23
Norway 1Aphodiinae Main (1)
Portugal 16Madeira
Aphodiinae Pal.3 (16) as A. nitidulus F.
Spain 8Canary Islands
Aphodiinae Pal.7 (8)
Spain 42Canary Islands
Aphodiinae (42) Pal.3+15 (as.A.nitidulus F.)

Spain 15Aphodiinae (15) Pal.7+23
Sweden 296  Skåne Blekinge Halland Öland Gotland Östergötland Västergötland Bohuslän Uppland
Alan Dufberg 48 (29)
Aphodiinae Main (267)
Switzerland 7Aphodiinae Pal.18 (7)
Tunisia 9Aphodiinae Pal.7 (9)
Turkey 5Aphodiinae Pal.27 (4)
Pal.30 (1) as Aphodius ictericus
Unknown 14Aphodiinae Pal.23 (14)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Aphodiinae Pal.7 (2)
Bodiloides ictericus ghardimaouensis (Balthasar,1929)   Italy 1Aphodiinae Pal.10 (1)
Portugal 3Madeira
Aphodiinae Pal.27 (3) as A. ghardimaoensis Balthasar
Spain 2Mallorca
Aphodiinae Pal.27 (2)
Spain 11Canary Islands
Aphodiinae Pal.10 (1)
Pal.27 (7) as Aphodius ghardimaoensis Balthasar
Pal.28 (3)
Spain 2Aphodiinae Pal.11 (2)
Tunisia 2Aphodiinae Pal.11 (2)
Bodilopsis rufa (Moll,1782)   Denmark 1Aphodiinae Main (1)
Finland 1Aphodiinae (1)
Norway 1Aphodiinae Main (1)
Sweden 524  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up Vr Dr Gä Me Jä Ån Nb
Alan Dufberg 48 (22)
Aphodiinae Main (502) as rufus
Tunisia 3Aphodiinae Pal.16 (2) as A. scybalarius F.
Pal.30 (1)
Unknown 26Aphodiinae Pal.23 (26)
Bodilopsis sordida (Fabricius,1775)   Norway 4Aphodiinae Main (4)
Sweden 365  Sk Bl Ha Sm Öl Go Ög Vg Bo Nä Sö Up Dr Ån
Alan Dufberg 48 (16)
Aphodiinae Main (348)
Aphodiinae Main (1) Type of A. bipustulatus G. Möller
Bodilus acutifrons  Petrovitz,1958   Botswana 2Aphodiinae Afr.16 (2) incl. 1 paratype
South Africa 2Aphodiinae Afr.16 (2)
Bodilus amabils Boheman,1857   South Africa 13Aphodiinae Afr.21 (6)
Afr.34 (7)
Bodilus ardens Harold,1866   South Africa 4Aphodiinae Afr.21 (1)
Afr.25 (1)
Afr.34 (2)
Bodilus cathedralensis (Endrödi,1991)   Lesotho 1Aphodiinae Afr.35 (1)
Bodilus cf. laterosetosus Petrovitz,1958   South Africa 3Aphodiinae Afr.32 (3)
Bodilus laterosetosus Petrovitz,1958   Botswana 1Aphodiinae Afr.16 (1)
South Africa 92Aphodiinae Afr.33 (92)
Bodilus lugens Creutzer,1799   Bulgaria 29Aphodiinae (22) Pal.5+10
Pal.29 (7)
Unknown 10Aphodiinae Pal.23 (10)
Bodilus sordescens Harold,1869   Russian Federation 1Aphodiinae Pal.18 (1)
Bodilus spp.    Lesotho 3Aphodiinae Afr.35 (3)
Tunisia 3Aphodiinae Pal.28 (3)
Bolax flavolineatus Mannerheim,1829   Brazil 5Rutelinae (5) S.Am.7
Bolax glabripennis Ohaus,1917   Unknown 1Rutelinae (1) S.Am.7
Bolax magnus Bates,1888   Colombia 1Rutelinae (1) S.Am.7
Bolax palliatus Burmeister,1844   Unknown 1Rutelinae (1) S.Am.7
Bolax phaleratus Burmeister,1844   Brazil 1Rutelinae (1) S.Am.7
Bolax salticola Ohaus,1908   Brazil 1Rutelinae (1) S.Am.7
Bolax sulcicollis (Germar,1824)   Brazil 1Rutelinae (1) S.Am.7
Bolax zoubkoffi Fisch.,1829   Brazil 1Rutelinae (1) S.Am.7
Colombia 1Rutelinae (1) S.Am.7
Unknown 2Rutelinae (2) S.Am.7
Bothynus cunctator (Mannerheim,1829)   Brazil 3Dynastinae S.Am.16 (3)
Bothynus cyclops (Burmeister,1847)   Brazil 1Dynastinae S.Am.16 (1)
Bothynus dasypleurus (Germar,1824)   Unknown 1Dynastinae S.Am.16 (1)
Bothynus entellus (Serville,1825)   Brazil 3Dynastinae S.Am.16 (3)
Unknown 2Dynastinae S.Am.16 (2)
Bothynus exaratus (Burmeister,1847)   Brazil 1Dynastinae S.Am.16 (1)
Bothynus herbivorus (Arow,1937)   Unknown 1Dynastinae S.Am.16 (1)
Bothynus medon (Germar,1824)   Brazil 3Dynastinae S.Am.16 (3)
Unknown 4Dynastinae S.Am.16 (4)
Bothynus minor Steinheil,1872   Argentina 2Dynastinae (2) S.Am.16+30
Bothynus spp.    Unknown 1Dynastinae S.Am.16 (1)
Bothynus striatellus (Fairmaire,1878)   Argentina 1Dynastinae S.Am.16 (1)
Unknown 1Dynastinae S.Am.16 (1)
Bothynus validus (Burmeister,1847)   Brazil 1Dynastinae S.Am.16 (1)
Brachiaphodius aureopilosus (Boucomont,1930)   Kenya 1Aphodiinae Afr.30 (1)
Brachiaphodius eccoptus (Bates,1889)   Japan 4Aphodiinae Pal.17 (4) +18
Brachiaphodius jeanneli (Paulian,1938)   Guinea 1Aphodiinae Afr.21 (1)
Kenya 5Aphodiinae Afr.21 (5)
South Africa 1Aphodiinae Afr.20 (1) 1 paratype
Brachiaphodius jendeki (Kral,2001)   Myanmar 5Aphodiinae Or.6 (5)
Brachiaphodius langeri (Bellman,2009)   Myanmar 2Type Coleoptera (2) Paratype 06785:2-3 of Aphodius langeri Bellman 2009
  Aphodius langeri Bellman 2009

Brachiaphodius pilosus (Harold,1874)   Nepal 3Aphodiinae Or.3 (3)
Brachysternus prasinus Guerin-Meneville,1831   Chile 3Rutelinae (3) S.Am.8
Brachysternus spectabilis Erichson,1847   Chile 1Rutelinae (1) S.Am.8
Brachysternus viridis (Solier,1851)   Chile 1Rutelinae S.Am.2 (1)
Brahmina sedakovii (Mannerheim,1849)   Mongolia 3Melolonthinae (3) Pal.5+12
Brancoplia leucaspis (Laporte,1840)   Unknown 2Rutelinae (2) Pal.8 (as Anisoplia l.)
Unknown 1Rutelinae Pal.9 (1)
Bricoptis variolosa (Goron & Percheron,1833)   Madagascar 1Cetoniinae Afr.7 (1)
Brindalus porcicollis (Illiger,1803)   Cyprus 1Aphodiinae Pal.12 (1)
Italy 13Aphodiinae (13) Pal.3+12
Portugal 5Azores
Type Landin (5) Holotype Paratype 03483:1-5 of Phycochus azoricus Landin 1960
  Phycochus azoricus Landin 1960

Portugal 8Madeira
Aphodiinae Pal.4 (8) as Psammodius p.
Spain 4Aphodiinae Pal.12 (4)
Spain 1Canary Islands
Aphodiinae Pal.26 (1) as Psammodius p.
Tunisia 1Aphodiinae Pal.12 (1)
Unknown 1Aphodiinae Pal.15 (1)
Brindalus rotundipennis (Reitter,1892)   Spain 1Aphodiinae (1) Pal.12
Brindalus schatzmayri (Pittino,1980)   Spain 2Canary Islands
Aphodiinae Pal.3 (2) as Psammodius porcicollis Ill.
Bruchaphodius ovalipennis (Harold,1871)   Argentina 1Aphodiinae S.Am.2 (1)
Bruneixenus squamosus Howden & Storey,1992   Borneo, Indonesia / Malaysia 1Aphodiinae Or.6 (1)
Bubas bison (Linnaeus,1767)   Italy 3Scarabaeinae (3) Pal.15+23
Spain 12Scarabaeinae (12) Pal.14+15
Spain 2Mallorca
Scarabaeinae Pal.38 (2)
Tunisia 1Scarabaeinae Pal.15 (1)
Unknown 1Scarabaeinae (1) Pal.34
Bubas bubaloides Janssens,1938   Cyprus 3Scarabaeinae Pal.51 (3) as Bubas bubalus ssp. bubaloides
Bubas bubalus (Olivier,1811)   Spain 4Scarabaeinae (4) Pal.24
Unknown 1Scarabaeinae (1) Pal.34
Caccobius brevis Waterhouse,1875   Japan 2Scarabaeinae Pal.34 (2)
Caccobius christophi Harold,1879   Russian Federation 1Scarabaeinae (1) Pal.34
Caccobius histeroides (Menetries,1832)   Armenia 4Scarabaeinae Pal.45 (4)
Greece 3Scarabaeinae Pal.12 (3)
Israel 1Scarabaeinae Pal.52 (1)
Caccobius jessoensis Harold,1867   Japan 2Scarabaeinae (2) Pal.34
Caccobius nikkoensis (Lewis,1895)   Japan 2Scarabaeinae (2) Pal.34
Japan 1Scarabaeinae Pal.35 (1) as Onthophagus n.
Caccobius schreberi (Linnaeus,1767)   Armenia 5Scarabaeinae Pal.45 (5)
Bulgaria 8 Alan Dufberg 47 (2)
Scarabaeinae Pal.13 (6)
France 1Scarabaeinae (1) Pal.13
Germany 1Scarabaeinae (1) Pal.33
Greece 13Scarabaeinae (13) Pal.12
Italy 23Scarabaeinae (23) Pal.13
Romania 1Scarabaeinae (1) Pal.13
Spain 10Scarabaeinae (10) Pal.12
Sweden 221 Skåne Öland
Scarabaeinae Main (221)
Turkey 2Scarabaeinae (2) Pal.13
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 10Scarabaeinae (10) Pal.13
Caccobius sordidus Harold,1886   Russian Federation 1Scarabaeinae (1) Pal.34
Caccobius spp.    Ethiopia 2Scarabaeinae Afr.1 (2)
Guinea-Bissau 1Scarabaeinae (1) Afr.22
Caccobius suzukii Matsumura,1936   Japan 1Scarabaeinae Pal.34 (1)
Caelius browni (Saylor,1934)   United States 4California
Aphodiinae N.Am.3 (4) as Aegialia browni
Caelius denticollis Lewis,1895   Japan 3Aphodiinae Pal.18 (2) as Aegialia denticollis
Pal.28 (1)
Caelius humeralis (Brown,1931)   United States 1Connecticut
Aphodiinae N.Am.8 (1)
Caelius montana (Brown,1931)   United States 1Alaska
Aphodiinae N.Am.4 (1) as Aegialia montana
Caelius rufescens (Horn,1887)   United States 1Alaska
Aphodiinae N.Am.4 (1) as Aegialia rufescens
Caelorrhina barthi Harold,1878   Congo, the democratic republic of the 1Cetoniinae (1) Afr.18
Caelorrhina semiviridis Bourgoin,1919   Tanzania 1Cetoniinae (1) Afr.18 (as Eccoptocnemis s.)

Caelorrhina spp.    Kenya 1Cetoniinae Afr.2 (1)
Calamosternus granarius (Linnaeus,1767)   Bulgaria 12Aphodiinae Pal.6 (11)
Pal.29 (1)
Chile 12Aphodiinae S.Am.1 (11)
S.Am.2 (1)
Cyprus 18Aphodiinae Pal.6 (15)
Pal.30 (3) as Aphodius granarius
Denmark 1Aphodiinae (1)
Greece 14Aphodiinae (13) Pal.6+11+13
Pal.30 (1) as Aphodius granarius
Portugal 124Azores
Aphodiinae Pal.1 (122)
Pal.27 (1)
Pal.28 (1)
South Africa 3Aphodiinae Afr.31 (3)
Spain 9Canary Islands
Aphodiinae Pal.3 (1)
Pal.10 (1)
Pal.27 (2)
Pal.28 (5)
Sweden 223  Skåne Blekinge Halland Öland Gotland Östergötland Västergötland Södermanland Uppland Värmland
Alan Dufberg 48 (29)
Aphodiinae Main (194)
Unknown 5Aphodiinae Pal.25 (5)
Calamosternus mayeri (Pilleri,1953)   Morocco 4Aphodiinae Pal.11 (4)
Calamosternus trucidatus (Harold,1863)   Italy 6Aphodiinae Pal.3 (6)
Turkey 1Aphodiinae Pal.28 (1)
Calamosternus uniplagiatus (Waterhouse,1875)   China 2Aphodiinae Pal.23 (2)
Japan 3Aphodiinae (3) Pal.18+24
Calicnemis latreillei Laporte,1832   Unknown 2Dynastinae Pal.3 (2)
Caligodorus opacus (LeConte,1872)   United States 5Washington
Aphodiinae N.Am.1 (5) as Aphodius
United States 7California
Aphodiinae (7) N.Am.1+5 as Aphodius
United States 1Oregon
Aphodiinae N.Am.1 (1) as Aphodius
United States 1Montana
Aphodiinae N.Am.8 (1)
Callistethus auronitens (Hope,1836)   India 2Rutelinae Or.7 (2)
Myanmar 4Rutelinae Or.7 (4)
Callistethus plagiicollis (Fairmaire,1886)   China 1Rutelinae (1) Pal.4 (as Anomala plagiicollis)
Callistethus pterygophorus Ohaus,1903   India 3Rutelinae Or.7 (3)
Callistopopillia iris (Candeze,1869)   India 3Rutelinae Or.8 (3)
Calloodes atkinsoni    Australia 3Rutelinae (3) Pac.1
Calloodes grayianus (White,1845)   Australia 1Rutelinae (1) Pac.1
Calomacraspis haroldi (Cand.,1869)   Unknown 1Rutelinae (1) S.Am.4
Calomacraspis splendens (Burmeister,1844)   Mexico 3Rutelinae N.Am.2 (3)
Camenta judanica Frey,   Sudan 1Melolonthinae Afr.2 (1) paratype
Campsiura cognata (Schaum,1841)   South Africa 1Cetoniinae Afr.5 (1)
Unknown 3Cetoniinae Afr.29 (3)
Campsiura javanica Gory & Percheron,1833   Unknown 2Cetoniinae Afr.28 (2)
Campsiura spp.    Cameroon 1Cetoniinae Afr.29 (1)
Campulipus limbatus (Olivier,1789)   Unknown 5Cetoniinae Afr.29 (5)
Campulipus rufipennis (Gory & Percheron, 1833)   Unknown 2Cetoniinae Afr.29 (2)
Canthidium aroldi Preudhomme,1886   Colombia 1Scarabaeinae S.Am.2 (1)
Canthidium breve Germar,1824   Argentina 1Scarabaeinae S.Am.8 (1)
Canthidium chrysis Fabricius,1801   Brazil 1Scarabaeinae S.Am.8 (1)
Canthidium decoratum Perty,1830   Brazil 1Scarabaeinae S.Am.8 (1)
Canthidium melanocephalum Olivier,1789   French Guiana 1Scarabaeinae S.Am.8 (1)
Canthidium spp.    Argentina 2Scarabaeinae S.Am.8 (2)
Brazil 4Scarabaeinae ()
S.Am.8 (4)
Ecuador 14Scarabaeinae S.Am.2 (14)
Honduras 8Scarabaeinae S.Am.2 (8)
Mexico 4Scarabaeinae N.Am.1 (4)
Peru 1Scarabaeinae S.Am.2 (1)
Canthidium trinodosum Boheman,1858   Brazil 1Scarabaeinae S.Am.8 (1)
French Guiana 1Scarabaeinae S.Am.8 (1)
Canthon aberrans (Harold,1868)   Ecuador 1Scarabaeinae S.Am.4 (1)
Canthon aequinoctiale Harold,1868   Ecuador 1Scarabaeinae S.Am.4 (1)
Canthon affinis Fabricius,1801   Brazil 4Scarabaeinae S.Am.9 (4)
Mexico 2Scarabaeinae S.Am.9 (2)
Paraguay 1Scarabaeinae S.Am.9 (1)
Unknown 3Scarabaeinae S.Am.9 (3)
Canthon cyanellus LeConte,1859   Nicaragua 1Scarabaeinae S.Am.6 (1)
Canthon femoralis Chevrolat,1834   Mexico 1Scarabaeinae N.Am.1 (1)
Canthon gagatinus var. amethystinus Harold,1863   Mexico 1Scarabaeinae S.Am.9 (1)
Unknown 1Scarabaeinae S.Am.9 (1)
Canthon humectus Say,1831   Brazil 1Scarabaeinae S.Am.9 (1)
Mexico 5Scarabaeinae S.Am.9 (5)
Unknown 1Scarabaeinae S.Am.9 (1)
Canthon humectus sayi Robinson,1948   Mexico 5Scarabaeinae N.Am.1 (5)
Canthon imitator Brown,1946   Mexico 3Scarabaeinae N.Am.1 (3)
Canthon indigaceus LeConte,1866   United States 6Arizona
Scarabaeinae N.Am.1 (6)
Canthon indigaceus chevrolati Harold,1868   Mexico 1Scarabaeinae S.Am.9 (1)
Canthon laevis (Drury,1773)   United States 2New York
Scarabaeinae N.Am.3 (2)
United States 1Kansas
Scarabaeinae N.Am.3 (1)
United States 1Pennsylvania
Scarabaeinae N.Am.3 (1)
United States 1New Jersey
Scarabaeinae N.Am.3 (1)
United States 1Wisconsin
Scarabaeinae N.Am.3 (1)
America borealis, United States/Canada 10Scarabaeinae N.Am.3 (10)
Unknown 13Scarabaeinae N.Am.3 (13)
America septentrionalis, United States/Canada 1Scarabaeinae N.Am.3 (1)
Canthon lividus seminitens Harold,1868   Brazil 2Scarabaeinae S.Am.9 (2)
Canthon luteicolle Erichson,1847   Ecuador 2Scarabaeinae S.Am.4 (2)
Canthon mutabilis Lucas,1857   Brazil 1Scarabaeinae S.Am.9 (1)
Unknown 1Scarabaeinae S.Am.9 (1)
Canthon ornatus thoracicus Harold,1868   Brazil 1Scarabaeinae S.Am.9 (1)
Canthon pilularius Linnaeus,1758   Unknown 2Scarabaeinae N.Am.3 (2)
Canthon probus Germar,1824   America borealis, United States/Canada 1Scarabaeinae N.Am.3 (1)
Canthon rutilans Laporte,1840   Brazil 5Scarabaeinae S.Am.10 (5)
Haiti 1Scarabaeinae S.Am.10 (1)
Unknown 3Scarabaeinae S.Am.10 (3)
Canthon septemmaculatus Latreille,1811   Brazil 1Scarabaeinae S.Am.9 (1)
Colombia 2Scarabaeinae S.Am.9 (2)
Unknown 2Scarabaeinae S.Am.9 (2)
Canthon simplex Leconte,1857   United States 2California
Scarabaeinae N.Am.3 (2)
Canthon smaragdulus (Fabricius,1781)   Brazil 14Scarabaeinae S.Am.10 (14)
Unknown 12Scarabaeinae S.Am.10 (12)
Canthon spp.    Brazil 6Scarabaeinae S.Am.10 (6)
Colombia 2Scarabaeinae S.Am.10 (2)
Ecuador 1Scarabaeinae S.Am.5 (1)
Mexico 2Scarabaeinae N.Am.1 (2)
Mexico 7Scarabaeinae S.Am.10 (7)
Suriname 1Scarabaeinae S.Am.10 (1)
Unknown 4Scarabaeinae S.Am.10 (4)
Canthon triangularis Drury,1770   Brazil 18Scarabaeinae S.Am.6 (5)
S.Am.11 (13)
French Guiana 1Scarabaeinae S.Am.11 (1)
Suriname 1Scarabaeinae S.Am.11 (1)
America meridionalis 2Scarabaeinae S.Am.11 (2)
Unknown 5Scarabaeinae S.Am.11 (5)
Canthon unicolor tristis Harold,1862   Brazil 2Scarabaeinae S.Am.10 (2)
Unknown 1Scarabaeinae S.Am.10 (1)
Canthon viduus Harold,1868   French Guiana 2Scarabaeinae S.Am.10 (2)
Canthon violaceus (Olivier,1789)   Saint Vincent and The Grenadines 1Scarabaeinae S.Am.9 (1)
Unknown 2Scarabaeinae S.Am.9 (2)
Canthon viridis Beauvois,1805   Mexico 1Scarabaeinae N.Am.3 (1)
Unknown 1Scarabaeinae N.Am.3 (1)
America borealis, United States/Canada 2Scarabaeinae N.Am.3 (2)
Canthonella spp.    Ecuador 1Scarabaeinae (1) S.Am.5
Catharsius abortivus (Boheman,1891)   South Africa 1Scarabaeinae Afr.35 (1)
Catharsius achates (Olivier,1789)   Guinea-Bissau 1Scarabaeinae Afr.6 (1)
Catharsius bradsawi Lansberge,1887   Namibia 2Scarabaeinae (2) Afr.35
Catharsius capucinus Fabricius,1781   Unknown 3Scarabaeinae Or.9 (3)
Catharsius chinai Ferreira,1960   Ethiopia 23Scarabaeinae (23) Afr.7+11
Catharsius disseptus Peringuey,1902   South Africa 2Scarabaeinae (2) Afr.35
Catharsius dux Harold,1878   Congo, the democratic republic of the 3Scarabaeinae Afr.46 (3)
Catharsius eteocles Castelnau,1840   Cameroon 5Scarabaeinae Afr.46 (5)
India 1Scarabaeinae Or.5 (1)
Catharsius furcillatus Guérin-méneville,1847   Tanzania 1Scarabaeinae Afr.57 (1)
Catharsius gorilla Thomson,1858   Cameroon 2Scarabaeinae Afr.46 (2)
Catharsius guineensis Ferreira,1963   Guinea-Bissau 9Scarabaeinae Afr.6 (9)
Catharsius heros  Boheman,1860   Namibia 1Scarabaeinae (1) Afr.35
Catharsius marcellus Kolbe,1893   South Africa 2Scarabaeinae (2) Afr.35
Catharsius molossus Linnaeus,1758   China 4Scarabaeinae Or.6 (4)
India 2Scarabaeinae Or.6 (2)
Indonesia 1Sumatra
Scarabaeinae Or.7 (1)
Indonesia 2Scarabaeinae Or.7 (2)
Indonesia 3Java
Scarabaeinae Or.7 (3)
Myanmar 3Scarabaeinae Or.7 (3)
Nepal 1Scarabaeinae Or.2 (1)
Sri Lanka 3Scarabaeinae Or.7 (3)
Borneo, Indonesia / Malaysia 1Scarabaeinae Or.6 (1)
Unknown 19Scarabaeinae Or.7 (3)
Or.8 (16)
Malacca 1Scarabaeinae Or.7 (1)
Ostindien 7Scarabaeinae Or.6 (7)
Catharsius pithecius Fabricius,1775   Afghanistan 3Scarabaeinae Or.5 (3)
Cameroon 1Scarabaeinae Afr.46 (1)
India 2Scarabaeinae Or.5 (2)
Unknown 4Scarabaeinae Or.5 (4)
Catharsius pseudovitulus Müller,1941   Tanzania 1Scarabaeinae Afr.57 (1)
Catharsius rhinoceros Klug,1855   Tanzania 2Scarabaeinae Afr.47 (2)
Catharsius sagax Quenstedt,1806   India 1Scarabaeinae Or.5 (1)
Unknown 2Scarabaeinae Or.5 (2)
Ind. Or. 1Scarabaeinae Or.5 (1)
Catharsius sesostris Waterhouse,1888   Cameroon 3Scarabaeinae Afr.47 (3)
Ethiopia 1Scarabaeinae Afr.47 (1)
Guinea-Bissau 4Scarabaeinae Afr.6 (4)
Unknown 3Scarabaeinae Afr.47 (3)
Catharsius tricornutus (DeGeer,1778)   South Africa 5Scarabaeinae Afr.35 (4)
Afr.47 (1)
Tanzania 1Scarabaeinae Afr.47 (1)
Unknown 3Scarabaeinae Afr.47 (3)
Catharsius ulysses Boheman,1857   Namibia 2Scarabaeinae (2) Afr.35
South Africa 1Scarabaeinae Afr.9 (1)
Cephalocyclus halffteri Dellacasa, Dellacasa & Gordon,2007   Mexico 1Aphodiinae N.Am.8 (1)
Cephalocyclus mexicanus (Harold,1862)   Mexico 1Aphodiinae N.Am.8 (1)
Cephalodesmius armiger Westwood,1842   Australia 2Queensland
Scarabaeinae Pac.1 (2)
Ceramida bedeaui (Erichson,1840)   Spain 1Melolonthinae Pal.3 (1)
Ceraspis bivulnerata (Germar,1824)   Brazil 1Melolonthinae S.Am.1 (1)
Cetonia asiatica Gory & Percheron,1833   Greece 4Cetoniinae Pal.4 (4) as C. oertzeni Reitter
Cetonia aurata (Linnaeus,1761)   Austria 1Cetoniinae (1) Pal.2 (as C. piligera Mulsant)

Cyprus 1Cetoniinae Pal.22 (1)
Estonia 1Cetoniinae (1) Pal.20
France 2Cetoniinae (2) Pal.16+
Germany 2Cetoniinae Pal.2 (2)
Greece 38Cetoniinae (38) Pal.4+6
Hungary 1Cetoniinae Pal.33 (1)
Italy 4Cetoniinae (4) Pal.2+16
Macedonia, The Former Yugoslav Republic of 3Cetoniinae (3) Pal.
Portugal 2Cetoniinae Pal.2 (2)
Romania 1Cetoniinae Pal.16 (1)
Russian Federation 4Cetoniinae Pal.32 (4)
Slovakia 3Cetoniinae Pal.35 (3)
Sweden 495  Sk Bl Ha Sm Öl Go GS Ög Vg Bo Ds Nä Sö Up Vs Vr Dr Gä
Alan Dufberg 48 (6)
Cetoniinae Main (489)
Switzerland 1Cetoniinae Pal.16 (1)
Turkey 3Cetoniinae (3) Pal.2+16
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 12Cetoniinae Pal.7 (12)
Cetonia aurata pallida (Drury,1773)   Russian Federation 1Cetoniinae Pal.16 (1) as var. undulata Reitter
Turkey 6Cetoniinae Pal.16 (6) as var. undulata Reitter
Cetonia aurata pisana Heer,1841   Italy 8Cetoniinae (8) Pal.10+11
Cetonia carthami Gory & Percheron,1833   Croatia 6Cetoniinae (6) Pal.
Portugal 1Cetoniinae Pal.3 (1)
Spain 1Cetoniinae Pal.3 (1)
Cetonia delagrangei Boucard,1893   Syria 2Cetoniinae (2) Pal.16
Cetonia pilifera (Motschulsky,1860)   Japan 2Cetoniinae (2) Pal.20
Cetonia roelofsi Harold,1880   Japan 3Cetoniinae (3) Pal.11 (as Eucetonia r.)

Chaetonyx robustus Schaum,1862   Greece 1Orphninae Pal.1 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Orphninae Pal.1 (1)
Chaetonyx robustus liguricus Schaum,1862   Italy 2Orphninae Pal.1 (2)
Chaetopteroplia segetum (Herbst,1783)   Austria 7Rutelinae Pal.2 (7)
Bulgaria 3Rutelinae Pal.11 (3) as Anisoplia s.
Germany 1Rutelinae Pal.8 (1)
Greece 7Rutelinae Pal.13 (7) as Anisplia s.
Turkey 63Rutelinae Pal.9 (33)
Pal.12 (19)
Pal.13 (11) partly as Anisplia s.
Unknown 1Rutelinae Pal.8 (1)
Chaetopteroplia segetum rhodiensis (Brullé,)   Greece 1Rutelinae Pal.2 (1)
Chaetopteroplia segetum straminea (Brullé,1832)   Greece 15Rutelinae Pal.2 (15)
Chaetopteroplia segetum velutina (Erichson,1847)   Bulgaria 8Rutelinae Pal.2 (8)
Turkey 3Rutelinae Pal.2 (2)
Pal.13 (1)
Chaetopteroplia spp.    Armenia 1Rutelinae Pal.11 (1)
Turkey 2Rutelinae Pal.11 (2)
Chaetopteroplia syriaca (Burmeister,1844)   Greece 1Rutelinae (1) Pal.8 (as Anisoplia s.)
Turkey 2Rutelinae (2) Pal.8 (as Anisoplia s.)
Unknown 1Rutelinae (1) Pal.9 (as Anisoplia s.)
Chalcocopris hesperus (Olivier,1789)   Brazil 4Scarabaeinae S.Am.4 (2)
S.Am.11 (2)
Unknown 2Scarabaeinae S.Am.11 (2)
Chalcosoma atlas (Linnaeus,1758)   Indonesia 1Celebes
Dynastinae Or.9 (1) as ssp. kirbyii Arrow
Philippines 1Dynastinae Or.13 (1)
Unknown 2Dynastinae Or.9 (2 ) as ssp. kirbyii Arrow
Unknown 4Dynastinae Or.8 (4)
Chalepides barbatus (Fabricius,1787)   Brazil 3Dynastinae S.Am.20 (3)
Saint Barthélemy 2Dynastinae S.Am.20 (2)
St. Domingo 1Dynastinae S.Am.20 (1)
Unknown 7Dynastinae S.Am.20 (7)
Chalepides hydrophiloides (Burmeister,1847)   Brazil 2Dynastinae (2) S.Am.36
Charadronota quadrisignata (Gory & Percheron,1833)   Senegal 1Cetoniinae (1) Afr.28
Uganda 1Cetoniinae (1) Afr.18
Chasmatopterus hirtulus (Illiger,1803)   Spain 2Melolonthinae Pal.13 (2)
Chasmatopterus hirtulus hirtulus (Illiger,1803)   Spain 8Melolonthinae Pal.1 (8)
Chasmatopterus spp.    Spain 15Melolonthinae Pal.1 (15)
Chasmatopterus villosulus (Illiger,1803)   Portugal 1Melolonthinae Pal.1 (1)
Spain 1Melolonthinae Pal.13 (1)
Chasme decora (Wiedemann,1823)   South Africa 4Melolonthinae Afr.1 (4) 2 as C. nobilitata Burm
Chasmodia badia (Perty,1830)   Brazil 1Rutelinae (1) S.Am 6
Unknown 3Rutelinae (3) S.Am 6
Chasmodia bipunctata MacLeay,1819   Brazil 9Rutelinae (9) S.Am 6
Unknown 1Rutelinae (1) S.Am 6
Chasmodia dilatata Burmeister,1844   Ecuador 1Rutelinae (1) S.Am 6
Unknown 1Rutelinae (1) S.Am 6
Chasmodia emarginata (Gyllenhal,1817)   Brazil 5Rutelinae (5) S.Am 6
Unknown 3Rutelinae (3) S.Am 6
Chasmodia obscurata (Ohaus,1905)   Ecuador 1Rutelinae (1) S.Am 6
Cheirolasia burkei Westwood,1843   Namibia 2Cetoniinae Afr.8 (2)
Cheironitis eumenes (Gebler,1860)   Turkmenistan 1Scarabaeinae Pal.34 (1)
Cheironitis furcifer (Rossi,1792)   Greece 6Scarabaeinae Pal.14 (6)
Italy 9Scarabaeinae (9) Pal.14+23
Tunisia 6Scarabaeinae Pal.14 (6)
Turkey 3Scarabaeinae Pal.47 (2)
Pal.50 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Scarabaeinae Pal.14 (1)
Cheironitis haroldi Ballion,1870   Afghanistan 9Scarabaeinae (9) Pal.5+29
Armenia 1Scarabaeinae Pal.46 (1)
Cheironitis hoplosternus (Harold,1868)   South Africa 2Scarabaeinae Afr.38 (2)
Cheironitis irroratus (Rossi,1790)   Israel 1Scarabaeinae (1) Pal.24
Italy 10Scarabaeinae (10) Pal.13
Cheironitis klapperischi Balthasar,1955   Afghanistan 4Scarabaeinae (4) Pal.4+29
Cheironitis moeris (Pallas,1781)   Afghanistan 1Scarabaeinae Pal.5 (1)
Cheironitis pamphilus (Menetries,1849)   Afghanistan 44Scarabaeinae (44) Pal.5+29+30
Turkey 2Scarabaeinae Pal.13 (1)
Pal.47 (1)
Cheironitis phoebus Reitter,1893   Afghanistan 1Scarabaeinae Pal.5 (1)
Cheironitis scabrosus (Fabricius,1776)   Namibia 2Scarabaeinae Afr.38 (2)
South Africa 14Scarabaeinae (14) Afr.18+38
Cheironitis ungaricus (Herbst,1789)   Greece 12Scarabaeinae (12) Pal.13
Spain 2Scarabaeinae (2) Pal.13
Cheiroplatys excavatus Lea,1917   Australia 1Dynastinae Pac.2 (1)
Cheiroplatys latipes (Guerin,1830)   Australia 1Dynastinae Pac.2 (1)
Cheirotonus macleayi Hope,1840   India 3Euchirinae Or.2 (3)
Chiloloba acuta Wiedemann,1823   India 2Cetoniinae (2) Or.5
Unknown 4Cetoniinae (4) Or.5
Chilothorax alternatus (Horn,1870)   United States 1Washington
Aphodiinae N.Am.2 (1)
Chilothorax cervorum (Fairmaire,1871)   France 1Aphodiinae Pal.20 (1)
Chilothorax comma (Reitter,1892)   Mongolia 1Aphodiinae Pal.20 (1)
Russian Federation 1Aphodiinae (1) Pal.17
Chilothorax conspurcatus (Linnaeus,1758)   Sweden 137  Sk Bl Ha Sm Öl Ög Vg Sö Up Vr Dr Hs Jä Vb
Alan Dufberg 47 (29)
Aphodiinae Main (108)
Chilothorax distinctus (O.F.Müller,1776)   Bulgaria 1Aphodiinae Pal.29 (1)
Denmark 9Aphodiinae Main (9)
Germany 4Aphodiinae Pal.20 (3)
Pal.28 (1)
Sweden 762  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up Vr Dr Gä
Alan Dufberg 47 (32)
Aphodiinae Main (730)
Chilothorax exclamationis (Motschulsky,1849)   Tunisia 4Aphodiinae Pal.11 (4)
Chilothorax grafi (Reitter,1901)   Russian Federation 1Aphodiinae (1) Pal.17
Chilothorax hieroglyphicus (Klug,1845)   Algeria 1Aphodiinae Pal.7 (1)
Spain 23Canary Islands
Aphodiinae Pal.3 (12)
Pal.27 (8)
Pal.28 (3)
Tunisia 1Aphodiinae Pal.7 (1)
Chilothorax jacobsoni (Koshantshikov,1911)   Afghanistan 1Aphodiinae Pal.8 (1)
Chilothorax kukunorensis (Semenov,1897)   Afghanistan 1Aphodiinae (1) Pal.14
China 5Aphodiinae (5) Pal.14
Chilothorax leopardus (Horn,1870)   Canada 7Ontario
Aphodiinae N.Am.1 (7)
Chilothorax lineolatus (Illiger,1803)   Greece 12Aphodiinae (12) Pal.6+20
Italy 1Aphodiinae Pal.10 (1)
Spain 32Aphodiinae (32) Pal.2+6+20
Spain 10Mallorca
Aphodiinae Pal.24 (10)
Syria 1Aphodiinae Pal.20 (1)
Tunisia 4Aphodiinae Pal.6 (4)
Chilothorax melanostichtus (W.Schmidt,1840)   Afghanistan 2Aphodiinae Pal.8 (2)
Austria 2Aphodiinae (2) Pal.16+20
Bulgaria 7Aphodiinae Pal.7 (7)
Russian Federation 3Aphodiinae Pal.7 (3)
Turkey 1Aphodiinae Pal.7 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Aphodiinae Pal.10 (1)
Unknown 6Aphodiinae Pal.20 (6)
Chilothorax melanostictus (W. Schmidt,1840)   Germany 1Aphodiinae Pal.28 (1)
Chilothorax pardalis (LeConte,1857)   Canada 1British Columbia
Aphodiinae N.Am.2 (1)
United States 2California
Aphodiinae N.Am.5 (2)
Chilothorax paykulli (Bedel,1907)   Sweden 124  Skåne Blekinge Halland Småland Öland Gotland Östergötland Västergötland Uppland
Alan Dufberg 47 (16)
Aphodiinae Main (108)
Chilothorax pictus (Sturm,1805)   Denmark 18Aphodiinae Main (18)
Germany 1Aphodiinae Pal.16 (1)
Netherlands 1Aphodiinae (1) Pal.16
Sweden 54  Skåne Blekinge
Alan Dufberg 47 (6)
Aphodiinae Main (48)
Chilothorax scuticollis (Semenov,1898)   China 6Aphodiinae (6) Pal.13
Turkestan, Kazakhstan / Uzbekistan / Kyrgyzstan / Tajikistan / Turkmenistan / Uzbekistan 4Aphodiinae (4) Pal.11+20
Chilothorax spp.    Unknown 1Aphodiinae Pal.28 (1)
Chilothorax sticticus (Panzer,1798)   Germany 3Aphodiinae Pal.15 (2) as A. equestris
Pal.29 (1)
Sweden 173  Skåne Blekinge Småland Öland Gotland Bohuslän Södermanland Uppland Västmanland
Alan Dufberg 47 (31)
Aphodiinae Main (142)
Chilothorax subaeneus (LeConte,1857)   United States 2California
Aphodiinae N.Am.5 (2)
Chioneosoma baadeni (Brenske,1886)   Russian Federation 1Melolonthinae Pal.3 (1)
Chioneosoma komarowi (Brenske,1886)   Unknown 1Melolonthinae Pal.2 (1)
Chioneosoma lopatini Medvedev,1962   Uzbekistan 1Melolonthinae Pal.12 (1)
Chioneosoma porosum (Fischer von Waldheim,1823)   Kazakhstan 1Melolonthinae Pal.13 (1)
Unknown 2Melolonthinae Pal.3 (2)
Chioneosoma reitteri (Brenske,1887)   Russian Federation 2Melolonthinae Pal.3 (2)
Chioneosoma senex (Semenov,1902)   Russian Federation 1Melolonthinae Pal.3 (1)
Chioneosoma spp.    Uzbekistan 5Melolonthinae Pal.8 (5)
Chioneosoma subporosum Reitter,1902   Russian Federation 2Melolonthinae Pal.3 (2)
Chioneosoma tschitscherini (Semenov,1895)   Russian Federation 4Melolonthinae Pal.3 (4)
Chioneosoma turcumanum Brenske,1888   Uzbekistan 1Melolonthinae Pal.12 (1)
Chioneosoma vulpinum (Gyllenhal,1817)   Russian Federation 1Melolonthinae Pal.3 (1)
Chiron cylindrus (Fabricius,1798)   Sri Lanka 5Chironinae Or.1 (5)
Chiron spp.    Senegal 32Chironinae Afr.1 (32)
Sudan 1Chironinae Afr.1 (1)
Chlorocala africana blanda (Burmeister,1847)   Gambia 10Cetoniinae (10) Afr.4 (as Diplognatha blanda)

Chlorota cuprea Burmeister,1844   Brazil 1Rutelinae (1) S.Am 4
Chlorota limbaticollis Blanchard,1850   Panama 1Rutelinae (1) S.Am 4
Chnaunanthus discolor Burmeister,1844   United States 4Arizona
Melolonthinae N.Am.2 (4)
Chondropyga dorsalis (Donovan,1805)   Australia 1Queensland
Cetoniinae Pac.1 (1) as Diaphonia d.
Chondrorrhina abyssinica? Thoms.   Gambia 8Cetoniinae Afr.7 (8)
Senegal 2Cetoniinae Afr.7 (2)
Chromoptilia diversipes Westwood,1842   Madagascar 1Cetoniinae (1) Afr.18
Chromovalgus peyroni Mulsant,1852   Turkey 50Cetoniinae Pal.23 (9)
Pal.26 (4)
Pal.28 (13)
Pal.30 (15)
Pal.32 (1) as Hemivalgus peyroni
Pal.35 (8)
Unknown 2Cetoniinae Pal.26 (2)
Chrysina peruviana Kirby,1828   Mexico 1Rutelinae N.Am.1 (1)
Chrysophora chrysochlora (Latreille,1811)   Ecuador 2Rutelinae S.Am.1 (2)
Unknown 2Rutelinae S.Am.1 (2)
Circellium bacchus (Fabricius,1781)   South Africa 1Scarabaeinae Afr.52 (1)
South Africa 32Scarabaeinae Afr. (26) Afr.26+27+28
Afr.49 (5)
Afr.52 (1)
Unknown 3Scarabaeinae Afr.52 (3)
Clinteria atra Wiedemann,1823   Indonesia 1Java
Cetoniinae Or.3 (1)
Clinteria chloronota Blanchard,1850   Sri Lanka 38Cetoniinae Or.2 (38)
Clinteria imperialis (Paykull,1817)   Sri Lanka 1Cetoniinae Or.2 (1)
Unknown 2Cetoniinae Or.3 (2)
Clinteria imperialis valida Lansberge,1887   India 3Cetoniinae Or.3 (3)
Clinteroides permutans (Burmeister,1842)   Malawi 1Cetoniinae Afr.5 (1)
South Africa 3Cetoniinae (3) Afr.5+9
Clitopa rufiventris (Boheman,1857)   South Africa 1Melolonthinae Afr.3 (1)
Clypeodrepanus striatus (Boucomont,1921)   Kenya 1Slide
Slide Coleoptera 5 (1) Genital + parts of metathorax
Kenya 24Scarabaeinae Afr.58 (24)
Clypeodrepanus strigatus (Janssens,1953)   Kenya 12Scarabaeinae Afr.58 (12)
Clyster itys (Olivier,1789)   Unknown 4Dynastinae Or.2 (4)
Cnemida lacerata (Germar,1824)   Brazil 1Rutelinae (1) S.Am 4
Coelosis bicornis (Leske,1779)   Brazil 6Dynastinae (6) S.Am.1+5
Unknown 4Dynastinae (4) S.Am.1+5
Coelosis biloba (Linnaeus,1767)   Brazil 2Dynastinae S.Am.4 (2)
Colombia 5Dynastinae S.Am.4 (5)
Ecuador 1Dynastinae (1) S.Am.34
Peru 1Dynastinae (1) S.Am.31
Unknown 3Dynastinae S.Am.4 (3)
Coelosis inermis Sternberg,1908   Brazil 2Dynastinae S.Am.4 (2)
Coelosis sylvanus (Fabricius,1775)   Brazil 2Dynastinae S.Am.4 (2)
Coelotrachelus rudis (LeConte,1878)   United States 1Arizona
Aphodiinae N.Am.8 (1)
Coenochilus spp.    Kenya 1Cetoniinae Afr.32 (1) as C. hiloventris
Coilodera diardi (Gory & Percheron,1833)   Malaysia 1Cetoniinae Or.4 (1)
Coilodera helleri Miksic,1972   Borneo, Indonesia / Malaysia 1Cetoniinae Or.2 (1)
Coilodera trisulcata (Gory & Percheron,1833)   Indonesia 3Java
Cetoniinae Or.2 (3)
Colobopterus brignolii (Carpaneto,1973)   Greece 1Aphodiinae Pal.30 (1) as Aphodius brignolii
Turkey 9Aphodiinae Pal.11 (9)
Colobopterus erraticus (Linnaeus,1758)   Bulgaria 1Aphodiinae Pal.29 (1)
Cyprus 1Aphodiinae Pal.30 (1) as Aphodius erraticus
Italy 1Sicilia
Aphodiinae Pal.24 (1)
Sweden 309  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up
Alan Dufberg 47 (20)
Aphodiinae Main (289)
Turkey 10Aphodiinae Pal.4 (2)
Pal.27 (5)
Pal.28 (3)
Colobopterus quadratus (Reiche,1850)   Japan 1Aphodiinae Pal.18 (1)
Colobopterus quadrispinosus (Landin,1974)   Uganda 1Type Landin (1) Holotype Paratype 03486:1-2 of Colobopterus quadrispinosus Landin 1974
  Colobopterus quadrispinosus Landin 1974

Zimbabwe 1Type Landin (1) Holotype Paratype 03486:1-2 of Colobopterus quadrispinosus Landin 1974
  Colobopterus quadrispinosus Landin 1974

Conradtia principalis Kolbe,1892   Tanzania 1Cetoniinae (1) Afr.22
Copris acutidens Motschulsky,1860   Japan 5Scarabaeinae Pal.27 (5)
Copris amyntor Klug,1855   Botswana 3Scarabaeinae Afr.47 (3)
Ethiopia 2Scarabaeinae Afr.47 (2)
Copris anceus Olivier,1789   Tanzania 2Scarabaeinae Afr.48 (2)
Copris caelatus Fabricius,1794   Lesotho 1Scarabaeinae (1) Afr.33
South Africa 6Scarabaeinae (6) Afr.33
Copris elphenor Klug,1855   Namibia 3Scarabaeinae Afr.33 (2)
Afr.48 (1)
South Africa 7Scarabaeinae Afr.33 (7)
Tanzania 1Scarabaeinae Afr.48 (1)
Unknown 2Scarabaeinae Afr.48 (2)
Zimbabwe 1Scarabaeinae Afr.33 (1)
Copris evanidus Klug,1855   Ethiopia 2Scarabaeinae Afr.47 (2)
South Africa 16Scarabaeinae Afr.33 (16)
Copris fallaciosus Gillet,1907   South Africa 3Scarabaeinae (3) Afr.34
Zimbabwe 1Scarabaeinae (1) Afr.34
Copris fidius Olivier,1789   South Africa 5Scarabaeinae Afr.34 (3)
Afr.47 (2)
Copris fricator (Fabricius,1787)   India 1n/a
Scarabaeinae Or.7 (1)
Indonesia 4Java
Scarabaeinae Or.7 (4)
Ind. Or. 1Scarabaeinae Or.7 (1)
Copris harrisi Waterhouse,1891   Kenya 22Scarabaeinae (22) Afr.9+11
Copris hispanus (Linnaeus,1764)   Afghanistan 4Scarabaeinae (4) Pal.5+33
Algeria 6Scarabaeinae Pal.17 (4)
Pal.37 (2)
Cameroon 1Scarabaeinae Afr.46 (1)
Croatia 1Scarabaeinae Pal.37 (1)
Greece 3Scarabaeinae Pal.37 (1)
Pal.50 (1)
Pal.53 (1) as C. hispaniola (?)
Israel 1Scarabaeinae Pal.46 (1)
Italy 5Scarabaeinae (5) Pal.17+23
Italy 1Sardinia
Scarabaeinae Pal.44 (1)
Italy 1Sicily
Scarabaeinae Pal.37 (1)
Libya 4Scarabaeinae Pal.44 (4)
Spain 11Scarabaeinae (9) Pal.17+27
Pal.37 (2)
Spain 2Mallorca
Scarabaeinae Pal.37 (2)
Syria 1Scarabaeinae Pal.37 (1)
Tunisia 15Scarabaeinae (10) Pal.17+18+26
Pal.37 (3)
Pal.46 (1)
Pal.50 (1)
Turkey 3Scarabaeinae Pal.49 (3)
Turkmenistan 1Scarabaeinae Pal.37 (1)
Unknown 13Scarabaeinae Pal.37 (13)
Copris hispanus cavolinii (Petagna,1792)   Cyprus 1Scarabaeinae Pal.18 (1)
Greece 6Scarabaeinae Pal.18 (6)
Copris incertus Say,1835   Ecuador 2Scarabaeinae S.Am.5 (2)
Guatemala 2Scarabaeinae S.Am.8 (2)
Copris integer Reiche,1847   Kenya 2Scarabaeinae Afr.48 (2)
Tanzania 5Scarabaeinae Afr.48 (5)
Copris jacchus Fabricius,1775   Lesotho 3Scarabaeinae (3) Afr.34
South Africa 2Scarabaeinae (2) Afr.34
Copris lugubris Boheman,1858   Costa Rica 1Scarabaeinae S.Am.2 (1)
Honduras 1Scarabaeinae S.Am.2 (1)
Copris lunaris (Linnaeus,1758)   Austria 5Scarabaeinae (5) Pal.18+24+35
Bulgaria 5Scarabaeinae (5) Pal.18
Croatia 2Scarabaeinae (2) Pal.24
Czechia 1Scarabaeinae Pal.53 (1)
Denmark 1Scarabaeinae (1)
Germany 1Scarabaeinae (1) Pal.33
Italy 1Scarabaeinae (1) Pal.24
Romania 2Scarabaeinae (2) Pal.18
Sweden 262  Skåne Blekinge Öland Gotland
Alan Dufberg 47 (6)
Scarabaeinae Main (256)
Switzerland 1Scarabaeinae (1) Pal.24
Turkey 1Scarabaeinae Pal.46 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 3Scarabaeinae (3) Pal.18
Copris mesacanthus Harold,1878   Namibia 1Scarabaeinae (1) Afr.34
South Africa 9Scarabaeinae (9) Afr.34
Copris misellus Peringuey,1901   South Africa 6Scarabaeinae (6) Afr.34
Copris obesus Boheman,1857   Lesotho 1Scarabaeinae (1) Afr.35
Copris ochus (Motschulsky,1860)   Japan 2Scarabaeinae (2) Pal.27
Copris orion Klug,1835   South Africa 6Scarabaeinae (6) Afr.34
Copris orphanus Guerin,1849   Kenya 4Scarabaeinae (4) Afr.11
Namibia 2Scarabaeinae (2) Afr.34
Copris pueli Boissy,1905   Tunisia 1Scarabaeinae (1) Pal.18
Copris pumilionis Wallengren,1881   South Africa 1Type Coleoptera 2769 (1) Typus Syntype 02769:1 of Copris pumilionis Wallengren 1881
  Copris pumilionis Wallengren 1881

Copris reflexus F. (Microcopris)   Indonesia 2Java
Scarabaeinae Or.2 (2)
Malaysia 1Scarabaeinae Or.2 (1)
Thailand 1Scarabaeinae Or.2 (1)
Copris sinon Olivier,1789   Gambia 4Scarabaeinae (4) Afr.12
Namibia 1Scarabaeinae (1) Afr.12
Senegal 5Scarabaeinae (5) Afr.12
Copris spp.    Cameroon 1Scarabaeinae (1) Afr.12
Ethiopia 2Scarabaeinae Afr.7 (2)
Gambia 1Scarabaeinae (1) Afr.12
Guatemala 1Scarabaeinae S.Am.6 (1)
Guinea-Bissau 2Scarabaeinae (2) Afr.7+12
Honduras 3Scarabaeinae (3) S.Am.2+5
Indonesia 2Java
Scarabaeinae Or.3 (2)
Indonesia 4Java
Scarabaeinae Or.2 (4)
Indonesia 1Java
Scarabaeinae Or.3 (1)
Kenya 11Scarabaeinae (11) Afr.12
Mexico 1Scarabaeinae S.Am.6 (1)
Rwanda 1Scarabaeinae (1) Afr.12
Senegal 2Scarabaeinae (2) Afr.12
South Africa 7Scarabaeinae (7) Afr.12
Sri Lanka 1Scarabaeinae Or.3 (1)
Tanzania 6Scarabaeinae (6) Afr.12+45
Thailand 6Scarabaeinae Or.3 (6)
Thailand 10Scarabaeinae (10) Or.2+3
Turkey 2Scarabaeinae Pal.34 (2)
Unknown 2Scarabaeinae Afr.48 (2)
Zambia 1Scarabaeinae (1) Afr.12
Copris tripartitus Waterhouse,1875   Korea, Republic of 1Scarabaeinae (1) Pal.18
Copris umbilicatus Abeille,1901   Italy 1Scarabaeinae Pal.34 (1)
Copris victorini Boheman,1857   South Africa 4Scarabaeinae (4) Afr.35
Copris wieswei Kolbe,1914   South Africa 1Scarabaeinae (1) Afr.35
Coprophanaeus ensifer (Germar,1821)   Brazil 2Scarabaeinae (2) S.Am.4
Coprophanaeus jasius (Olivier,1789)   Brazil 2Scarabaeinae S.Am.4 (2)
Coprophanaeus lancifer (Linnaeus,1767)   Brazil 2Scarabaeinae S.Am.4 (2)
Coprophanaeus morenoi Arnaud,1982   Ecuador 2Scarabaeinae (2) S.Am.4
Coprophanaeus spitzi (Pessoa,1934)   Brazil 2Scarabaeinae (2) S.Am.4
Coprophanaeus spp.    Honduras 1Scarabaeinae S.Am.2 (1)
Coptochirus brachyperus Harold,1868   South Africa 1Aphodiinae Afr.24 (1)
Coptochirus cf. perlucidus Bordat,1989   South Africa 3Aphodiinae Afr.32 (3)
Coptochirus emarginatus Germar,1824   South Africa 1Aphodiinae Afr.24 (1)
Coptochirus vulgatus Harold,1868   South Africa 4Aphodiinae (4) Afr.16+24
Coptodactyla subaenea Harold,1877   Australia 3Scarabaeinae Pac.1 (3)
Coptomia granulata Burmeister,1842   Madagascar 1Cetoniinae (1) Afr.18
Coptomia mauritania Gory & Percheron,1833   Madagascar 1Cetoniinae (1) Afr.18 (as C. fasciata)

Coptorhina klugi Hope,1830   South Africa 4Scarabaeinae Afr.33 (4)
Coptorhina nitidipennis Boheman,1857   South Africa 1Scarabaeinae Afr.33 (1)
Corynotrichius bicolor Kolbe,1892   Cameroon 32Cetoniinae Afr.31 (32)
Corythoderus loripes Klug,1845   Senegal 1Aphodiinae (1) Afr.13
Corythoderus marshalli Brauns,1900   South Africa 1Aphodiinae (1) 1 cotype Afr.20
Cosmesthes lineaticollis Kraatz,1880   Cameroon 6Cetoniinae (6) Afr.26
Cosmiomorpha modesta Saunders,1872   China 3Cetoniinae (3) Pal.17
Cosmiophaena impar (Gory & Percheron,1833)   Côte D'ivoire 1Cetoniinae (1) Afr.18
Cosmiophaena rubescens (Brancsik,1914)   Zambia 2Cetoniinae Afr.5 (2)
Cotalpa crassa Blanchard,1850   Unknown 3Rutelinae (3) S.Am 1
Cotalpa lanigera (Linnaeus,1764)   United States 1Connecticut
Rutelinae N.Am.1 (1)
United States 2Michigan
Rutelinae N.Am.1 (2)
United States 1New York
Rutelinae N.Am.1 (1)
America borealis, United States/Canada 11Rutelinae N.Am.1 (11)
Cotinis mutabilis (Gory & Percheron,1833)   Guatemala 1Cetoniinae S.Am.1 (1)
Mexico 1Cetoniinae S.Am.1 (1)
Cremastocheilus angularis Leconte,1857   United States 2California
Cetoniinae N.Am.3 (2)
Cremastocheilus castaneae Knoch,1801   Unknown 2Cetoniinae N.Am.3 (2)
Cremastocheilus crinitus Leconte,1874   United States 1Washington
Cetoniinae N.Am.1 (1)
Cremastocheilus knochii Leconte,1853   United States 1Montana
Cetoniinae N.Am.3 (1)
Cremastocheilus spp.    Canada 1Nova Scotia
Cetoniinae N.Am.1 (1)
Unknown 2Cetoniinae N.Am.3 (2)
Cryptocanthon brevisetosus Howden,1973   Guatemala 7Scarabaeinae S.Am.2 (7)
Cryptodus matthewsi Endrödi,1976   Australia 1Dynastinae Pac.2 (1)
Cryptodus paradoxus Macleay,1819   Australia 1Dynastinae Pac.2 (1)
Cryptodus spp.    Australia 5Dynastinae Pac.2 (5)
Cyclidius elongatus (Olivier,1789)   Ecuador 3Cetoniinae S.Am.1 (3)
Cyclocephala amazona (Linnaeus,1767)   Colombia 8Dynastinae S.Am.18 (8)
Panama 2Dynastinae S.Am.18 (2)
Cyclocephala borealis Arrow,1911   United States 1Arizona
Dynastinae N.Am.4 (1)
United States 35Maryland
Dynastinae N.Am.4 (35)
Cyclocephala castanea (Olivier,1789)   Suriname 1Dynastinae S.Am.18 (1)
Unknown 2Dynastinae S.Am.18 (2)
Cyclocephala discolor (Hebst,1792)   Colombia 5Dynastinae S.Am.18 (5)
Cyclocephala flavipennis Arrow,1914   Brazil 2Dynastinae S.Am.19 (2)
Cyclocephala fulgurata Burmeister,1847   Brazil 1Dynastinae S.Am.19 (1)
Cyclocephala gregaria Heyne & Taschenberg,1907   Colombia 2Dynastinae S.Am.18 (2)
Unknown 2Dynastinae S.Am.18 (2)
Cyclocephala halffteriana Martinez,1968   Mexico 3Dynastinae N.Am.4 (3)
Cyclocephala hardyi Endrödi,1975   Brazil 3Dynastinae (3) S.Am.18 (incl. 3 paratypes)

Cyclocephala immaculata (Olivier,1789)   United States 6Kansas
Dynastinae N.Am.2 (6)
Cyclocephala longula LeConte,1863   United States 4California
Dynastinae N.Am.3 (4)
Cyclocephala lunulata Burmeister,1847   Brazil 3Dynastinae S.Am.19 (3)
Unknown 1Dynastinae S.Am.19 (1)
Cyclocephala lurida (Bland,1863)   United States 4West Virginia
Dynastinae N.Am.4 (4)
Cyclocephala marginalis Kirsch,1870   Unknown 1Dynastinae S.Am.18 (1)
Cyclocephala melanocephala (Fabricius,1775)   Argentina 5Dynastinae S.Am.19 (5)
Brazil 5Dynastinae S.Am.19 (5)
Colombia 3Dynastinae S.Am.19 (3)
Unknown 1Dynastinae S.Am.19 (1)
Cyclocephala metrica Steinheil,1874   Brazil 1Dynastinae S.Am.19 (1)
Cyclocephala modesta Burmeister,1847   Brazil 3Dynastinae S.Am.17 (3)
Cyclocephala notata (Illiger,1806)   Dominican Republic 3Dynastinae S.Am.18 (3)
Cyclocephala ocellata Burmeister,1847   Guyana 2Dynastinae (2) S.Am.32
Cyclocephala pubescens Burmeister,1847   Brazil 1Dynastinae S.Am.19 (1)
Cyclocephala putrida Burmeister,1847   Argentina 10Dynastinae S.Am.19 (10)
Unknown 2Dynastinae S.Am.19 (2)
Cyclocephala rorulenta Höhne,1923   Unknown 1Dynastinae S.Am.18 (1)
Cyclocephala sexpunctata Castelnau,1840   Brazil 1Dynastinae S.Am.18 (1)
Colombia 2Dynastinae S.Am.18 (2)
Cyclocephala signaticollis Burmeister,1847   Colombia 1Dynastinae S.Am.19 (1)
Cyclocephala stictica Burmeister,1847   Unknown 2Dynastinae S.Am.18 (2)
Cyclocephala tarsalis Dechambre,1979   Guyana 2Dynastinae (2) S.Am.32
Cyclocephala toulgoeti Dechambre,1992   Guyana 2Dynastinae (2) S.Am.32
Cyclocephala tridentata (Fabricius,1801)   Guadeloupe 1Dynastinae (1) S.Am.36
Cyclocephala tucumana Brethes,1904   Argentina 3Dynastinae S.Am.20 (3)
Cyclocephala tutilina Burmeister,1847   Venezuela 2Dynastinae S.Am.18 (2)
Cyclocephala tylifera Höhne,1923   Brazil 2Dynastinae S.Am.18 (2)
Cyclocephala undata (Olivier,1789)   Guyana 2Dynastinae (2) S.Am.32
Cyclocephala weidneri Endrödi,1964   Guatemala 2Dynastinae S.Am.19 (2)
Cymophorus leucostictus Schaum,1841   Ethiopia 2Cetoniinae (2) Afr.27
Cymophorus undatus Kirby,1827   South Africa 2Cetoniinae (2) Afr.17
Cyphonistes vallatus (Wiedemann,1823)   South Africa 2Dynastinae Afr.5 (2)
Tanzania 2Dynastinae Afr.5 (2)
Cyphonoxia brenskei Reitter,1895   Afghanistan 1Melolonthinae Pal.10 (1)
Cyprolais aurata (Westwood,1841)   Cameroon 1Cetoniinae (1) Afr.18
Cyprolais cornuta Heath,1904   Tanzania 1Cetoniinae (1) Afr.18
Cyptochirus ambiguus Kirby,1828   South Africa 6Scarabaeinae Afr.58 (6)
Cyriopertha arcuata (Gebler,1832)   China 1Rutelinae (1) Pal.6 (as Phylloperta a.)

Cyriopertha glabra (Gebler,1841)   China 1Rutelinae (1) Pal.6 (as Phyllopertha g.)

Turkestan, Kazakhstan / Uzbekistan / Kyrgyzstan / Tajikistan / Turkmenistan / Uzbekistan 2Rutelinae (2) Pal.6 (as Phyllopertha g.)

Uzbekistan 1Rutelinae Pal.10 (1)
Cyrtothyrea marginalis (Swartz,1817)   Ethiopia 4Cetoniinae (4) Afr.27
South Africa 61Cetoniinae (61) Afr.16 (as Oxythyrea m.)
Unknown 4Cetoniinae (4) Afr.27
Cyrtothyrea testaceoguttata Blanchard,1850   Lesotho 2Cetoniinae (2) Afr.16 (as Oxythyrea t.)
South Africa 23Cetoniinae (23) Afr.16 (as Oxythyrea t.)
Danielssonia minuta Bordat & Howden,1995   Malaysia 1Type Coleoptera 3036 (1) Holotype 03040:1 of Danielssonia minuta Bordat & Howden 1995
  Danielssonia minuta Bordat & Howden 1995

Dasygnathus globosus Arrow,1914   Australia 2Dynastinae Pac.2 (2)
Dasygnathus trituberculatus Blackburn,1889   Australia 2Dynastinae Pac.2 (2)
Deltochilum brasiliense Castelnau,1840   Brazil 1Scarabaeinae S.Am.11 (1)
Deltochilum dentipes Eschscholtz,1822   Brazil 3Scarabaeinae S.Am.11 (3)
Deltochilum furcatum Castelnau,1840   Brazil 1Scarabaeinae S.Am.11 (1)
Deltochilum gibbosum Fabricius,1775   Brazil 1Scarabaeinae S.Am.11 (1)
Deltochilum lobipes Bates,1887   Mexico 1Scarabaeinae N.Am.1 (1)
Deltochilum mexicanum Burmeister,1848   Honduras 9Scarabaeinae S.Am.3 (9)
Deltochilum spp.    Colombia 1Scarabaeinae S.Am.8 (1)
Ecuador 1Scarabaeinae S.Am.1 (1)
Unknown 2Scarabaeinae S.Am.8 (2)
Democrates burmeisteri Reiche,1852   Ecuador 2Dynastinae S.Am.18 (2)
Dendropaemon denticollis Felsche,1909   Brazil 1Scarabaeinae S.Am.4 (1)
Dendropaemon renatii Olsoufieff,1924   Brazil 1Scarabaeinae S.Am.4 (1)
Dendropaemon vazdemelloi Génier & Arnaud,2016   Brazil 1Type Coleoptera 5094 (1) Paratype 05343:2 of Dendropaemon vazdemelloi Génier & Arnaud 2016
  Dendropaemon vazdemelloi Génier & Arnaud 2016

Denticnema morbillosa (Burmeister,1844)   South Africa 2Hopliini Afr.6 (2)
Denticnema squamosa (Burmeister,1844)   South Africa 3Hopliini Afr.6 (3)
Denticnema striata (Burmeister,1844)   South Africa 3Hopliini Afr.6 (3)
Diabroctis mimas (Linnaeus,1758)   Brazil 1Scarabaeinae (1) S.Am.4
Peru 2Scarabaeinae S.Am.1 (2)
Dialithus magnificus Parry,1849   Mexico 2Cetoniinae S.Am.1 (2)
Dialithus scintillans Howden,   Colombia 2Cetoniinae S.Am.1 (2)
Dialytellus humeralis (LeConte,1878)   United States 1North Carolina
Aphodiinae N.Am.2 (1)
Dialytes striatulus (Say,1825)   Canada 1Quebec
Aphodiinae (1) N.Am.2 (6-N.Am.2)

Dialytes ulkei Horn,1875   United States 1Virginia
Aphodiinae N.Am.2 (1)
Diaplochelus crassipes Burmeister,1844   South Africa 1Hopliini (1) Afr.12
Diaplochelus longipes Fabricius,1787   South Africa 3Hopliini (3) Afr.12
Diaplochelus squamulatus Burmeister,1844   South Africa 1Hopliini (1) Afr.12=paratype of transvaalensis Per.?
Diastellopalpus johnstoni Waterhouse,1885   Tanzania 2Scarabaeinae (2) Afr.18
Diastellopalpus lamellicollis Quedenfeldt,1884   Malawi 1Scarabaeinae (1) Afr.18
Diastellopalpus spp.    Guinea-Bissau 1Scarabaeinae Afr.3 (1)
Diastellopalpus thompsoni (Bates,1888)   Malawi 1Scarabaeinae (1) Afr.18
Diastellopalpus tridens (Fabricius,1781)   Gambia 2Scarabaeinae (2) Afr.18
Guinea-Bissau 1Scarabaeinae (1) Afr.18
Diastictus vulneratus (Sturm,1805)   Estonia 1Aphodiinae (1) Pal.15
Germany 4Aphodiinae (4) Pal.13
Sweden 110  Skåne Öland Gotland
Alan Dufberg 48 (15)
Aphodiinae Main (95)
Diathermus globosus Schoch,1897   Lesotho 2Cetoniinae (2) Afr.15
South Africa 2Cetoniinae (2) Afr.15
Dicaulocephalus fruhstorferi Felsche,1901   Thailand 1Rutelinae Or.8 (1)
Dichecephala ovata (Fåhraeus in Boheman, 1857)   South Africa 3Melolonthinae Afr.3 (3)
Dichelonyx backi (Kirby,1837)   Canada 2British Columbia
Melolonthinae N.Am.2 (2)
United States 4Oregon
Melolonthinae N.Am.2 (4)
Dichelonyx canadensis (Horn,1876)   Canada 8Quebec
Melolonthinae N.Am.2 (8)
Dichelonyx elongatula (Schönherr,1817)   Canada 1Quebec
Melolonthinae N.Am.2 (1)
United States 2New York
Melolonthinae N.Am.2 (2)
United States 6Virginia
Melolonthinae N.Am.2 (6)
Dichelonyx elongatula Schoenherr,1817   United States 2California, Pennsylvania
Melolonthinae N.Am.3 (2) as Dichelonycha elongata F.
Dichelonyx fulgida (LeConte,1856)   United States 4California
Melolonthinae N.Am.2 (4)
Dichelonyx robusta (Fall,1901)   United States 1Oregon
Melolonthinae N.Am.2 (1)
Dichelonyx subvittata (LeConte,1856)   United States 1Alabama
Melolonthinae N.Am.1 (1)
Dichelonyx truncata (LeConte,1856)   United States 5California
Melolonthinae (5) N.Am.2+3
Dichelonyx vaga (Fall,1901)   United States 1California
Melolonthinae N.Am.2 (1)
Dichelonyx valida (LeConte,1856)   United States 1Arizona
Melolonthinae N.Am.2 (1)
United States 3Oregon
Melolonthinae N.Am.2 (3)
United States 1Washington
Melolonthinae N.Am.2 (1)
Dichelus acanthopus Burmeister,1844   South Africa 2Hopliini Afr.8 (2)
Dichelus dentipes Fabricius,1781   South Africa 95Hopliini (95) Afr.7+8
Dichelus duplosquamosus Schein,1958   South Africa 1Hopliini Afr.8 (1)
Dichelus expansus Peringuey,1902   South Africa 2Hopliini Afr.8 (2)
Dichelus holosericeus Burmeister,1844   South Africa 2Hopliini Afr.8 (2)
Dichelus laticollis Burmeister,1844   South Africa 2Hopliini Afr.7 (2)
Dichelus villosus Burmeister,1844   South Africa 2Hopliini Afr.8 (2)
Dichelus villosus ssp.luteopygus Burmeister,1844   South Africa 2Hopliini Afr.8 (2)
Dichelus villosus ssp.minor Burmeister,1844   South Africa 1Hopliini (1) Type 2015:2
Dicheros bicornis borneensis Nagai,1985   Borneo, Indonesia / Malaysia 1Cetoniinae Or.4 (1)
Dicheros decorus Gory & Percheron,1833   Indonesia 2Java
Cetoniinae Or.2 (2)
Dicheros spp.    Indonesia 1Nias
Cetoniinae (1) Or.4
Dichodontus croesus (Fabricius,1801)   Indonesia 1Java
Dynastinae Or.2 (1)
Dichodontus grandis Ritsema,1882   Borneo, Indonesia / Malaysia 2Dynastinae Or.9 (2)
Dichotomius amplicollis Harold,1869   Brazil 1Scarabaeinae S.Am.12 (1)
Dichotomius assifer (Eschscholtz,1822)   Brazil 2Scarabaeinae S.Am.3 (2)
Dichotomius bicuspis Germar,1824   Brazil 4Scarabaeinae S.Am.12 (4)
Dichotomius bos (Blanchard,1843)   Brazil 4Scarabaeinae S.Am.3 (2)
S.Am.13 (2)
Unknown 1Scarabaeinae S.Am.13 (1)
Dichotomius carbonarius Mannerheim,1829   Brazil 1Scarabaeinae S.Am.12 (1)
Unknown 1Scarabaeinae S.Am.12 (1)
Dichotomius carolinus (Linnaeus,1767)   Guatemala 1Scarabaeinae S.Am.5 (1)
Mexico 5Scarabaeinae S.Am.13 (5)
America borealis, United States/Canada 2Scarabaeinae S.Am.13 (2)
Unknown 9Scarabaeinae S.Am.13 (9)
South America 1Scarabaeinae S.Am.13 (1)
Dichotomius crinicollis Germar,1824   Brazil 2Scarabaeinae S.Am.12 (2)
Paraguay 1Scarabaeinae S.Am.12 (1)
Dichotomius divergens Luedereald,1923   Ecuador 1Scarabaeinae (1) S.Am.4
Dichotomius fissus (Harold,1867)   Brazil 3Scarabaeinae S.Am.3 (2)
S.Am.12 (1)
Dichotomius laevicollis Felsche,1901   Unknown 1Scarabaeinae S.Am.12 (1)
Dichotomius lucasi Harold,1869   Guadeloupe 1Scarabaeinae S.Am.6 (1)
Dichotomius mormon (Ljungh,1799)   Brazil 10Scarabaeinae S.Am.3 (2)
S.Am.12 (7)
Type Coleoptera 2718 (1) Typus Syntype 02720:1 of Scarabaeus mormon Ljungh 1799
  Scarabaeus mormon Ljungh 1799

Unknown 6Scarabaeinae S.Am.12 (6)
Dichotomius nisus (Olivier,1789)   Brazil 3Scarabaeinae S.Am.3 (2)
S.Am.11 (1)
French Guiana 2Scarabaeinae S.Am.11 (2)
Dichotomius prietoi Martinez & Martinez,1982   Ecuador 2Scarabaeinae S.Am.4 (2)
Dichotomius problematicus Luederwaldt,1924   Ecuador 2Scarabaeinae S.Am.3 (2)
Dichotomius satanas Harold,1867   Colombia 1Scarabaeinae S.Am.11 (1)
Dichotomius schiffleri Vaz-de-Mello & Louzada & Gavino,2001   Brazil 4Scarabaeinae (4) 4 paratypes S.Am.2
Dichotomius semissquamosus (Curtis,1845)   Brazil 1Scarabaeinae S.Am.2 (1)
Dichotomius spp.    ? 1Scarabaeinae S.Am.7 (1)
Brazil 2Scarabaeinae S.Am.13 (2)
Costa Rica 1Scarabaeinae S.Am.1 (1)
Ecuador 4Scarabaeinae (4) S.Am.1 +7
Guatemala 7Scarabaeinae (7) S.Am.6+7
Kenya 1Scarabaeinae Afr.53 (1)
Mexico 1Scarabaeinae S.Am.7 (1)
South Africa 1Scarabaeinae Afr.53 (1)
Dichotomius telamon Harold,1869   Suriname 1Scarabaeinae S.Am.11 (1)
Unknown 1Scarabaeinae S.Am.11 (1)
Dichotomius torulosus (Eschscholtz,1822)   Chile 11Scarabaeinae S.Am.14 (7)
Type Coleoptera (4) Typus Holotype Paratype 02775:1-4 of Pinotus dahli Landin 1955
  Pinotus dahli Landin 1955

Dicranocnemus andreaei Schein,1958   South Africa 1Type Schein (1) Paratype 02034:2 of Dicranocnemus andreaei Schein 1958
  Dicranocnemus andreaei Schein 1958

Dicranocnemus hirtipes Schein,1958   South Africa 1Type Schein (1) Paratype 02035:2 of Dicranocnemus hirtipes Schein 1958
  Dicranocnemus hirtipes Schein 1958

Dicranocnemus hypocrita Peringuey,1902   South Africa 3Hopliini (3) Afr.12
Dicranocnemus mendicus Peringuey,1902   South Africa 2Hopliini (2) Afr.12
Dicranocnemus pulcher Peringuey,1902   South Africa 3Hopliini (3) Afr.12
Dicranocnemus squamosus Burmeister,1844   South Africa 2Hopliini (2) Afr.12
Dicranocnemus squamulatus Burmeister,1844   South Africa 3Hopliini (3) Afr.12
Dicranocnemus sulcicollis Schein,1958   South Africa 4Hopliini (4) Afr.12
Dicranocnemus sulcicollis ssp.fürschi Schein,1958   South Africa 2Hopliini (2) Afr.12
Dicranoplia deserticola (Lucas,1859)   Algeria 1Rutelinae Pal.8 (1)
Dicronocephalus adamsi (Pascoe,1863)   China 1Cetoniinae (1) Pal.20
Dicronorrhina derbyana Westwood,1843   Namibia 1Cetoniinae Afr.2 (1)
Namibia 11Cetoniinae (11) Afr.7+8 (as ssp. layaardi Peringuey)

South Africa 2Cetoniinae Afr.7 (2)
Digitonthophagus gazella (Fabricius,1787)   Sierra Leone 1Scarabaeinae Afr.3 (1)
Diloboderus abderus (Sturm,1826)   Argentina 5Dynastinae S.Am.17 (5)
Brazil 8Dynastinae S.Am.17 (8)
Paraguay 2Dynastinae (2) S.Am.30
Unknown 8Dynastinae S.Am.17 (8)
Dilortomaeus excultus (Péringuey,1908)   Zambia 1Aphodiinae Afr.30 (1)
Diorygopyx asciculifer Matthews,1974   Australia 1New South Wales
Scarabaeinae Pac.1 (1)
Dipelicus hircus (Fabricius,1775)   Sri Lanka 1Dynastinae Or.3 (1)
Dipelicus optatus (Sharp,1875)   Australia 2Dynastinae Pac.2 (2)
Diplognatha gagates (Forster,1771)   Cameroon 3Cetoniinae (3) Afr.28
Congo, the democratic republic of the 3Cetoniinae (3) Afr.28
Ethiopia 1Cetoniinae (1) Afr.28
France 2Cetoniinae Pal.25 (2)
Malawi 1Cetoniinae Afr.4 (1)
Mozambique 1Cetoniinae Afr.4 (1)
Sierra Leone 1Cetoniinae (1) Afr.28
South Africa 6Cetoniinae (6) Afr.4+5+28
Tanzania 4Cetoniinae (4) Afr.5+28
Uganda 1Cetoniinae (1) Afr.28
Diplognatha gagates silicea (Forster,1771)   Cameroon 1Cetoniinae Afr.29 (1)
Namibia 2Cetoniinae (2) Afr.17
South Africa 6Cetoniinae (6) Afr.17
Tanzania 7Cetoniinae Afr.29 (7)
Unknown 6Cetoniinae Afr.29 (6)
Diplognatha striata (Janson,1877)   Malawi 1Cetoniinae (1) Afr.5 (as Apocnosis s.)

Diplognatha viridula Janson,1877   Congo 1Cetoniinae (1) Afr.18
Diplotaxis abnormis Fall,1909   United States 2Arizona
Melolonthinae N.Am.3 (2)
Diplotaxis ambigua Fall,1909   United States 2Arizona
Melolonthinae N.Am.3 (2)
Diplotaxis angularis LeConte,1856   Mexico 2Melolonthinae N.Am.3 (2)
Diplotaxis arctifrons Bates,1888   United States 2New Mexico
Melolonthinae N.Am.3 (2)
Diplotaxis arizonica Schaeffer,1907   United States 2Arizona
Melolonthinae N.Am.3 (2)
Diplotaxis atlantis Fall,1909   Canada 2Ontario
Melolonthinae N.Am.2 (2)
Diplotaxis boops Bates,1888   United States 2New Mexico
Melolonthinae N.Am.3 (2)
Diplotaxis brevicollis LeConte,1856   Canada 1British Columbia
Melolonthinae N.Am.2 (1)
Diplotaxis brevidens LeConte,1856   United States 5Arizona
Melolonthinae N.Am.1 (5)
Diplotaxis carbonata LeConte,1856   Mexico 2Melolonthinae N.Am.3 (2)
Diplotaxis chiricahuae Fall,1909   Mexico 1Melolonthinae N.Am.3 (1)
United States 1Arizona
Melolonthinae N.Am.3 (1)
Diplotaxis conformis Fall,1909   United States 2Arizona
Melolonthinae N.Am.3 (2)
Diplotaxis cribulosa LeConte,1856   United States 1Arizona
Melolonthinae N.Am.2 (1)
Diplotaxis dentella Fall,1909   United States 2Arizona
Melolonthinae N.Am.2 (2)
Diplotaxis fimbriata Fall,1909   United States 1California
Melolonthinae N.Am.2 (1)
Diplotaxis frondicola (Say,1825)   United States 1Maryland
Melolonthinae N.Am.3 (1)
United States 1Texas
Melolonthinae N.Am.3 (1)
Diplotaxis fulvus (LeConte,1856)   United States 1Maryland
Melolonthinae N.Am.2 (1)
Diplotaxis haydenii LeConte,1856   United States 1New Mexico
Melolonthinae N.Am.3 (1)
Diplotaxis ingenua Fall,1909   United States 2Arizona
Melolonthinae N.Am.3 (2)
Diplotaxis knausii Schaeffer,1907   United States 2Arizona
Melolonthinae N.Am.3 (2)
Diplotaxis liberta (Germar,1824)   United States 1North Carolina
Melolonthinae N.Am.3 (1)
Diplotaxis magna Bates,1888   United States 3Arizona
Melolonthinae N.Am.3 (3)
Diplotaxis marginicollis Fall,1909     United States 2Arizona
Melolonthinae N.Am.3 (2)
Diplotaxis megapleura Vaurie,1960   Mexico 1Melolonthinae N.Am.3 (1)
Unknown 1Melolonthinae N.Am.3 (1)
Diplotaxis moerens LeConte,1856   United States 1Arizona
Melolonthinae N.Am.3 (1)
Diplotaxis obscura LeConte,1859   United States 1New Mexico
Melolonthinae N.Am.3 (1)
Diplotaxis parallela Fall,1909   United States 2Arizona
Melolonthinae N.Am.3 (2)
Diplotaxis rufiola Fall,1909   United States 12Texas
Melolonthinae N.Am.3 (12)
Diplotaxis sordida (Say,1825)   Canada 3Ontario
Melolonthinae N.Am.3 (3)
United States 2Massachusetts
Melolonthinae N.Am.3 (2)
Diplotaxis subangulata LeConte,1856   Mexico 1Melolonthinae N.Am.2 (1)
United States 1Texas
Melolonthinae N.Am.2 (1)
Diplotaxis tristis Kirby,1837   Canada 1Ontario
Melolonthinae N.Am.3 (1)
Diplotaxis truncatula LeConte,1856   Unknown 2Melolonthinae N.Am.3 (2)
Dischista cincta (DeGeer,1778)   Central African Republic 5Cetoniinae (5) Afr.24
Ethiopia 2Cetoniinae (2) Afr.24
Namibia 7Cetoniinae (7) Afr.5+10+24
South Africa 10Cetoniinae (10) Afr.5+10+24
Unknown 6Cetoniinae (6) Afr.24
Zimbabwe 1Cetoniinae (1) Afr.24
Dolichostethus levis (Janson,1877)   South Africa 1Cetoniinae (1) Afr.6 (as Dulcinea l.)

Drepanocanthus brevitarsis Peringuey,1901   South Africa 4Aphodiinae Afr.24 (4)
Drepanocanthus lineatus (Wiedemann,1823)   South Africa 41Aphodiinae Afr.24 (4)
Afr.31 (37)
Drepanocanthus mineti Bordat,1993   South Africa 2Aphodiinae Afr.34 (2)
Drepanocerus kirbyi Kirby,1828   Kenya 6Scarabaeinae Afr.58 (6)
South Africa 8Scarabaeinae Afr.58 (8)
Drepanopodus costatus Wiedeman,1823   South Africa 2Scarabaeinae (2) Afr.26
Dynastes hercules (Linnaeus,1758)   Brazil 1Dynastinae (1) S.Am.22
Colombia 1Dynastinae (1) S.Am.23
Costa Rica 1Dynastinae (1) S.Am.24
Unknown 4Dynastinae (4) S.Am.22+23
Dynastes hyllus (Chevrolat,1843)   Unknown 2Dynastinae (2) S.Am.24
Dynastes neptunus (Quensel,1905)   Colombia 1Dynastinae (1) S.Am.25
Venezuela 1Dynastinae (1) S.Am.25
Dynastes papauana??    Indonesia 1New Guinea
Dynastinae Or.10 (1)
Dynastes tityus (Linnaeus,1763)   United States 1Alabama
Dynastinae N.Am.2 (1)
Unknown 2Dynastinae N.Am.2 (2)
Dyscinetus dubius (Olivier,1789)   Brazil 2Dynastinae S.Am.20 (2)
Colombia 10Dynastinae S.Am.20 (10)
Honduras 1Dynastinae S.Am.20 (1)
Unknown 7Dynastinae S.Am.20 (7)
Dyscinetus gagates (Burmeister,1847)   Argentina 1Dynastinae (1) S.Am.30
Dyscinetus hydrophiloides Arrow,1937   Brazil 2Dynastinae S.Am.20 (2)
Dyscinetus laevipunctatus Bates,1888   Unknown 1Dynastinae S.Am.19 (1)
Dyscinetus morator (Fabricius,1798)   United States 2District of Columbia
Dynastinae N.Am.2 (2)
United States 1Kansas
Dynastinae N.Am.2 (1)
United States 1Pennsylvania
Dynastinae N.Am.2 (1)
Unknown 4Dynastinae N.Am.2 (4)
Dyscinetus olivaceus Höhne,1923   Bolivia 1Dynastinae (1) S.Am.30
Unknown 5Dynastinae S.Am.20 (5)
Dyscinetus picipes (Burmeister,1847)   United States 1Idaho
Dynastinae N.Am.2 (1)
Unknown 1Dynastinae S.Am.19 (1)
Dyscinetus rugifrons (Burmeister,1847)   Argentina 4Dynastinae (4) S.Am.19+30
Unknown 4Dynastinae S.Am.19 (4)
Ectinohoplia balthasari Tesar,1963   China 1Type Coleoptera (1) Paratype 05842:2 of Ectinohoplia balthasari Tesar 1963
  Ectinohoplia balthasari Tesar 1963

Ectinohoplia formosana arrowi Miwa,1931   Taiwan 1Hopliini Pal.3 (1)
Ectinohoplia nigrotincta Fairmaire,1897   China 1Hopliini Pal.3 (1)
Ectinohoplia obducta (Motschulsky,1858)   Japan 5Hopliini Pal.2 (5)
Ectinohoplia yoi Sawada,1939   Taiwan 2Hopliini Pal.3 (2)
Elaphinis cinereonebulosa (DeGeer,1778)   South Africa 2Cetoniinae (2) Afr.14
Unknown 4Cetoniinae (4) Afr.25
Emadiellus equestris Endrödi,1964   Congo 1Aphodiinae (1) Afr.25 (incl.1 paratype)

Enaria melanictera (Klug,1833)   Madagascar 1Melolonthinae Afr.4 (1)
Encya strigiscutata Fairmaire,1883   Madagascar 15Melolonthinae Afr.2 (13)
Afr.4 (2)
Enema pan (Fabricius,1775)   Brazil 24Dynastinae (24) S.Am.6+7
Ecuador 7Dynastinae (7) S.Am.30+34
Peru 5Dynastinae (5) S.Am.14+30+31
Unknown 1Dynastinae S.Am.7 (1)
Eodrepanus bechynei (Janssens,1953)   Gambia 3Scarabaeinae Afr.58 (3)
Kenya 5Scarabaeinae Afr.58 (5)
Eodrepanus fastiditus (Peringuey,1901)   Zimbabwe 1Scarabaeinae Afr.58 (1)
Eodrepanus parallelus (Raffray,1877)   Kenya 2Scarabaeinae Afr.58 (2)
Eophileurus javanus Prell,1913   Indonesia 1Java
Dynastinae Or.10 (1)
Epadoretus reitteri (Semenov,1890)   Russian Federation 2Rutelinae Pal.9 (2)
Epectinaspis mexicana (Burmeister,1844)   Mexico 2Rutelinae N.Am.1 (2)
Epilissus spp.    Unknown 2Scarabaeinae Afr.47 (2)
Epirinus asper Peringuey,1901   South Africa 1Scarabaeinae Afr.19 (1)
Epirinus flagellatus (Fabricius,1775)   Lesotho 2Scarabaeinae Afr.32 (2)
South Africa 5Scarabaeinae Afr.32 (3)
Afr.46 (2)
Epirinus gratus Peringuey,1901   South Africa 1Scarabaeinae Afr.32 (1)
Epirinus hilaris Peringuey,1901   South Africa 3Scarabaeinae Afr.32 (3)
Epirinus obtusus Boheman,1857   South Africa 2Scarabaeinae Afr.32 (2)
Epirinus validus Peringuey,1901   South Africa 2Scarabaeinae Afr.19 (2)
Epixanthis novempunctata (Gory & Percheron,1835)   Madagascar 1Cetoniinae (1) Afr.18 (as E. novemmaculata?)

Eremazus unistriatus Mulsant,1851   Israel 1Aphodiinae Pal.28 (1)
Mauritania 1Aphodiinae Pal.30 (1) as Aphodius
Oman 92Aphodiinae Pal.17 (92)
Spain 17Canary Islands
Aphodiinae Pal.3 (17)
Turkey 1Aphodiinae Pal.11 (1)
Eriesthis decora Peringuey,1902   South Africa 6Hopliini Afr.5 (6)
Eriesthis fallax Burmeister,1844   South Africa 11Hopliini (11) Afr.5+19
Eriesthis guttata Burmeister,1844   South Africa 20Hopliini Afr.5 (20)
Eriesthis hypocrita Peringuey,1902   South Africa 3Hopliini Afr.5 (3)
Eriesthis semihirta Burmeister,1844   South Africa 5Hopliini (5) Afr.5+20
Eriesthis stigmatica Billberg,1817   South Africa 2Hopliini Afr.5 (2)
Eriesthis vestita Burmeister,1844   South Africa 3Hopliini Afr.5 (3)
Unknown 2Hopliini Afr.20 (2)
Eriesthis vulpina Burmeister,1844   South Africa 2Hopliini Afr.5 (2)
Erioscelis emarginata Mannerheim,1829   Brazil 3Dynastinae S.Am.20 (3)
Erioscelis proba Sharp,1877   Colombia 1Dynastinae S.Am.20 (1)
Peru 4Dynastinae (4) S.Am.20+32
Esymus pusillus (Herbst,1789)   Denmark 4Aphodiinae Main (4)
Japan 1Aphodiinae Pal.28 (1)
Sweden 614  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up Vs Vr Dr Gä Hs Me Ån
Alan Dufberg 47 (37)
Aphodiinae Main (577)
Etischiza infantilis (Peringuey,1904)   Tanzania 1Melolonthinae Afr.2 (1)
Euchirus longimanus Linnaeus,1758   Indonesia 3Euchirinae Or.1 (3)
Indonesia 1Java
Euchirinae Or.1 (1)
Unknown 2Euchirinae Or.1 (2)
Euchroea auropigmenta (Gory & Percheron,1835)   Madagascar 1Cetoniinae (1) Afr.18
Euchroea aurostellata Fairmaire,1898   Madagascar 1Cetoniinae (1) Afr.18
Euchroea clementi riphaeus Fairmaire,1897   Madagascar 1Cetoniinae (1) Afr.18
Eucranium arachnoides Brulle,1834   Argentina 3Scarabaeinae S.Am.2 (3)
Eucranium spp.    Argentina 5Scarabaeinae S.Am.2 (5)
Eudicella daphnis (Buquet,1835)   Gambia 28Cetoniinae (28) Afr.1+2 (incl. Neotype)
Kenya 1Cetoniinae Afr.2 (1)
Eudicella darwiniana Kraatz,1880   Togo 2Cetoniinae Afr.20 (2)
Eudicella gralli (Buquet,1836)   Uganda 2Cetoniinae (2) Afr.20
Unknown 2Cetoniinae (2) Afr.19
Eudicella hereroensis Kraatz,1900   Namibia 1Cetoniinae Afr.7 (1)
Eudicella morgani White,1839   Cameroon 1Cetoniinae Afr.20 (1)
Unknown 1Eastafrica
Cetoniinae (1) Afr.22
Eudicella pauperata Kolbe,1884   Congo, the democratic republic of the 2Cetoniinae Afr.20 (2)
Eudicella schultzeorum Kolbe,1906   Cameroon 1Cetoniinae (1) Afr.19
Unknown 3Cetoniinae (3) Afr.19
Eudicella smithi (MacLeay,1838)   Mozambique 1Cetoniinae Afr.1 (1)
Tanzania 2Cetoniinae Afr.19 (2)
Eudicella tetraspilota Harold,1879   Tanzania 2Cetoniinae Afr.19 (2)
Eudicella tetraspilota euthalia Harold,1879   Burundi 2Cetoniinae Afr.20 (2)
Uganda 2Cetoniinae Afr.20 (2)
Euglypta francolina (Burmeister,1842)   Philippines 1Cetoniinae Or.4 (1) as Astraea f.
Euheptaulacus carinatus (Germar,1824)   Slovakia 1Aphodiinae (1) Pal.16
Unknown 3Aphodiinae (3) Pal.15
Euheptaulacus carinatus brutius (Luigione,1920)   Italy 6Aphodiinae Pal.11 (6)
Euheptaulacus carinatus esuriens (Dellacasa,1983   Italy 3Aphodiinae (3) Pal.16
Euheptaulacus sus (Herbst,1783)   Bulgaria 9Aphodiinae Pal.10 (9)
Estonia 15Aphodiinae (15) Pal.15
Sweden 142  Skåne Småland Öland Gotland Uppland
Alan Dufberg 48 (34) as Heptaulacus s.
Aphodiinae Main (108)
Turkey 1Aphodiinae (1) Pal.16
Euheptaulacus villosus (Gyllenhal,1806)   Denmark 2Aphodiinae (2)
Sweden 43  Skåne Halland Öland Gotland Västergötland
Alan Dufberg 48 (3)
Aphodiinae Main (40)
Eumaladera opaciventris (Moser)   Korea, Republic of 1Sericini Pal.1 (1)
Euoniticellus africanus (Harold,1873)   Lesotho 4Scarabaeinae Afr.31 (4)
South Africa 18Scarabaeinae (18) Afr.1+30+31
Euoniticellus fulvus (Goeze,1777)   Afghanistan 16Scarabaeinae Pal.8 (16)
Armenia 6Scarabaeinae Pal.45 (6)
Bulgaria 7Scarabaeinae (7) Pal.9+13+30
Greece 23Scarabaeinae (23) Pal.9+13
Italy 29Scarabaeinae (29) Pal.9+13
Romania 2Scarabaeinae (2) Pal.13
Spain 14Scarabaeinae (14) Pal.9+13
Tunisia 8Scarabaeinae (8) Pal.26
Turkey 5Scarabaeinae Pal.13 (1)
Pal.47 (4)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 3Scarabaeinae Pal.13 (3)
Euoniticellus inaequalis (Reiche,1850)   Ethiopia 5Scarabaeinae Afr.3 (5)
Russian Federation 1Scarabaeinae Pal.9 (1)
Euoniticellus intermedius (Reiche,1850)   Ethiopia 8Scarabaeinae (8) Afr.18
Gambia 47Scarabaeinae Afr.2 (47)
Kenya 6Scarabaeinae Afr.2 (6)
Namibia 2Scarabaeinae (2) Afr.31
Senegal 1Scarabaeinae Afr.2 (1)
South Africa 17Scarabaeinae (17) Afr.31
Euoniticellus pallens (Olivier,1789)   Algeria 4Scarabaeinae (4) Pal.13
Cyprus 1Scarabaeinae (1) Pal.13
Egypt 5Scarabaeinae Afr.1 (5)
Greece 16Scarabaeinae (16) Pal.13
Italy 3Scarabaeinae (3) Pal.13
Morocco 1Scarabaeinae Afr.1 (1)
Somalia 1Scarabaeinae Afr.1 (1)
South Africa 1Scarabaeinae Afr.1 (1)
Spain 8Scarabaeinae (8) Pal.9+13
Tunisia 10Scarabaeinae (9) Pal.9+13
Pal.51 (1)
Turkey 1Scarabaeinae (1) Pal.13
Euoniticellus pallipes (Fabricius,1781)   Afghanistan 8Scarabaeinae Pal.8 (8)
Cyprus 3Scarabaeinae Pal.51 (3)
Greece 11Scarabaeinae Pal.9 (11)
Spain 7Scarabaeinae Pal.9 (7)
Turkey 7Scarabaeinae Pal.9 (7)
Euoniticellus parvus (Kraatz,1895)   Gambia 22Scarabaeinae Afr.1 (22)
Euoniticellus triangulatus (Harold,1873)   Kenya 5Scarabaeinae Afr.1 (5)
South Africa 4Scarabaeinae (4) Afr.1+31
Unknown 2Scarabaeinae (2) Afr.31
Euonthophagus amyntas (Olivier,1789)   Armenia 5Scarabaeinae Pal.45 (5) as Onthophagus a.
Greece 2Crete
Scarabaeinae Pal.49 (2)
Greece 10Scarabaeinae Pal.26 (5)
Pal.52 (5)
Turkey 4Scarabaeinae Pal.49 (3)
Pal.52 (1)
Unknown 1Scarabaeinae Pal.35 (1)
Euonthophagus amyntas alces (Fabricius,1792)   Afghanistan 16Scarabaeinae Pal.5 (16) as Onthophagus amyntas ssp. subviolaceus
Greece 13Scarabaeinae (13) Pal.12+34
Euonthophagus atramentarius (Menetries,1832)   Afghanistan 14Scarabaeinae Pal.6 (14)
Cyprus 12Scarabaeinae Pal.12 (8)
Pal.51 (2)
Pal.53 (2)
Greece 31Scarabaeinae Pal.12 (31)
Turkey 8Scarabaeinae Pal.47 (8)
Euonthophagus bedeli (Reitter,1891)   Afghanistan 14Scarabaeinae Pal.5 (14)
Tunisia 5Scarabaeinae Pal.30 (2)
Pal.52 (3)
Euonthophagus crocatus (Mulsant & Godart,1873)   Morocco 2Scarabaeinae Pal.12 (2)
Tunisia 21Scarabaeinae Pal.25 (18)
Pal.51 (2)
Pal.52 (1)
Euonthophagus gibbosus (Scriba,1790)   Armenia 7Scarabaeinae Pal.45 (7) as Onthophagus g.
Bulgaria 1Scarabaeinae (1) Pal.12
Tajikistan 1Scarabaeinae (1) Pal.35 (as Onthophagus g.)

Turkey 2Scarabaeinae Pal.45 (2) as Onthophagus g.
Euonthophagus koshantschikoffi Reitter,1891   Afghanistan 44Scarabaeinae (44) Pal.6 (as Onthophagus k.)


Afghanistan 7Scarabaeinae (7) Pal.6 (as Onthophagus k. ssp. semirufus))

Euonthophagus spp.    Tunisia 6Scarabaeinae (6) Pal.12
Euonthophagus tissoni (Reitter,1906)   Israel 1Scarabaeinae Pal.52 (1) as Onthophagus hannas Sahlberg
1908
Euorodalus coenosus (Panzer,1798)   Denmark 2Aphodiinae Main (2) as Aphodius
Germany 3Aphodiinae Pal.21 (3) as Aphodius
Netherlands 1Aphodiinae Pal.16 (1) as Aphodius
Sweden 39  Skåne Småland Öland
Alan Dufberg 47 (20) as Aphodius
Aphodiinae Main (19) as Aphodius
Euorodalus paracoenosus Balthasar & Hrubant,1960   Armenia 2Aphodiinae Pal.25 (2) as Aphodius
Cyprus 1Aphodiinae Pal.30 (1)
Greece 3Aphodiinae Pal.6 (3) as Aphodius
Turkey 1Aphodiinae Pal.27 (1) as Aphodius
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Aphodiinae Pal.6 (1) as Aphodius
Euorodalus tersus Erichson,1848   Morocco 7Aphodiinae Pal.6 (7) as Aphodius
Spain 1Aphodiinae Pal.6 (1) as Aphodius
Unknown 1Aphodiinae Pal.21 (1) as Aphodius
Euparia castanea Lepeletier & Serville,1828   Mexico 1Aphodiinae N.Am.2 (1)
Eupatorus gracilicornis Arrow,1908   India 1Dynastinae Or.7 (1)
Malaysia 1Dynastinae Or.11 (1)
Thailand 2Dynastinae Or.13 (2)
Euphoria abronea Janson,1878   Brazil 1Cetoniinae S.Am.2 (1)
Euphoria basalis Gory & Percheron,1833   Mexico 18Cetoniinae N.Am.2 (18)
Unknown 2Cetoniinae N.Am.2 (2)
Euphoria biguttata Gory & Percheron,1833   Unknown 2Cetoniinae N.Am.1 (2)
Euphoria dimidiata (Gory & Percheron,1833)   Mexico 5Cetoniinae N.Am.1 (5)
Unknown 2Cetoniinae N.Am.1 (2)
Euphoria fulgida Fabricius,1775   Canada 3Cetoniinae N.Am.2 (3)
United States 2Illinois
Cetoniinae N.Am.2 (1)
N.Am.3 (1)
Unknown 4Cetoniinae N.Am.2 (4)
Euphoria geminata Chevrolat,1835   Mexico 1Cetoniinae N.Am.1 (1)
Euphoria herbacea (Olivier,1789)   United States 1North Carolina
Cetoniinae N.Am.1 (1)
Unknown 2Cetoniinae N.Am.2 (2)
Euphoria inda Linnaeus,1758   United States 3Connecticut
Cetoniinae N.Am.2 (3)
United States 3Massachusetts
Cetoniinae N.Am.2 (3)
United States 6America Borealis
Cetoniinae N.Am.2 (6)
United States 1New York
Cetoniinae N.Am.4 (1)
Unknown 15Cetoniinae N.Am.2 (15)
Euphoria kerni Haldeman,1852   Mexico 1Cetoniinae N.Am.1 (1) as E. retusa Casey
Euphoria lineoligera Blanchard,1850   Mexico 1Cetoniinae N.Am.2 (1)
Euphoria lurida Fabricius,1775   Argentina 2Cetoniinae S.Am.2 (2)
Brazil 10Cetoniinae S.Am.2 (10)
Chile 1Cetoniinae S.Am.2 (1)
Unknown 3Cetoniinae S.Am.2 (3)
Euphoria melancholica Gory & Percheron, 1833   Unknown 4Cetoniinae N.Am.2 (4)
Euphoria pulchella Gory & Percheron, 1833   Mexico 1Cetoniinae N.Am.1 (1) as Euphoria childreni G.o.P.
Mexico 1Cetoniinae N.Am.2 (1)
Euphoria sepulcralis (Fabricius,1801)   United States 1Arizona
Cetoniinae N.Am.1 (1)
Unknown 1Cetoniinae N.Am.2 (1)
Euphoria spp.    Argentina 2Cetoniinae S.Am.2 (2)
Brazil 10Cetoniinae S.Am.2 (10)
Denmark 1Cetoniinae Pal.25 (1)
Indonesia 1Java
Cetoniinae Or.6 (1)
Mexico 7Cetoniinae N.Am.3 (7)
United States 1Connecticut
Cetoniinae N.Am.3 (1)
Unknown 1Cetoniinae N.Am.3 (1)
Unknown 4Cetoniinae S.Am.2 (4)
Euphoria submaculosa Gory & Percheron,1833   Mexico 1Cetoniinae N.Am.2 (1)
Unknown 2Cetoniinae N.Am.2 (2)
Eupoecila australasiae (Donovan,1805)   Australia 1Cetoniinae Pac.1 (1)
Eurysternus calligrammus Dalman,1824   Brazil 1Scarabaeinae S.Am.9 (1)
Suriname 2Scarabaeinae S.Am.9 (2)
Eurysternus caribaeus (Herbst,1789)   Brazil 2Scarabaeinae S.Am.5 (2)
Eurysternus cayennensis Laporte,1840   Ecuador 1Scarabaeinae S.Am.2 (1)
Peru 2Scarabaeinae S.Am.2 (2)
Eurysternus hirtellus-group    Ecuador 3Scarabaeinae S.Am.2 (3)
Peru 1Scarabaeinae S.Am.2 (1)
Eurysternus impressicollis Castelnau,1840   Colombia 1Scarabaeinae S.Am.9 (1)
Eurysternus inflexus Germar,1824   Brazil 1Scarabaeinae S.Am.9 (1)
Eurysternus parallelus Dalman,1840   Brazil 2Scarabaeinae S.Am.5 (2)
Eurysternus plebejus Harold,1880   Peru 1Scarabaeinae S.Am.2 (1)
Eurysternus spp.    Unknown 2Scarabaeinae S.Am.9 (2)
Euselates ornatus (Saunders,1852)   Thailand 1Cetoniinae Or.4 (1)
Euselates perraudierii (Fairmaire,1893)   Myanmar 1Cetoniinae Or.4 (1)
Euserica lucipeta Baraud,1965   Spain 1Sericini Pal.2 (1)
Euserica modesta Fairmaire,1881   Turkey 1Melolonthinae Pal.18 (1)
Euserica spp.    Morocco 2Sericini Afr.1 (2)
Eutheola bidentata (Burmeister,1847)   Argentina 1Dynastinae S.Am.15 (1)
Colombia 1Dynastinae S.Am.15 (1)
Mexico 2Dynastinae (2) N.Am.4+
Trinidad and Tobago 1Trinidad
Dynastinae S.Am.15 (1)
Eutheola humilis (Burmeister,1847)   Argentina 2Dynastinae S.Am.15 (2)
Brazil 6Dynastinae S.Am.15 (6)
United States 1Mississippi
Dynastinae (1) N.Am.
Unknown 8Dynastinae S.Am.15 (8)
Eutheola humilis rugiceps (Burmeister,1847)   United States 2Florida
Dynastinae N.Am.6 (2)
United States 4Georgia
Dynastinae N.Am.6 (4)
United States 1Maryland
Dynastinae N.Am.6 (1)
Exomala arenicola (Mulsant & Pellet,1870)   Russian Federation 2Rutelinae Pal.6 (2) as Phyllopertha a.
Exomala campestris (Latreille,1804)   Portugal 3Rutelinae (3) Pal.6 (as Phyllopertha c.)

Exomala conspurcata (Harold,1878)   China 1Rutelinae (1) Pal.6 (as Phyllopertha c.)

Korea, Republic of 1Rutelinae (1) Pal.6 (as Phyllopertha c.)

Exomala hirtella (Brullé,1832)   Turkey 1Rutelinae Pal.3 (1)
Exomala orientalis (Waterhouse,1875)   Japan 2Rutelinae (2) Pal.10 (as Blitopertha o.)

Exomala pallidipennis (Reitter,1903)   Japan 1Rutelinae (1) Pal.6 (as Phyllopertha p.)

Exopholis hypoleuca (Wiedemann,1819)   Indonesia 7Java
Melolonthinae Or.8 (6)
Or.10 (1)
Borneo, Indonesia / Malaysia 2Melolonthinae Or.10 (2)
Firminus ciliatus ciliatus (Reiche,1862)   Italy 3Melolonthinae Pal.12 (3)
Firminus ciliatus vexilis (Reitter,1902)   Italy 1Melolonthinae Pal.12 (1)
Frankenbergerius armatus (Boheman,1857)   South Africa 2Scarabaeinae Afr.47 (2)
Fruhstorferia javana Kolbe,1894   Indonesia 1Java
Rutelinae Or.8 (1)
Gametis andrewesi Janson,1901   Unknown 1Cetoniinae Or.5 (1)
Gametis forticula (Janson,1881)   Japan 1Cetoniinae Pal.20 (1) Pal.20
Gametis jucunda (Faldermann,1835)   China 1Cetoniinae Pal.24 (1)
China 2Manchuria
Cetoniinae Pal.24 (2)
India 1Cetoniinae Pal.24 (1)
Japan 2Cetoniinae Pal.20 (1)
Pal.25 (1)
Russian Federation 1Amur
Cetoniinae Pal.24 (1)
Unknown 17Cetoniinae Pal.24 (17)
Gametis jucunda kuperi (Faldermann,1835)   China 5Cetoniinae Pal.25 (5)
Unknown 11Cetoniinae Pal.25 (11)
Gametis spp.    India 1Cetoniinae Or.6 (1)
Indonesia 2New Guinea
Cetoniinae Or.6 (2)
Japan 1Cetoniinae Or.6 (1)
Unknown 2Cetoniinae Or.6 (2)
Gametis versicolor Fabricius,1775   India 3Cetoniinae Or.6 (3)
Unknown 8Cetoniinae Or.6 (8)
Gametis versicolor variegata Fabricius,1775   Réunion 1Cetoniinae Or.6 (1)
Unknown 4Cetoniinae Or.6 (4)
Garreta azureus (Fabricius,1801)   Guinea 2Scarabaeinae Afr.53 (2)
South Africa 1Scarabaeinae Afr.19 (1)
Tanzania 4Scarabaeinae Afr.53 (4)
Zimbabwe 1Scarabaeinae Afr.30 (1)
Garreta fastiditus (Harold,1867)   Lesotho 1Scarabaeinae (1) Afr.30
Garreta morosus (Fairmaire4,1886)   China 1Scarabaeinae (1) Pal.3 (as Gymnopleurus m. )
Garreta nitens (Olivier,1789)   Guinea-Bissau 3Scarabaeinae Afr.6 (3)
Namibia 2Scarabaeinae (2) Afr.19
Namibia 1Scarabaeinae (1) Afr.30
Rwanda 1Scarabaeinae (1) Afr.19
South Africa 4Scarabaeinae (4) Afr.9+30
Unknown 1Scarabaeinae (1) Afr.30
Garreta unicolor (Fahraeus,1857)   South Africa 2Scarabaeinae (2) Afr.30
Swaziland 3Scarabaeinae Afr.6 (3)
Gastroserica brevicornis Lewis,1895   Japan 2Sericini Pal.2 (2)
Gastroserica herzi Heyden,1887   China 1Sericini Pal.1 (1)
Gastroserica marginalis (Brenske,1894)   China 1Sericini Pal.1 (1)
Geniates barbatus Kirby,1818   Brazil 2Rutelinae (2) S.Am 7
Unknown 1Rutelinae (1) S.Am 7
Genuchus hottentottus (Fabricius,1775)   South Africa 1Cetoniinae Afr.29 (1)
Unknown 4Cetoniinae Afr.29 (4)
Genyodonta flavomaculata (Fabricius,1798)   South Africa 4Cetoniinae Afr.8 (4)
Geotrogus euphytus (Buquet,1840)   Algeria 1Melolonthinae (1) Pal.1 (as Pseudapterogynas e.)

Italy 1Melolonthinae (1) Pal.1 (as Pseudapterogynas e.)

Geotrogus gabalus (Buquet,1840)   Algeria 1Melolonthinae Pal.3 (1)
Geotrogus gonoderus (Fairmaire,1860)   Algeria 1Melolonthinae Pal.3 (1)
Geotrogus inflatus (Buquet,1840)   Algeria 1Melolonthinae Pal.3 (1)
Geotrogus numidicus (Lucas,1849)   Algeria 2Melolonthinae (2) Pal.1 (as Pseudoapterygona n.)

Geotrogus psilopus (Fairmaire,1883)   Morocco 1Melolonthinae Pal.3 (1)
Geotrogus sicelis (Blanchard,1851)   Italy 1Melolonthinae Pal.12 (1)
Geotrogus tusculus (Buquet,1840)   Algeria 1Melolonthinae (1) Pal.1 (as Pseudoapterygona n.)

Gilletianus cf. reichei (Harold,1859)   India 1Aphodiinae Or.7 (1)
Indonesia 2Aphodiinae Or.7 (2)
Gilletianus costatulus (A. Schmidt,1908)   India 1Aphodiinae Or.7 (1)
Glycyphana brunnipes Kirby,1818   Unknown 3Cetoniinae Or.5 (3)
Glycyphana fulvistemma Motschulsky,1858   China 2Cetoniinae Pal.24 (2)
Japan 3Cetoniinae (3) Pal.12
Russian Federation 1Sibiria
Cetoniinae Pal.24 (1)
Glycyphana gracilis Sawada,1942   Japan 2Cetoniinae Pal.11 (2)
Glycyphana horsfieldi Hope,1831   Unknown 2Cetoniinae Or.5 (2)
Glycyphana macquarti sumatrensis van Vollenhoven,1864   Indonesia 1Sumatra
Cetoniinae Or.4 (1)
Glycyphana modesta Fabricius,1792   Unknown 1Cetoniinae Or.5 (1)
Glycyphana quadricolor (Wiedemann,1823)   Indonesia 1Sulawesi
Cetoniinae Or.4 (1)
Glycyphana spp.    Unknown 1Cetoniinae Or.5 (1)
Glycyphana stolata (Fabricius,1781)   Australia 1Queensland
Cetoniinae (1) Pac.1
Glyphoderus spp.    Brazil 2Scarabaeinae S.Am.2 (2)
Gnathocera hirta Burmeister,1842   Lesotho 2Cetoniinae Afr.9 (2)
South Africa 8Cetoniinae (8) Afr.6+9
Gnathocera valida Jansen,1884   Gambia 4Cetoniinae Afr.6 (4)
Gnorimella maculosa (Knoch,1801)   United States 1Virginia
Cetoniinae N.Am.1 (1)
Gnorimus nobilis (Linnaeus,1758)   Austria 6Cetoniinae Pal.21 (6)
France 1Cetoniinae Pal.21 (1)
Germany 1Cetoniinae Pal.21 (1)
Greece 15Cetoniinae Pal.21 (14)
Pal.35 (1)
Latvia 1Cetoniinae Pal.26 (1)
Macedonia, The Former Yugoslav Republic of 1Cetoniinae (1) Pal.
Norway 4Cetoniinae (4)
Romania 4Cetoniinae Pal.25 (2)
Pal.35 (2)
Sweden 205  Sk Bl Ha Sm Öl Go GS Ög Vg Bo Ds Sö Up
Alan Dufberg 49 (9)
Cetoniinae Main (196)
Unknown 4Cetoniinae Pal.25 (4)
Gnorimus subcostatus Ménétriés,1832   Iran 1Cetoniinae Pal.31 (1)
Gnorimus subopacus Motschulsky,1860   Japan 2Cetoniinae Pal.21 (2)
Gnorimus variabilis (Linnaeus,1758)   Bulgaria 1Cetoniinae Pal.21 (1)
Germany 3Cetoniinae (3) Pal.21+25
Sweden 99  Skåne Blekinge Småland Öland Östergötland Västergötland Uppland
Alan Dufberg 49 (3)
Cetoniinae Main (96)
Unknown 2Cetoniinae Pal.25 (2)
Goliathus albosignatus Boheman,1857   South Africa 1Cetoniinae Afr.4 (1)
Goliathus goliatus (Linnaeus,1771)   Congo, DRC 1Cetoniinae Afr.33 (1)
Unknown 1Cetoniinae Afr.30 (1) without lable
Goliathus regius Klug,1835   Côte D'ivoire 1Cetoniinae Afr.33 (1)
Golofa claviger (Linnaeus,1771)   Unknown 2Dynastinae (2) S.Am.29
Golofa eacus Burmeister,1847   Colombia 1Dynastinae (1) S.Am.29
Ecuador 15Dynastinae (15) S.Am.29+35+36
Grenada 2Dynastinae (2) S.Am.29
Peru 1Dynastinae (1) S.Am.31
Unknown 2Dynastinae (2) S.Am.29
Golofa imbellis Bates,1888   Colombia 1Dynastinae (1) S.Am.24
Golofa imperialis Thomson,1858   Ecuador 1Dynastinae (1) S.Am.30
Mexico 1Dynastinae N.Am.4 (1)
Golofa inermis Thomson,1859   Ecuador 2Dynastinae (2) S.Am.24
Golofa pelagon Burmeister,1847   Bolivia 1Dynastinae (1) S.Am.35
Ecuador 1Dynastinae (1) S.Am.30
Golofa pizarro Hope,1837   Mexico 2Dynastinae N.Am.2 (2)
Golofa porteri Hope,1837   Colombia 1Dynastinae (1) S.Am.29
Unknown 2Dynastinae (2) S.Am.30
Venezuela 3Dynastinae (3) S.Am.30+38
Gonaphodiellus arcanus Dellacasa, Dellacasa & Gordon,2012   Honduras 9Aphodiinae S.Am.2 (9)
Gonaphodiellus bimaculosus (Schmidt,1909)   Mexico 1Aphodiinae N.Am.3 (1) as Aphodius
Gonaphodiellus opisthius (Bates,1887)   Mexico 1Aphodiinae N.Am.8 (1)
Gonaphodiellus sexguttatus (Sshmidt,1916)   Mexico 13Aphodiinae N.Am.3 (13) as Aphodius
Gromphas lacordairei Brulle,1843   Bolivia 1Scarabaeinae S.Am.6 (1)
Gymnetosoma holoserica (Voet,1779)   Brazil 1Cetoniinae S.Am.2 (1)
Gymnopleurus aciculatus Gebler,1841   Afghanistan 106Scarabaeinae (106) Pal.3+4+27+30
Kazakhstan 2Scarabaeinae (2) Pal.16
Tajikistan 1Scarabaeinae Pal.16 (1)
Turkmenistan 2Scarabaeinae Pal.41 (2)
Uzbekistan 2Scarabaeinae (2) Pal.16
Gymnopleurus aenescens Wiedeman,1821   South Africa 1Scarabaeinae (1) Afr.29
Gymnopleurus assamensis Waterhouse,1890   India 10Scarabaeinae Or.4 (1) as Paragymnopleurus sinuatus assamensis Waterhouse
1890
Or.9 (9)
Gymnopleurus bicolor Latreille,1823   Unknown 22 ()
Gymnopleurus caffer Fahraeus,1857   South Africa 1Scarabaeinae Afr.53 (1)
Gymnopleurus coerulescens (Olivier,1789)   Ethiopia 1Scarabaeinae Afr.45 (1)
Senegal 1Scarabaeinae Afr.53 (1)
Sierra Leone 1Scarabaeinae Afr.53 (1)
Gymnopleurus flagellatus (Fabricius,1787)   Afghanistan 110Scarabaeinae (63) Pal.28+29
(47) Pal.2+3 as G. coriarius (Herbst
1789)
Algeria 2Scarabaeinae Pal.17 (1)
Pal.29 (1)
Armenia 3Scarabaeinae Pal.46 (3)
France 1Scarabaeinae Pal.24 (1) as G. coriarius (Herbst
1789)
Morocco 2Scarabaeinae Pal.43 (2)
Russian Federation 1Dagestan
Scarabaeinae Pal.43 (1) as var. serratus
Spain 3Scarabaeinae Pal.43 (3)
Tunisia 9Scarabaeinae Pal.24 (1) as G. coriarius (Herbst
1789)
Pal.29 (1)
Pal.43 (4)
Pal.50 (3)
Turkey 1Scarabaeinae Pal.17 (1)
Turkey 3Scarabaeinae Pal.43 (3)
Unknown 4Scarabaeinae Pal.43 (3)
Pal.43 (1) as var. asperatus
Caucasus, Ryssland / Georgien / Azerbajdzjan / Armenien / Iran / Turkiet 1Scarabaeinae Pal.43 (1) as var. serratus
Gymnopleurus fulgidus (Olivier,1789)   Guinea-Bissau 7Scarabaeinae Afr.6 (7)
Senegal 4Scarabaeinae Afr.53 (4)
Sudan 3Scarabaeinae Afr.18 (3)
Unknown 3Scarabaeinae Afr.53 (3)
Gymnopleurus gemmatus Harold,1871   India 1Scarabaeinae Or.10 (1)
Gymnopleurus geoffroyi (Fuessly,1775)   Afghanistan 33Scarabaeinae (33) Pal.27+28
Bulgaria 3Scarabaeinae Pal.47 (3)
Cyprus 1Scarabaeinae Pal.50 (1)
France 2Scarabaeinae (2) Pal.24
Greece 4Scarabaeinae Pal.15 (2)
Pal.41 (1)
Pal.50 (1)
Israel 3Scarabaeinae Pal.15 (3)
Italy 1Scarabaeinae Pal.41 (1)
Romania 1Scarabaeinae (1) Pal.24
Syria 1Scarabaeinae (1) Pal.24
Turkey 6Scarabaeinae Pal.15 (2)
Pal.41 (1)
Pal.50 (3)
Turkey 23Scarabaeinae Pal.41 (4)
Pal.46 (3)
Pal.47 (16)
Unknown 2Scarabaeinae Pal.41 (1)
Pal.46 (1)
Unknown 1Scarabaeinae Pal.41 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Scarabaeinae (1) Pal.15
Gymnopleurus humanus MacLeay,1821   Botswana 5Scarabaeinae Afr.52 (5)
Namibia 9Scarabaeinae Afr.49 (9)
South Africa 6Scarabaeinae Afr.29 (6)
Unknown 3Scarabaeinae Afr.52 (3)
Gymnopleurus koenigii (Fabricius,1775)   India 6Scarabaeinae Or.10 (6)
Sri Lanka 4Scarabaeinae Or.2 (2)
Or.10 (2)
Tunisia 1Scarabaeinae (1) Pal.17
Unknown 12Scarabaeinae Or.10 (12)
Gymnopleurus leei (Fabricius,1792)   South Africa 16Scarabaeinae (16) Afr.29
Gymnopleurus melanarius Harold,1867   Hong Kong 3Scarabaeinae Pal.43 (3)
Gymnopleurus miliaris (Fabricius,1775)   Afghanistan 1Scarabaeinae Pal.3 (1)
China 1Tibet
Scarabaeinae Or.10 (1)
India 2Scarabaeinae Or.10 (2)
Unknown 1Scarabaeinae Or.10 (1)
Gymnopleurus mopsus (Pallas,1781)   Afghanistan 22Scarabaeinae Pal.4 (22)
Bulgaria 7Scarabaeinae (7) Pal.16+17
France 1Scarabaeinae Pal.42 (1)
Greece 1Rhodos
Scarabaeinae Pal.42 (1)
Greece 39Scarabaeinae Pal.16 (37)
Pal.42 (2)
Hungary 1Scarabaeinae Pal.42 (1)
Italy 8Scarabaeinae Pal.16 (3)
Pal.42 (5)
Morocco 1Scarabaeinae Pal.42 (1)
Russian Federation 5Buryatiya
Scarabaeinae Pal.42 (5)
Spain 1Scarabaeinae Pal.42 (1)
Syria 4Scarabaeinae Pal.24 (4)
Tunisia 3Scarabaeinae Pal.50 (3)
Turkey 7Scarabaeinae Pal.42 (7)
Turkey 22Scarabaeinae Pal.16 (3)
Pal.42 (16)
Pal.47 (3)
Ukraine 2Scarabaeinae Pal.42 (2)
Unknown 11Scarabaeinae Pal.35 (1)
Pal.43 (10)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Scarabaeinae Pal.16 (1)
Gymnopleurus sericatus Erichson,1843   Namibia 7Scarabaeinae (7) Afr.29
Gymnopleurus sericeifrons Fairmaire,1887   South Africa 2Scarabaeinae Afr.29 (2)
Sudan 4Scarabaeinae Afr.19 (4)
Tanzania 1Scarabaeinae Afr.19 (1)
Tanzania 3Scarabaeinae Afr.50 (3)
Gymnopleurus sinuatus Olivier,1789   China 8Scarabaeinae Or.9 (8)
Hong Kong 2Scarabaeinae Or.9 (2)
India 3Scarabaeinae Or.9 (3)
Indonesia 1Celebes
Scarabaeinae Or.9 (1)
Unknown 4Scarabaeinae Or.9 (4)
Ostindien 5Scarabaeinae Or.9 (5)
Gymnopleurus smaragdinus Fahraeus,1857   South Africa 1Scarabaeinae Afr.52 (1)
Gymnopleurus sparsus ?   Malaysia 1Scarabaeinae Or.3 (1)
Gymnopleurus spp.    Guinea-Bissau 1Scarabaeinae Afr.6 (1)
Kazakhstan 2Scarabaeinae Pal.41 (2)
South Africa 1Scarabaeinae Afr.53 (1)
Tanzania 2Scarabaeinae Afr.53 (2)
Unknown 9Scarabaeinae Afr.53 (9)
Unknown 1Scarabaeinae Pal.41 (1)
Gymnopleurus sturmii Mac Leay,1821   Algeria 2Scarabaeinae Pal.16 (2)
France 2Scarabaeinae Pal.24 (2)
Italy 8Scarabaeinae (8) Pal.16+30
Morocco 3Scarabaeinae Pal.16 (1)
Pal.42 (2)
Spain 6Scarabaeinae Pal.16 (2)
Pal.42 (4)
Spain 4Mallorca
Scarabaeinae Pal.38 (3)
Pal.42 (1)
Tunisia 31Scarabaeinae (31) Pal.25+26+35
Turkey 2Scarabaeinae Pal.42 (2)
Unknown 24Scarabaeinae Pal.34 (1)
Pal.42 (23)
Gymnopleurus thoracicus Harold,1868   Unknown 4Scarabaeinae Afr.50 (4)
Gymnopleurus tuxeni Petrovitz,1955   Afghanistan 28Scarabaeinae (28) Pal.3+29
Gymnopleurus virens Erichson,1843   Botswana 3Scarabaeinae Afr.52 (3)
Namibia 19Scarabaeinae (19) Afr.19+29
Unknown 1Scarabaeinae Afr.52 (1)
Gymnopyge hopliaeformis Linell,1896   United States 1Arizona
Melolonthinae N.Am.1 (1)
Gymnoschiza serrata (Aulmann)   Cameroon 1Melolonthinae Afr.2 (1)
Gyronotus fimetarius Kolbe,1894   Kenya 4Scarabaeinae (4) Afr.19
Gyronotus pumilus (Boheman,1857)   South Africa 3Scarabaeinae (3) Afr.32
Haploscapanes barbarossa (Fabricius,1775)   Australia 2Dynastinae Pac.1 (2)
Harmogaster exarata Harold,1861   South Africa 1Aphodiinae Afr.32 (1)
Harmogaster opacula Harold,1869   South Africa 3Aphodiinae (3) Afr.24
Harmogaster spp.    South Africa 2Aphodiinae (2) Afr.16
Harmogaster sulcatula Schmidt,1911   South Africa 1Aphodiinae (1) Afr.16
Haroldiellus sallei (Harold,1863)   Costa Rica 2Aphodiinae S.Am.2 (2)
Hatamus gressitti Endrödi,1971   Indonesia 1Dynastinae Or.9 (1)
Heliocopris andersoni Bates,1868   Egypt 1Scarabaeinae Afr.54 (1)
Kenya 1Scarabaeinae Afr.54 (1)
Namibia 2Scarabaeinae Afr.10 (1)
Afr.36 (1)
South Africa 1Scarabaeinae Afr.36 (1)
Unknown 1Scarabaeinae Afr.54 (1)
Zimbabwe 1Scarabaeinae Afr.54 (1)
Heliocopris antenor (Olivier,1789)   Gambia 3Scarabaeinae (3) Afr.22
Guinea-Bissau 1Scarabaeinae Afr.7 (1)
Kenya 1Scarabaeinae (1) Afr.22
Namibia 1Scarabaeinae (1) Afr.22
Rwanda 1Scarabaeinae (1) Afr.22
Heliocopris bucephalus (Fabricius,1775)   Indonesia 2Java
Scarabaeinae Or.11 (2)
Thailand 1Scarabaeinae Or.2 (1)
Unknown 6Scarabaeinae Or.11 (6)
Ind. Or. 4Scarabaeinae Or.11 (4)
Heliocopris cf. hamadryas (Fabricius,1775)   Gambia 2Scarabaeinae Afr.44 (2)
Heliocopris cf. japetus Klug,1855   Gambia 1Scarabaeinae Afr.44 (1)
Heliocopris dilloni Guerin,1847   Kenya 4Scarabaeinae (4) Afr.21
Heliocopris faunus Boheman,1857   Namibia 6Scarabaeinae Afr.36 (4)
Afr.54 (2)
Namibia 1Scarabaeinae Afr.54 (1)
Heliocopris felschei Kolbe,1904   Tanzania 1Scarabaeinae Afr.56 (1)
Heliocopris gigas (Linnaeus,1758)   Egypt 1Scarabaeinae Afr.55 (1)
Namibia 3Scarabaeinae Afr.37 (3)
South Africa 2Scarabaeinae Afr.36 (2)
Unknown 5Scarabaeinae Afr.55 (5)
Heliocopris hamadryas (Fabricius,1775)   Cameroon 1Scarabaeinae Afr.56 (1)
Ethiopia 2Scarabaeinae Afr.56 (2)
Namibia 1Scarabaeinae Afr.10 (1)
South Africa 4Scarabaeinae Afr.37 (4)
Tanzania 1Scarabaeinae Afr.56 (1)
Unknown 3Scarabaeinae Afr.56 (3)
Heliocopris haroldi Kolbe,1893   Cameroon 1Scarabaeinae Afr.56 (1)
Tanzania 4Scarabaeinae Afr.20 (4)
Heliocopris hunteri Waterhouse,1891   Kenya 1Scarabaeinae Afr.55 (1)
Namibia 1Scarabaeinae Afr.55 (1)
Tanzania 1Scarabaeinae Afr.55 (1)
Unknown 1Scarabaeinae Afr.55 (1)
Heliocopris japetus Klug,1855   South Africa 1Scarabaeinae Afr.38 (1)
Unknown 2Scarabaeinae Afr.55 (2)
Heliocopris midas Fabricius,1775   India 2Scarabaeinae Or.10 (2)
Heliocopris myrmidion Kolbe,1893   Kenya 1Scarabaeinae (1) Afr.20
Heliocopris neptunus Boheman,1857   Kenya 1Scarabaeinae Afr.55 (1)
Unknown 1Scarabaeinae Afr.55 (1)
Zimbabwe 1Scarabaeinae Afr.55 (1)
Heliocopris neptunus-group    Zambia 1Scarabaeinae Afr.20 (1)
Heliocopris pirmal (Fabricius,1798)   Kenya 1Scarabaeinae (1) Afr.19
Namibia 1Scarabaeinae (1) Afr.38
Heliocopris spp.    China 1Scarabaeinae Pal.44 (1)
India 1Scarabaeinae Or.12 (1)
Kenya 1Scarabaeinae Afr.55 (1)
Tanzania 1Scarabaeinae Afr.45 (1)
Unknown 9Scarabaeinae Or.12 (3)
Pal.44 (6)
Uzbekistan 1Scarabaeinae Pal.41 (1)
Hemichaetoplia pallidipennis (Gyllenhal,1817)   Greece 1Rutelinae (1) Pal.8 (as Anisoplia) p
Hemictenius simplicitarsis Reitter,1897   Tajikistan 1Melolonthinae Pal.12 (1)
Hemicyclium lateritius (Roth,1851)   Ethiopia 3Aphodiinae Afr.24 (1)
Afr.29 (2)
Hemicyclium unicornutus (Schmidt,1909)   Zimbabwe 5Aphodiinae Afr.16 (4)
Afr.35 (1)
Hemiphileurus variolosus (Burmeister,1847)   Colombia 1Dynastinae (1) S.Am.22
Unknown 1Dynastinae (1) S.Am.22
Venezuela 2Dynastinae (2) S.Am.22
Hemiphileurus vicarius Prell,1936   Colombia 1Dynastinae (1) S.Am.22
Heptaulacus alpinus Drapiez,1819   Turkey 1Aphodiinae Pal.6 (1)
Heptaulacus brancoi Baraud,1976   Spain 5Aphodiinae Pal.11 (5)
Heptaulacus testudinarius (Fabricius,1775)   Germany 1Aphodiinae (1) Pal.15
Sweden 2 Skåne
Aphodiinae Main (2)
Exotica 1Aphodiinae Pal.16 (1)
Heptophylla picea Motschulsky,1857   Japan 4Melolonthinae (4) Pal.5 +10
Heterochelus adspersus Peringuey,1902   South Africa 2Hopliini (2) 2 paratypes Afr.10
Heterochelus albosetosus Kulzer,1960   South Africa 1Hopliini (1) Types 2021:2
Heterochelus armipes Boheman   South Africa 2Hopliini Afr.10 (2)
Heterochelus arthriticus Fabricius,1781   South Africa 13Hopliini (13) Afr.10+14
Heterochelus auricollis Burmeister,1844   South Africa 1Hopliini Afr.10 (1)
Heterochelus barkeri Kulzer,1960   South Africa 1Hopliini (1) Types 2022:2
Heterochelus binotatus Thunberg,1818   South Africa 7Hopliini Afr.10 (7)
Heterochelus bivittatus Burmeister,1844   South Africa 1Hopliini Afr.10 (1)
Heterochelus brincki Kulzer,1960   South Africa 35Hopliini (35) Types 2016:1-35
Heterochelus capicola Fabricius,   South Africa 2Hopliini Afr.10 (2)
Heterochelus carus Kulzer,1960   South Africa 3Hopliini (3) Types 2019:1-3
Heterochelus coccineus Burmeister,1844   South Africa 1Hopliini (1) Afr.11
Heterochelus comosus Peringuey,   South Africa 2Hopliini (2) Afr.11
Heterochelus consanguineus Peringuey,   South Africa 2Hopliini (2) Afr.11
Heterochelus cristaticeps Peringuey,   South Africa 2Hopliini (2) Afr.11
Heterochelus denticeps Wiedemann,   South Africa 3Hopliini (3) Afr.11
Heterochelus detritus Burmeister,1844   South Africa 2Hopliini (2) Afr.11
Heterochelus dissidens Peringuey,1902   South Africa 2Hopliini (2) 2 paratypes Afr.11
Heterochelus ditus Peringuey,1902   South Africa 11Hopliini (11) Afr.11+15
Heterochelus diversus Kulzer,1960   South Africa 1Hopliini (1) Types 2023:2
Heterochelus elegans Peringuey,   South Africa 1Hopliini (1) 1 paratype Afr.11
Heterochelus fallasciosa Kulzer,1960   South Africa 5Hopliini (5) Types 2017:1-5
Heterochelus forcipatus Burmeister,1844   South Africa 35Hopliini (35) Afr.8+17
Heterochelus fraternus Peringuey,   South Africa 3Hopliini (3) Afr.11
Heterochelus gonager (Fabricius,1781)   South Africa 22Hopliini (22) Afr.9+18
Heterochelus gracilis Kulzer,1960   South Africa 2Hopliini (2) Types 2030:2-3
Heterochelus guillarmodi Schein,1959   South Africa 17Hopliini Afr.18 (15)
Type Schein (2) Holotype Paratype 02018:1-2 of Heterochelus guillarmodi Schein 1959
  Heterochelus guillarmodi Schein 1959

Heterochelus haykei Kulzer,1960   South Africa 1Hopliini (1) Type 2031:2
Heterochelus hessei Kulzer,1960   South Africa 1Hopliini (1) Type 2024:2
Heterochelus hybridus Burmeister,1844   South Africa 4Hopliini (4) Afr.11
Heterochelus icterinus Peringuey,   South Africa 2Hopliini (2) Afr.11
Heterochelus insignus Peringuey,   South Africa 1Hopliini (1) Afr.11
Heterochelus jucundulus Peringuey,   South Africa 1Hopliini (1) 1 paratype? Afr.11
Heterochelus kochi Schein,   South Africa 2Hopliini (2) Afr.11
Heterochelus leoninus Peringuey,1902   South Africa 6Hopliini Afr.10 (6)
Heterochelus longicollis Kulzer,1960   South Africa 1Hopliini (1) Type 2025:2
Heterochelus mimus Peringuey,   South Africa 1Hopliini (1) 1 paratype? Afr.11
Heterochelus miserabilis Blanch.,   South Africa 1Hopliini (1) Afr.11
Heterochelus molestus Peringuey,   South Africa 1Hopliini (1) Afr.11
Heterochelus multidentatus Burmeister,1844   South Africa 35Hopliini (35) Afr.11+18
Heterochelus murinus Burmeister,1844   South Africa 44Hopliini (44) Afr.11+15
Heterochelus natalensis Peringuey,   South Africa 1Hopliini (1) 1 paratype? Afr.11
Heterochelus nubilis Burmeister,1844   South Africa 5Hopliini Afr.10 (5)
Heterochelus oreopygus Burmeister,1844   South Africa 29Hopliini (29) Afr.11+17
Heterochelus pachyglutus Wiedemann,1823   South Africa 232Hopliini (232) Afr.9+15+16+17
Heterochelus parilis Peringuey,1902   South Africa 7Hopliini Afr.9 (7)
Heterochelus pavidus Peringuey,   South Africa 2Hopliini (2) 2 paratypes Afr.11
Heterochelus persimilis Peringuey,   South Africa 2Hopliini (2) Afr.11
Heterochelus placatus Peringuey,   South Africa 2Hopliini (2) Afr.11
Heterochelus podagricus Fabricius,   South Africa 2Hopliini (2) Afr.11
Heterochelus rudebecki Kulzer,1960   South Africa 12Hopliini (12) Types 2020:1-12
Heterochelus rufimanus Castelnau,1840   South Africa 304Hopliini (304) Afr.9+10+13+14
Heterochelus saldanhensis Kulzer,1960   South Africa 1Hopliini (1) Type 2026:2
Heterochelus sexlineatus Thunberg,1818   South Africa 35Hopliini (35) Afr.8+14
Heterochelus simulator Peringuey,   South Africa 1Hopliini (1) Afr.11
Heterochelus sobrinus Burmeister,1844   South Africa 5Hopliini Afr.9 (5)
Heterochelus spp.    South Africa 1Hopliini Afr.10 (1)
Heterochelus stuckenbergi Kulzer,1960   South Africa 13Hopliini (13) Afr.18
Heterochelus subvittatus Burmeister,1844   South Africa 2Hopliini (2) Afr.11
Heterochelus sulphureus Boheman,   South Africa 1Hopliini (1) Afr.11
Heterochelus tristis Peringuey,   South Africa 1Hopliini (1) Afr.11
Heterochelus unguiculatus Burmeister,1844   South Africa 2Hopliini (2) Afr.11
Heterochelus viridicollis Blanchard,   South Africa 1Hopliini (1) Afr.12
Heterochelus vittipennis Burmeister,1844   South Africa 1Hopliini (1) Afr.12
Heterochelus vulpecula Burmeister,1844   South Africa 1Hopliini Afr.10 (1)
Heteroclita raeuperi (Schaum,1841)   Lesotho 2Cetoniinae Afr.9 (2)
Heterocnemis graeca (Brulle,1832)   Greece 8Cetoniinae Pal.17 (6)
Pal.28 (2)
Heteroconus muticus (Burmeister,1847)   Madagascar 1Dynastinae Afr.5 (1)
Heterogomphus achilles Burmeister,1847   Brazil 3Dynastinae S.Am.9 (3)
Heterogomphus aidoneus (Perty,1830)   Unknown 1Dynastinae S.Am.9 (1)
Heterogomphus bispinosus Burmeister,1847   Brazil 1Dynastinae S.Am.8 (1)
Heterogomphus bourcieri Guerin-Meneville,1851   Ecuador 5Dynastinae (5) S.Am.32
Heterogomphus chevrolati Burmeister,1847   Mexico 1Dynastinae S.Am.8 (1)
Heterogomphus dilaticollis Burmeister,1847   Peru 2Dynastinae (2) S.Am.31
Unknown 3Dynastinae S.Am.10 (3)
Heterogomphus eteocles Burmeister,1847   Brazil 13Dynastinae S.Am.8 (13)
Unknown 2Dynastinae S.Am.8 (2)
Heterogomphus peruanus Endrödi,1976   Unknown 1Dynastinae S.Am.9 (1)
Heterogomphus rugicollis Prell,1912   Unknown 1Dynastinae S.Am.8 (1)
Heterogomphus schoenherri Burmeister,1847   Colombia 1Dynastinae S.Am.9 (1)
Ecuador 10Dynastinae (10) S.Am.31+33
Unknown 3Dynastinae S.Am.9 (3)
Heterogomphus thoas Burmeister,1847   Unknown 1Dynastinae S.Am.10 (1)
Heterogomphus ulysses Burmeister,1847   Brazil 1Dynastinae S.Am.9 (1)
Heteroligus geotrupinus (Quedenfeldt,1884)   Senegal 1Dynastinae Afr.5 (1)
Unknown 1Dynastinae Afr.5 (1)
Heteroligus meles Billberg,1815   Angola 1Dynastinae Afr.8 (1)
Cameroon 1Dynastinae Afr.8 (1)
Congo 1Dynastinae (1) Afr.19
Gambia 1Dynastinae Afr.8 (1)
Ghana 1Dynastinae Afr.8 (1)
Guinea-Bissau 1Dynastinae (1) Afr.18
Senegal 5Dynastinae Afr.8 (5)
Sierra Leone 5Dynastinae (5) Afr.18
Heteroligus peringueyi Arrow,1911   Malawi 3Dynastinae (3) Afr.18
Heteroligus planifrons Prell,1914   Sierra Leone 3Dynastinae (3) Afr.16
Heteroligus spp.    South Africa 1Dynastinae (1) Afr.11
Heteronitis castelnaui (Harold,1862)   South Africa 2Scarabaeinae (2) Afr.38
Heteronychus amplipennis Benderitter,1915   South Africa 1Dynastinae (1) Afr.12
Heteronychus amplus Kolbe,1900   Tanzania 2Dynastinae Afr.7 (2)
Heteronychus arator (Fabricius,1775)   Ethiopia 2Dynastinae Afr.8 (2)
South Africa 54Dynastinae (54) Afr.8+11+12
Unknown 3Dynastinae Afr.8 (3)
Heteronychus arator australis (Fabricius,1775)   Australia 5Dynastinae Pac.2 (5)
Heteronychus atratus obtusifrons Fairmaire,1893   Ethiopia 49Dynastinae (49) Afr.16
Heteronychus bituberculatus Kolbe,1900   Madagascar 5Dynastinae Afr.7 (5)
Heteronychus citernii Paoli,1934   Unknown 1Dynastinae Afr.8 (1)
Heteronychus congoensis Kolbe,1900   Madagascar 2Dynastinae Afr.7 (2)
Heteronychus consimilis Kolbe,1900   Kenya 1Dynastinae Afr.7 (1)
Heteronychus cricetus (Hausmann,1807)   South Africa 2Dynastinae (2) Afr.11
Heteronychus fossor Reiche,1849   Guinea-Bissau 16Dynastinae (16) Afr.19
Tanzania 12Dynastinae Afr.7 (12)
Heteronychus impudens Jack,1923   Ethiopia 2Dynastinae (2) Afr.16
Heteronychus licas (Klug,1835)   Guinea-Bissau 1Dynastinae (1) Afr.18
South Africa 1Dynastinae (1) Afr.11
Heteronychus lioderes Redtenbacher,1868   India 1Dynastinae Or.10 (1)
Indonesia 2Java
Dynastinae Or.10 (2)
Nepal 1Dynastinae Or.10 (1)
Heteronychus mozambicus Peringuey,1908   Guinea-Bissau 3Dynastinae (3) Afr.18
Senegal 27Dynastinae Afr.7 (27)
Sierra Leone 8Dynastinae (8) Afr.19
South Africa 1Dynastinae Afr.7 (1)
Heteronychus parumpunctatus Burmeister,1847   Guinea-Bissau 2Dynastinae (2) Afr.16
Heteronychus parvus Burmeister,1847   Gambia 114Dynastinae (114) Afr.6+19
Senegal 1Dynastinae Afr.6 (1)
Heteronychus pauperatus Peringuey,1901   Ethiopia 2Dynastinae (2) Afr.16
Tanzania 10Dynastinae Afr.8 (10)
Heteronychus rusticus (Klug,1832)   Cameroon 1Dynastinae Afr.8 (1)
Heteronychus tenuestriatus Fairmaire,1893   South Africa 1Dynastinae (1) Afr.11
Tanzania 1Dynastinae Afr.8 (1)
Heteronychus transvalicus Kolbe,1900   Namibia 1Dynastinae (1) Afr.12
South Africa 7Dynastinae (7) Afr.12
Heteronyx agrestis Burmeister,1855   Australia 1Western Australia
Melolonthinae Pac.1 (1)
Heteronyx elongatus Blanchard,1850   Australia 1Western Australia
Melolonthinae Pac.1 (1)
Heteronyx exsectus Blackburne,1909   Australia 1Western Australia
Melolonthinae Pac.1 (1)
Heteronyx spp.    Australia 3Western Australia
Melolonthinae Pac.1 (3)
Australia 1Melolonthinae Pac.1 (1)
Australia 1Melolonthinae Pac.1 (1) As H. nigellus Erichson
Heterorrhina elegans (Fabricius,1781)   Unknown 3Cetoniinae Or.1 (3)
Heterorrhina macleayi (Kirby,1818)   Philippines 2Cetoniinae Or.1 (2)
Heterorrhina obesa Janson,1884   India 4Cetoniinae Or.1 (3)
Or.7 (1)
Heterorrhina punctatissima Westwood,1842   India 4Cetoniinae Or.1 (4)
Heterorrhina sexmaculata (Fabricius,1801)   Unknown 1Cetoniinae Or.1 (1)
Heterorrhina spp.    Indonesia 1Cetoniinae Or.7 (1)
Hexodon unicolor Olivier,1789   Madagascar 2Dynastinae Afr.9 (2)
Hilyotrogus formosanus (Niijima & Kinoshita,1927)   Taiwan 1Melolonthinae Pal.12 (1)
Holocephalus eridanus (Olivier,1789)   Unknown 3Scarabaeinae (3) S.Am.5+6
Holochelus aequinoctialis (Herbst,1790)   Austria 3Melolonthinae (3) Pal.4+9+11
Hungary 6Melolonthinae Pal.4 (6)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 4Melolonthinae Pal.9 (4)
Holochelus cf. aequinoctialis (Herbst,1790)   Turkey 1Melolonthinae Pal.16 (1) as Miltotrogus
Holochelus pilicollis (Gyllenhal,1817)   Hungary 2Melolonthinae Pal.4 (2)
Holochelus rusticus Faldermann,1835   Armenia 1Melolonthinae Pal.16 (1) as Miltotrogus r.
Holochelus tataricus (Faldermann,1835)   Russian Federation 1Melolonthinae (1) Pal.4 (as H. arcilabris Marseul)

Holochelus vernus (Germar,1823)   Romania 1Melolonthinae Pal.9 (1)
Russian Federation 1Melolonthinae Pal.4 (1)
Unknown 1Melolonthinae Pal.11 (1)
Holotrichia bidentata (Burmeister,1855)   Indonesia 1Sumatra
Melolonthinae Or.10 (1)
Holotrichia brunneipennis Moser,1916   India 3Melolonthinae Or.8 (3)
Holotrichia danielssoni Bunalski,1965   Sri Lanka 35Type Bunalski 1 (35) Holotype Paratype 02723:1-36 of Holotrichia danielssoni Bunalski 1995
  Holotrichia danielssoni Bunalski 1995

Holotrichia diomphalia (Bates,1888)   China 1Melolonthinae Pal.3 (1)
Korea, Republic of 1Melolonthinae Pal.3 (1)
Russian Federation 1Melolonthinae Pal.12 (1)
Holotrichia hirtipectus (Walker,1859)   Sri Lanka 1Melolonthinae Or.6 (1)
Holotrichia inducta (Walker,1859)   Sri Lanka 1Melolonthinae Or.6 (1)
Holotrichia kiotonensis Brenske,1894   Japan 1Melolonthinae Pal.3 (1) as H. inelegans Lewis
Holotrichia paralella (Motschulsky,1854)   China 5Melolonthinae Pal.3 (5)
Holotrichia reynaudi (Blanchard,1850)   Sri Lanka 1Melolonthinae Or.6 (1)
Holotrichia rufoflava Brenske,1894   Sri Lanka 2Melolonthinae Or.6 (2)
Holotrichia schmitzi Frey,1973   Sri Lanka 3Melolonthinae Or.6 (3)
Holotrichia serrata (Fabricius,1787)   Sri Lanka 9Melolonthinae Or.6 (9)
Holotrichia seticollis Moser,1913   India 1Melolonthinae Or.10 (1)
Holotrichia sichotana Brenske,1896   Russian Federation 1Melolonthinae Pal.3 (1)
Holotrichia spp.    China 1Melolonthinae Pal.3 (1)
Holotrichia staudingeri Brenske,1892   India 2Melolonthinae Or.8 (2)
Homalothyrea franki (Janson,1888)   Namibia 1Cetoniinae (1) Afr.15
Unknown 1Cetoniinae (1) Afr.27
Homothyrea helenae (Schaum,1848)   Djibouti 1Cetoniinae (1) Afr.18
Hoplia africana africana Escalera,1914   Morocco 2Hopliini Pal.2 (1)
Pal.4 (1)
Hoplia africana kocheri Escalera,1914   Algeria 1Hopliini Pal.2 (1)
Hoplia argentea (Poda,1761)   Austria 48Hopliini Pal.1 (48)
Bulgaria 1Hopliini Pal.1 (1)
France 60Hopliini Pal.1 (56)
Pal.3 (2) Old spms
partly poor labeling.
Pal.4 (2)
Germany 2Hopliini Pal.3 (2) Old spms
partly poor labeling.
Greece 3Hopliini Pal.1 (3)
Italy 4Hopliini Pal.1 (3)
Pal.3 (1) Old spm
poor labeling.
Sweden 2 Skåne
Hopliini Main (2)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 4Hopliini Pal.1 (4)
Unknown 9Hopliini Pal.3 (9) Old spms
partly poor labeling.
Hoplia attilioi Massa,1979   Italy 1Hopliini Pal.2 (1)
Hoplia aulica (Linnaeus,1767)   Spain 2Hopliini Pal.3 (2)
Unknown 5Hopliini Pal.3 (5)
Hoplia aureola (Pallas,1781)   Russian Federation 2Hopliini Pal.2 (1)
Pal.3 (1)
Hoplia bilineata (Fabricius,1801)   Spain 23Hopliini Pal.2 (23)
Hoplia chlorophana Erichson,1847   Spain 1Hopliini Pal.2 (1)
Hoplia coerulea (Drury,1773)   France 10Hopliini Pal.2 (3)
Pal.3 (4)
Pal.3 (3) Old spms
originally as H. farinosa auct.
Italy 6Hopliini Pal.3 (6)
Spain 10Hopliini Pal. (1)
Pal.2 (5)
Pal.4 (4)
Unknown 1Hopliini Pal.3 (1) Old spms
originally as H. farinosa auct.
Hoplia communis Waterhouse,1875   Japan 3Hopliini Pal.2 (3)
Hoplia dubia (Rossi,1790)   Italy 2Hopliini Pal.2 (2)
Hoplia graminicola (Fabricius,1792)   Germany 5Hopliini Pal.2 (5)
Unknown 2Hopliini Pal.3 (2)
Hoplia hauseri Reitter,1904   Uzbekistan 2Hopliini Pal.3 (2)
Hoplia hungarica Burmeister,1844   Bulgaria 5Hopliini Pal.2 (5)
Greece 9Hopliini Pal.2 (9)
Hoplia magnifica    Borneo, Indonesia / Malaysia 1Hopliini Or.1 (1)
Hoplia minuta (Panzer,1789)   Italy 2Hopliini Pal.2 (2)
Hoplia philanthus (Fuesslin,1775)   Austria 2Hopliini Pal.2 (2) as H. farinosa
Germany 2Hopliini Pal.2 (2)
Germany 1Hopliini Pal.2 (1) as H. farinosa
Italy 7Hopliini Pal.2 (7) as H. farinosa
Spain 11Hopliini (11) Pal.2+3
Sweden 12 Skåne
Hopliini Main (8)
Reference Coleoptera (4)
Switzerland 6Hopliini Pal.2 (6) as H. farinosa
Unknown 11Hopliini Pal.3 (11)
Hoplia pollinosa Krynicki,1832   Russian Federation 1Hopliini Pal.2 (1)
Hoplia praticola Duftschmid,1805   France 1Hopliini Pal. (1)
Hoplia pubicollis Küster,1849   Italy 6Hopliini Pal.2 (6)
Hoplia retusa Klug,1832   Madagascar 3Hopliini Afr.20 (3)
Hoplia spp.    Austria 5Hopliini Pal.4 (1) as H. farinosa
Pal.4 (4) Old spms
partly poor labeling
Croatia 1Hopliini Pal.4 (1)
Germany 2Hopliini Pal.4 (2) Old spms
partly poor labeling
Kyrgyzstan 2Hopliini Pal.4 (2)
Poland 1Hopliini Pal.4 (1) Old spm
Romania 1Hopliini Pal.4 (1)
Slovenia 2Hopliini Pal.4 (2)
Spain 2Hopliini Pal.4 (2)
Turkey 2Hopliini Pal.3 (2)
Unknown 10Hopliini Pal.4 (10) Old spms
partly poor labeling
Hoplia stenolepis Apfelbeck,1912   Greece 8Hopliini Pal.3 (8)
Hoplocnemis auriventris (Burmeister,1844)   South Africa 2Hopliini Afr.4 (2)
Hoplocnemis crassipes (Olivier,1789)   South Africa 2Hopliini (2) Afr.13
Hoplocnemis fuliginosa (Burmeister,1844)   South Africa 2Hopliini Afr.6 (2)
Hoplocnemis hylax (Fabricius,1775)   South Africa 2Hopliini (2) Afr.13
Hoplocnemis karrooana Peringuey,1908   South Africa 2Hopliini Afr.13 (1)
Type Coleoptera (1) Paralectotype 05843:2 of Hoplocnemis karrooana Peringuey 1908
  Hoplocnemis karrooana Peringuey 1908

Hoplocnemis lightfooti Peringuey,1902   South Africa 1Hopliini (1) Afr.19
Hoplocnemis mutica (Burmeister,1844)   South Africa 2Hopliini Afr.6 (2)
Hoplopus atriplicis (Fabricius,1787)   Tunisia 3Rutelinae (3) Pal.5+11 (as Anomala atriplicis F.)
Hoplopyga brasiliensis (Gory & Percheron,1833)   Ecuador 1Cetoniinae S.Am.1 (1)
Hybaloides foveolatus Quedenfeldt,1884   Central African Republic 2Orphninae Afr.1 (2)
Hybalus barbarus Castelnau,1840   Algeria 1Orphninae Pal.1 (1)
Hybalus benoitii Tournier,1864   Italy 1Scarabaeinae Pal.34 (1) (IN SCARABAEINAE)
Hybalus bigibber Reitter,1893   Italy 1Scarabaeinae Pal.25 (1) (IN SCARABAEINAE)
Hybalus graecus Sturm,1843   Morocco 1Orphninae Pal.1 (1)
Tunisia 1Orphninae Pal.1 (1)
Hybalus spp.    Tunisia 1Orphninae Pal.1 (1)
Hyboschema contractum Boheman,1857   South Africa 1Dynastinae Afr.19 (1)
Hylamorpha cylindrica Arrow,1899   Chile 2Rutelinae (2) S.Am 4
Hylamorpha elegans (Burmeister,1844)   Chile 10Rutelinae (10) S.Am 7
Hymenoplia spp.    Portugal 4Sericini Pal.1 (4)
Spain 7Sericini Pal.1 (7)
Hypselogenia geotrupina (Billberg,1817)   South Africa 2Cetoniinae Afr.9 (2)
Hypselogenia geotrupina concava (Billberg,1817)   South Africa 1Cetoniinae Afr.9 (1)
Iapurea attenuata (Harold,1870)   Mexico 8Aphodiinae N.Am.3 (8)
Idiochelyna pectoralis (Walker,1859)   Sri Lanka 1Melolonthinae Or.8 (1)
Idutywa collaris Peringuey,1902   South Africa 19Hopliini (19) Afr.19+20
Inanda trivialis Peringuey,1902   South Africa 3Hopliini (3) Afr.12
Inca besckei Schaum,1840   Brazil 3Cetoniinae S.Am.1 (3)
Inca bonplandi (Gyllenhal,1817)   Brazil 3Cetoniinae S.Am.1 (3)
Inca pulverulentus (Olivier,1789)   Brazil 4Cetoniinae S.Am.1 (4)
Ischiopsopha latreille Gory & Percheron,1833   Borneo, Indonesia / Malaysia 2Cetoniinae Or.1 (2)
Ischiopsopha lucivorax Kraatz,1890   Indonesia 3New Guinea
Cetoniinae Or.1 (3)
Ischiopsopha wallacei J.Thomson,1857   Indonesia 1Aru Islands
Cetoniinae Or.1 (1)
Ischnestoma luridipennis Burmeister,1842   South Africa 1Cetoniinae Afr.9 (1)
Ischnochelus andreaei Kulzer,1960   South Africa 2Hopliini (2) Types 2029:2-3
Ischnochelus emeritus Peringuey,   South Africa 2Hopliini (2) Afr.12
Ischnochelus fulvipennis Kulzer,1960   South Africa 2Hopliini (2) Types 2028:2-3
Ischnochelus minutus Burmeister,1844   South Africa 2Hopliini (2) Afr.12
Ischnochelus praestabilis Peringuey,   South Africa 2Hopliini (2) Afr.12
Ischnochelus puerilis Peringuey,   South Africa 2Hopliini (2) Afr.12
Ischnochelus salcatus Burmeister,1844   South Africa 2Hopliini (2) Afr.12
Ischnochelus tibialis Kulzer,1960   South Africa 2Hopliini (2) Types 2027:2-3
Ischnopopillia foveata?    China 1Rutelinae Pal.8 (1)
Korea, Republic of 1Rutelinae Pal.8 (1)
Ischnopopillia moorei Kraatz,1892   India 2Rutelinae Pal.8 (2)
Isocopris inhiata (Germar,1824)   Brazil 2Scarabaeinae S.Am.4 (2)
Isonychus arizonensis Howden,1959   United States 1Arizona
Melolonthinae N.Am.1 (1)
Ixodina abyssinicus Roth,1851   Ethiopia 3Scarabaeinae Afr.58 (3)
Kenya 11Scarabaeinae Afr.58 (11)
Tanzania 3Scarabaeinae Afr.58 (3)
Zimbabwe 2Scarabaeinae Afr.58 (2)
Ixodina endroedyi Endrödi,1976   Gambia 29Scarabaeinae Afr.58 (29)
Ixodina runicus (Arrow,1909)   Thailand 9Scarabaeinae Or.1 (9)
Ixorida celebensis Wallace,1868   Indonesia 1Sulawesi
Cetoniinae Or.4 (1)
Kheper aegyptiorum (Latreille,1827)   Ethiopia 1Scarabaeinae Afr.8 (1)
Kheper bonelli MacLeay,1821   South Africa 1Scarabaeinae (1) Afr.25
Kheper devotus Redtenbacher,   Unknown 1Scarabaeinae Or.3 (1)
Kheper festivus (Harold,1868)   Mozambique 1Scarabaeinae (1) Afr.25
Togo 1Scarabaeinae Afr.45 (1)
Kheper lemarchi (MacLeay,)   Unknown 1Scarabaeinae (1) Afr.25
Kheper nigroaeneus (Boheman,1857)   South Africa 1Scarabaeinae (1) Afr.25
Zimbabwe 3Scarabaeinae (3) Afr.25+45
Kheper nigroaeneus var.usurpator (Boheman,1857)   Mozambique 1Scarabaeinae (1) Afr.25
South Africa 1Scarabaeinae (1) Afr.25
Kheper spp.    Sierra Leone 1Scarabaeinae Afr.7 (1)
Khoina andreaei Schein,1959   South Africa 2Hopliini (2) Types 2013:2-3
Khoina bilateralis (Thunberg,1818)   South Africa 44Hopliini (44) Afr.5+19
Khoina haafi Schein,1959   South Africa 4Hopliini Afr.4 (3)
Type Schein (1) Paratype 02014:2 of Khoina haafi Schein 1959
  Khoina haafi Schein 1959

Labarrus lividus (Olivier,1789)   Greece 5Aphodiinae Pal.6 (4)
Pal.30 (1)
Portugal 4Aphodiinae Pal.28 (4)
Sweden 1 Uppland
Aphodiinae Main (1)
Turkey 2Aphodiinae Pal.6 (1)
Pal.28 (1)
United States 5Arizona
Aphodiinae N.Am.1 (1)
N.Am.8 (4)
Labarrus madagassicus (Petrovitz,1961)   Madagascar 1Aphodiinae Afr.30 (1)
Labarrus madagassicus Petrovitz,1961   Madagascar 1Aphodiinae Afr.4 (1) as Aphodius
Labarrus meruensis Petrovitz,1961   Ethiopia 1Aphodiinae Afr.3 (1) as Aphodius
Kenya 7Aphodiinae Afr.3 (7) as Aphodius
Labarrus paralividus Balthasar,1941   Ethiopia 50Aphodiinae Afr.3 (43)
Afr.27 (2)
Afr.29 (5)
Labarrus parnaguaensis (Petrovitz,1961)   Madagascar 5Aphodiinae Afr.30 (5)
Namibia 4Aphodiinae Afr.35 (4)
Labarrus pseudolividus (Balthasar,1941)   Mexico 1Aphodiinae N.Am.8 (1)
United States 2Hawaii
Aphodiinae N.Am.8 (2)
Labarrus pseudolividus Balthasar,1941   Australia 2Aphodiinae Pac.1 (2) as Aphodius
Botswana 5Aphodiinae Afr.3 (5) as Aphodius
Chile 1Aphodiinae S.Am.2 (1)
Kenya 17Aphodiinae Afr.3 (17) as Aphodius
South Africa 11Aphodiinae Afr.31 (11) as Aphodius
Labarrus sublimbatus Motschulsky,1860   China 1Aphodiinae Pal.22 (1) as Aphodius
Japan 2Aphodiinae Pal.18 (2) as Aphodius
Labarrus sublividus Balthasar,1941   Kenya 3Aphodiinae Afr.4 (3) as Aphodius
Labarrus umbratus Petrovitz,1961   Cameroon 3Aphodiinae Afr.4 (3) as Aphodius
Mozambique 2Aphodiinae Afr.24 (2) as Aphodius
Lachnota henningi (Fischer von Waldheim,1823)   Russian Federation 3Melolonthinae Pal.5 (3)
Lagochile sparsa Ohaus,1903   Panama 1Rutelinae S.Am.7 (1)
Lagochile trigona (Herbst,1790)   Ecuador 2Rutelinae S.Am.7 (2)
Lasiexis dilaticollis (Ballion,1871)   Russian Federation 3Melolonthinae Pal.5 (3)
Lasiopsis canina (Zoubkov,1829)   Russian Federation 3Melolonthinae Pal.5 (3)
Lasiotrichius succinctus (Pallas,1781)   Japan 3Cetoniinae Pal.21 (3) as Trichius s.
Latodrepanus laticollis (Fåhraeus,1857)   Gambia 2Scarabaeinae Afr.58 (2)
Kenya 2Scarabaeinae Afr.58 (2)
Leiopsammodius abyssinicus (Muller,1942)   Tanzania 1Aphodiinae Afr.21 (1)
Leiopsammodius japonicus Harold,1878   Japan 1Aphodiinae Pal.18 (1)
Leiopsammodius laevis (Paulian,1942)   Sudan 1Aphodiinae Afr.24 (1)
Leiopsammodius spp.    South Africa 8Aphodiinae Afr.16 (8)
Leiopsammodius subciliatus (Harold,1869)   Kenya 1Aphodiinae Afr.35 (1)
Leiopsammodius substriatus (Balthasar,1941)   Namibia 5Aphodiinae Afr.35 (5)
Lepidiota "bimaculata" (Sounders,1879)   Indonesia 6Java
Melolonthinae Or.9 (6)
Lepidiota elegans Brenske,1892   Borneo, Indonesia / Malaysia 1Melolonthinae Or.10 (1)
Lepidiota ferruginosa (Walker,1859)   Sri Lanka 7Melolonthinae Or.4 (7)
Lepidiota fruhstorferi Mos.   Viet Nam 3Melolonthinae Or.9 (2)
Or.10 (1)
Lepidiota stigma (Fabricius,1801)   Malaysia 1Melolonthinae Or.10 (1)
Lepidiota stigma Fabricius,1798   Indonesia 1Java
Melolonthinae Or.10 (1)
Lepisia rupicola Fabricius,1775   Unknown 3Hopliini Afr.20 (3)
Lepithrix abbreviatus Fabricius,1775   South Africa 1Hopliini Afr.4 (1)
Lepithrix dichropus Blanchard,1850   South Africa 2Hopliini Afr.4 (2)
Lepithrix freudei Schein,1959   South Africa 2Hopliini Afr.4 (2)
Lepithrix fulvipes (Thunberg,1818)   South Africa 2Hopliini Afr.4 (2) as Lepithrix pseudomodesta Schein
South Africa 18Hopliini Afr.4+20 (18) as Lepithrix modesta Peringuey
Lepithrix gentilis Peringuey,1902   South Africa 5Hopliini (5) Afr.4+20
Lepithrix hilaris Peringuey,1902   South Africa 2Hopliini Afr.4 (2)
Lepithrix kochi Schein,1959   South Africa 2Hopliini Afr.4 (2)
Lepithrix kulzeri Schein,1959   South Africa 2Hopliini Afr.4 (2) as Lepithrix freyi Schein
Lepithrix lineata (Fabricius,1775)   South Africa 19Hopliini (19) Afr.4+20
Lepithrix longitarsis Schein,1959   South Africa 4Hopliini (4) Afr.4+20 (as Lepithrix pseudolineata Schein)

South Africa 2Hopliini Afr.4 (2) as Lepithrix pseudolineata ab.nigrescens Schein
Lepithrix namaqua Peringuey,1902   South Africa 2Hopliini Afr.4 (2)
Lepithrix ornatella Peringuey,1902   South Africa 4Hopliini (4) Afr.4+20
Lepithrix pseudogentilis Schein,1959   South Africa 2Hopliini (2) Types 2012:2-3
Lepithrix stigma (DeGeer,1778)   South Africa 20Hopliini Afr.4 (20)
Lepithrix xanthoptera Burmeister,1844   South Africa 2Hopliini Afr.4 (2)
Lepitrichula lanata (Boheman,1857)   South Africa 1Hopliini Afr.4 (1)
Lepitrichula setosa (Burmeister,1844)   South Africa 2Hopliini Afr.4 (2)
Leptothyrea perroudi Schaum,1844   South Africa 1Cetoniinae Afr.5 (1)
Leucocelis adspersa Fabricius,1801   Lesotho 2Cetoniinae (2) Afr.15 (as Oxythyrea a.)
South Africa 8Cetoniinae (8) Afr.15 (as Oxythyrea a.)
Unknown 3Cetoniinae (3) Afr.27
Leucocelis aeneicollis Schaum,1844   Ethiopia 3Cetoniinae (3) Afr.27
South Africa 1Cetoniinae Afr.27 (1)
Unknown 1Cetoniinae Afr.27 (1)
Leucocelis albopilosa Kraatz,1899   Tanzania 1Cetoniinae Afr.6 (1)
Leucocelis amethystina (McLeay,1838)   Central African Republic 3Cetoniinae (3) Afr.27
South Africa 1Cetoniinae (1) Afr.6+18
Unknown 4Cetoniinae (4) Afr.27
Zimbabwe 2Cetoniinae (2) Afr.27
Leucocelis cupricollis Kraatz,1880   Zimbabwe 1Cetoniinae (1) Afr.27
Leucocelis haemorrhoidalis (Fabricius,1775)   Central African Republic 1Cetoniinae (1) Afr.27
Ethiopia 7Cetoniinae (7) Afr.27
South Africa 4Cetoniinae (4) Afr.6+27
Unknown 5Cetoniinae (5) Afr.27 (as ab. dysenterica M'Leay)
Unknown 6Cetoniinae (6) Afr.27
Leucocelis nitidula (Olivier,1789)   Malawi 2Cetoniinae Afr.6 (2)
Senegal 1Cetoniinae (1) Afr.27
Leucocelis planifrons ?   Greece 1Cetoniinae Pal.20 (1)
Leucocelis plebeja Kolbe,1895   Tanzania 2Cetoniinae Afr.6 (2)
Leucocelis rubra (Gory & Percheron,1833)   South Africa 5Cetoniinae Afr.5 (5)
South Africa 5Cetoniinae (5) Afr.15 (as Erythroderma marginata Schoch)

Unknown 2Cetoniinae (2) Afr.26
Leucocelis similis Kraatz,1896   Tanzania 3Cetoniinae Afr.6 (3)
Leucocelis spp.    Angola 1Cetoniinae Afr.32 (1)
Leucocelis testaceoguttata (Blanchard,1850)   Ethiopia 1Cetoniinae (1) Afr.27
South Africa 14Cetoniinae Afr.6 (14)
Leucocelis vitticollis Boheman,1857   Ethiopia 2Cetoniinae (2) Afr.27 (possible type as Oxythyrea v.)
Unknown 5Cetoniinae (5) Afr.27
Zimbabwe 2Cetoniinae (2) Afr.27
Leucopholis emarginata Burmeister,1855   Borneo, Indonesia / Malaysia 3Melolonthinae (3) Or.8+10
Leucopholis pinguis Burmeister,1855   Sri Lanka 9Melolonthinae (9) Or.4+5
Leucopholis plagiata Blanchard,1850   Malaysia 3Melolonthinae Or.8 (3)
Singapore 1Melolonthinae Or.8 (1)
Leucopholis rorida Fabricius,1801   Indonesia 1Sumatra
Melolonthinae Or.10 (1)
Unknown 4Melolonthinae Or.10 (4)
Leucopholis tristicula Brenske,1896   Indonesia 2Java
Melolonthinae Or.9 (2)
Leucothyreus flavipes (Eschscholtz,1822)   Brazil 1Rutelinae S.Am.7 (1)
Leucothyreus kirbyanus MacLeay,1819   Brazil 3Rutelinae S.Am.7 (3)
Leucothyreus marginaticollis Blanchard,1843   Brazil 2Rutelinae S.Am.7 (2)
Leucothyreus niveicollis Laporte,1840   Unknown 2Rutelinae S.Am.7 (2)
Leucothyreus splendidus Burmeister,1844   Trinidad and Tobago 3Trinidad
Rutelinae S.Am.7 (3)
Leucothyreus suturalis Laporte,1840   Brazil 1Rutelinae S.Am.7 (1)
Liatongus affinis (Arrow,1908)   Unknown 1Scarabaeinae Pal.8 (1)
Liatongus cf. incurvaticornis    China 1Scarabaeinae (1) Pal.34
Liatongus femoratus (Illiger,1800)   Indonesia 5Java
Scarabaeinae (5) Or.1+2
Malaysia 1Scarabaeinae Or.1 (1)
Borneo, Indonesia / Malaysia 2Scarabaeinae Or.1 (2)
Liatongus festivus (Steven,1809)   Unknown 2Scarabaeinae Pal.8 (2)
Liatongus gagatinus (Hope,1831)   Unknown 4Scarabaeinae Pal.8 (4)
Liatongus militaris (Laporte,1840)   South Africa 32Scarabaeinae (32 ) Afr.1+30
Swaziland 1Scarabaeinae Afr.3 (1)
Liatongus phanaeoides (Westwood,1839)   India 1Scarabaeinae Or.1 (1)
Japan 2Scarabaeinae (2) Pal.24
Liatongus rhadamistus (Fabricius,1775)   Thailand 2Scarabaeinae Or.4 (2) as Pseudoniticellus r.
Liatongus spathulatus (Roth,1851)   Ethiopia 9Scarabaeinae (9) Afr.1+3
Unknown 2Scarabaeinae Afr.1 (2)
Liatongus tuberculicollis (Felsche,1909)   Ethiopia 20Scarabaeinae Afr.3 (20)
Liatongus venator (Fabricius,1801)   Indonesia 1Java
Scarabaeinae Or.1 (1)
Liatongus vertagus (Fabricius,1798)   Unknown 4Scarabaeinae Or.1 (4)
Ligyrus bituberculatus (Beauvois,1805)   Colombia 8Dynastinae S.Am.15 (8)
Guatemala 1Dynastinae (1) S.Am. 32
Peru 7Dynastinae (7) S.Am.15+31+32
Trinidad and Tobago 1Trinidad
Dynastinae S.Am.15 (1)
Unknown 4Dynastinae S.Am.15 (4)
Ligyrus bituberculatus maximus (Beauvois,1805)   Brazil 1Dynastinae S.Am.15 (1)
Ligyrus burmeisteri Steinheil,1872   Argentina 2Dynastinae (2) S.Am.11+30
Ligyrus cuniculus (Fabricius,1801)   United States 1Connecticut
Dynastinae N.Am.1 (1)
Unknown 3Dynastinae N.Am.1 (3)
Ligyrus fossor (Latreille,1833)   Argentina 3Dynastinae S.Am.15 (3)
Unknown 3Dynastinae S.Am.15 (3)
Ligyrus gibbosus (DeGeer,1774)   United States 3District of Columbia
Dynastinae N.Am.1 (3)
United States 2New York
Dynastinae N.Am.1 (2)
United States 1Montana
Dynastinae N.Am.5 (1)
United States 2Arizona
Dynastinae N.Am.2 (2)
United States 1Georgia
Dynastinae N.Am.1 (1)
Unknown 2Dynastinae N.Am.1 (2)
Ligyrus gyas (Erichson,1848)   Brazil 1Dynastinae S.Am.11 (1)
Peru 1Dynastinae (1) S.Am.32
Ligyrus maimon (Erichson,1847)   Chile 1Dynastinae (1) S.Am.22
Ligyrus maternus Prell,1937   Peru 3Dynastinae (3) S.Am.5+32
Ligyrus neglectus (LeConte,1847)   United States 2South Carolina
Dynastinae N.Am.2 (2)
United States 1District of Columbia
Dynastinae N.Am.2 (1)
Unknown 11Dynastinae N.Am.2 (11)
Ligyrus relictus (Say,1825)   Canada 1Quebec
Dynastinae N.Am.4 (1)
United States 4New York
Dynastinae (4) N.Am.1+4
United States 1Montana
Dynastinae N.Am.4 (1)
United States 15Connecticut
Dynastinae (15) N.Am.4+5
Unknown 3Dynastinae N.Am.1 (3)
Ligyrus rosettae Endrödi,1968   Argentina 2Dynastinae S.Am.15 (2)
Ligyrus sallei Bates,1888   Mexico 1Dynastinae N.Am.1 (1)
United States 2Connecticut
Dynastinae N.Am.1 (2)
United States 1Illinois
Dynastinae N.Am.1 (1)
United States 2New York
Dynastinae N.Am.1 (2)
Unknown 7Dynastinae N.Am.1 (7)
Ligyrus similis Endrödi,1968   Guyana 1Dynastinae (1) S.Am.32
Ligyrus spp.    Brazil 2Dynastinae S.Am.15 (2)
Liogenys spp.    Argentina 1Melolonthinae S.Am.1 (1)
Liothorax alternatus (Horn,1870)   Canada 1Alberta
Aphodiinae N.Am.8 (1)
Liothorax innexus (Say,1835)   Mexico 1Aphodiinae N.Am.8 (1)
Liothorax kraatzi Harold,1868   Afghanistan 18Aphodiinae Pal.14 (18) as Aphodius
Greece 1Aphodiinae Pal.30 (1) as Aphodius
Turkey 1Aphodiinae Pal.6 (1) as Aphodius
Unknown 2Aphodiinae Pal.22 (2) as Aphodius
Uzbekistan 2Aphodiinae Pal.22 (2) as Aphodius
Liothorax niger (Panzer,1798)   Austria 3Aphodiinae Pal.11 (3) as Aphodius
Finland 2Aphodiinae (2) as Aphodius
Italy 5Aphodiinae Pal.6 (5) as Aphodius muscorum Adam
Russian Federation 1Aphodiinae Pal.6 (1) as Aphodius muscorum Adam
Sweden 64  Skåne Halland Öland Östergötland Västergötland Uppland Värmland
Alan Dufberg 48 (9)
Aphodiinae Main (55) as Aphodius
United Kingdom 1England
Aphodiinae Pal.15 (1) as Aphodius
Unknown 1Aphodiinae Pal.22 (1) as Aphodius
Liothorax plagiatus (Linnaeus,1767)   Denmark 3Aphodiinae (3) as Aphodius
Finland 4Aphodiinae (4) as Aphodius
Greece 2Aphodiinae Pal.6 (2) as Aphodius
Hungary 9Aphodiinae (9) Pal.16+22 as Aphodius
Russian Federation 1Aphodiinae Pal.6 (1) as Aphodius
Sweden 281  Sk Bl Ha Sm Öl Go Ög Vg Bo Vs Vr
Alan Dufberg 48 (28)
Aphodiinae Main (253) as Aphodius
Turkey 1Aphodiinae Pal.25 (1) as Aphodius
Unknown 10Aphodiinae Pal.22 (10) as Aphodius
Lissogenius conspersus Burmeister,1847   South Africa 3Cetoniinae (3) Afr.17
Loboparius auriculatus (Schmidt,1908)   South Africa 1Aphodiinae Afr.32 (1)
Loboparius globulus Harold,1859   Unknown 2Aphodiinae Pal.18 (2) as Aphodius g.
Lomaptera adelpha Thomson,1857   Indonesia 1Aru Islands
Cetoniinae Or.4 (1)
Lonchotus latus Arrow,1911   Madagascar 1Dynastinae Afr.7 (1)
Lorditomaeus aequus Schmidt,1908   Kenya 2Aphodiinae Afr.27 (2)
Lorditomaeus bifidus Schmidt,1908   Gambia 19Aphodiinae Afr.13 (19)
Ghana 23Aphodiinae Afr.17 (23)
Lorditomaeus gridellii Endrödi,1964   Kenya 1Aphodiinae Afr.23 (1) Paratype. As Aphodius g.
Lorditomaeus harpalinus (Gerstaecker,1884)   Kenya 1Aphodiinae Main.coll.types (1) Syntype: 577 of Brachiaphodius aureopilosus Boucomont 1930
  Brachiaphodius aureopilosus Boucomont 1930

Lorditomaeus horni (Balthasar,1937)   Tanzania 2Aphodiinae Afr.30 (2)
Zambia 1Aphodiinae Afr.30 (1)
Lorditomaeus infuscatus Schmidt,1908.   Ethiopia 1Aphodiinae Afr.13 (1)
Kenya 6Aphodiinae (6) Afr.13+16
Lorditomaeus invenustus Schmidt,1908   Ghana 26Aphodiinae Afr.17 (26)
Lorditomaeus lunulatus Schmidt,1908   Kenya 1Aphodiinae Afr.13 (1)
Tanzania 1Aphodiinae Afr.13 (1)
Lorditomaeus opatroides (Klug,1855)   Tanzania 2Aphodiinae Afr.13 (1)
Afr.30 (1)
Lorditomaeus spp.    Ethiopia 1Aphodiinae Afr.17 (1) as L. bordati n.sp.
Ghana 4Aphodiinae Afr.17 (4) as L. collartoides n.sp.
originaly as L. opatroides
Lorditomaeus tenuis Schmidt,1908   Gambia 1Aphodiinae Afr.13 (1)
Lycomedes buckleyi Waterhouse,1880   Colombia 29Dynastinae (29) S.Am.1+2+3+4
Ecuador 1Dynastinae S.Am.3 (1)
Macraspis chalcea Burmeister,1844   Trinidad and Tobago 3Trinidad
Rutelinae (3) S.Am 4
Macraspis chrysis (Linnaeus,1764)   Colombia 2Rutelinae (2) S.Am 4
Grenada 1Rutelinae (1) S.Am 4
Unknown 1Rutelinae (1) S.Am 4
Macraspis cincta (Drury,1782)   Brazil 10Rutelinae (10) S.Am 5
Unknown 3Rutelinae (3) S.Am 5
Macraspis clavata (Olivier,1789)   Brazil 1Rutelinae (1) S.Am 5
Unknown 3Rutelinae (3) S.Am 5
Macraspis cupripes Kirsch.,1870   Colombia 1Rutelinae (1) S.Am 5
Unknown 2Rutelinae (2) S.Am 5
Macraspis dichroa Mannerheim,1829   Brazil 3Rutelinae (3) S.Am 5
Unknown 1Rutelinae (1) S.Am 5
Macraspis gagatina (Waterhouse,1881)   Brazil 1Rutelinae (1) S.Am 6
Ecuador 1Rutelinae (1) S.Am 6
Macraspis hirtiventris Bates,1888   Unknown 1Rutelinae (1) S.Am 5
Macraspis lateralis (Olivier,1789)   Suriname 1Rutelinae (1) S.Am 5
Macraspis mixta (Blanchard,1850)   Unknown 1Rutelinae (1) S.Am 5
Macraspis morio Burmeister,1844   Brazil 3Rutelinae (3) S.Am 6
Unknown 3Rutelinae (3) S.Am 6
Macraspis pantochloris (Blanchard,1850)   Brazil 2Rutelinae (2) S.Am 6
Unknown 2Rutelinae (2) S.Am 6
Macraspis pseudochrysis Landin,1956   Brazil 1Type Coleoptera (1) Holotype 03251:1 of Macraspis pseudochrysis Landin 1956
  Macraspis pseudochrysis Landin 1956

Macraspis tetradactyla (Linnaeus,1771)   Brazil 3Rutelinae (3) S.Am 4
Unknown 1Rutelinae (1) S.Am 4
Macraspis thoracica Mannerheim,1829   Brazil 3Rutelinae (3) S.Am 5
Unknown 3Rutelinae (3) S.Am 5
Macrodactylus silaonus Bates,1887   Mexico 7Melolonthinae N.Am.2 (7)
Macrodactylus subspinosa (Fabricius,1775)   Canada 10Ontario
Melolonthinae N.Am.2 (10)
United States 2Illinois
Melolonthinae N.Am.2 (2)
United States 1Maryland
Melolonthinae N.Am.2 (1)
United States 8Massachusetts
Melolonthinae N.Am.2 (8)
Macrodactylus uniformis Horn,1876   United States 1Arizona
Melolonthinae N.Am.1 (1)
Macrodactylus variipes Bates,1887   Mexico 1Melolonthinae N.Am.1 (1)
Macropopillia arrowi Ohaus,1905   India 4Rutelinae Or.8 (4)
Macroretrus confusus Harold,1862   Somalia 2Aphodiinae Afr.27 (2)
Macrotetrus singularis Peringuey,1908   Tanzania 1Aphodiinae Afr.13 (1)
Madotrogus glabricollis (Reitter,1888)   Russian Federation 1Melolonthinae Pal.5 (1)
Madotrogus glabripennis (Ballion,1871) (?)   Russian Federation 1Melolonthinae Pal.5 (1)
Madotrogus kirgisicus (Semenov,1895)   Kyrgyzstan 1Melolonthinae Pal.9 (1)
Maechidius major Blackburne,1888   Australia 9Western Australia
Melolonthinae Pac.1 (9)
Maladera anderssoni Ahrens in litt.,   Sri Lanka 2Sericini Or.2 (2) 2 paratypes
Maladera arenicola Solsky,1876   Russian Federation 1Sericini (1) Pal.1 (as Leucoserica a.)

Turkmenistan 1Sericini Pal.2 (1)
Syr Darya, Kyrgyzstan / Uzbekistan / Tajikistan / Kazakhstan 1Sericini Pal.2 (1)
Maladera atratula Dalla Torre,1912   Unknown 2Sericini Pal.2 (2)
Maladera breviata (Brenske,1898)   Sri Lanka 4Sericini Or.2 (4)
Maladera calcarata (Brenske,1898)   Sri Lanka 1Sericini Or.2 (1)
Maladera cardoni (Brenske,1896)   Sri Lanka 2Sericini Or.1 (2)
Maladera castanea (Arrow,1913)   Japan 2Sericini Pal.2 (2)
Maladera cinnaberina (Brenske,1898)   Sri Lanka 2Sericini Or.2 (2)
Maladera euphorbiae (Burmeister,1855)   Russian Federation 2Sericini Pal.1 (2)
Syr Darya, Kyrgyzstan / Uzbekistan / Tajikistan / Kazakhstan 1Sericini Pal.2 (1)
Maladera gibbiventris (Brenske,1897)   Korea, Republic of 1Sericini Pal.1 (1)
Maladera graeca Petrovitz,1969   Greece 1Sericini Pal.2 (1)
Maladera holosericea (Scopoli,1772)   Denmark 1Sericini Main (1)
France 1Sericini Pal.1 (1)
Germany 12Sericini Pal.2 (12)
Hungary 1Sericini Pal.2 (1)
Poland 4Sericini Pal.2 (4)
Romania 3Sericini Pal.1 (3)
Sweden 71  Skåne
Alan Dufberg 48 (11)
Sericini Main (60)
Unknown 1Sericini Pal.2 (1)
Maladera hortonensis Ahrens in litt.,   Sri Lanka 1Sericini (1) 1 paratype Or.2
Maladera insanabilis (Brenske,1894)   Afghanistan 1Sericini (1) Pal.1 (as Autoserica I)

Afghanistan 1Sericini (1) Pal.1 (as Autoserica adjuncta Brenske)

Maladera insanabilis (Brenske,1894) (Cephaloserica)   Sri Lanka 1Sericini Or.1 (1)
Maladera japonica (Motschulsky,1860)   Japan 2Sericini Pal.2 (2)
Maladera lindulana Ahrens in litt.,   Sri Lanka 3Sericini (3) 3 paratypes Or.2
Maladera marginella (Hope,1831)   Kashmir: Pakistan / India / China 1Sericini Or.2 (1)
Maladera mollis (Walker,1859)   Sri Lanka 22Sericini Or.1 (22)
Maladera nagporeana (Brenske,1898)   Afghanistan 13Sericini Pal.1 (13)
Maladera orientalis (Motschulsky,1858)   China 3Sericini (3) Pal.1+2
Japan 1Sericini Pal.2 (1)
Maladera pubescens (Arrow,1916)   Sri Lanka 74Sericini Or.1 (74)
Maladera punctatissima Faldermann,1835   Iran 1Sericini Pal. (1)
Maladera rotundata (Walker,1859)   Sri Lanka 2Sericini Or.2 (2)
Maladera rufocuprea (Blanchard,1850)   Sri Lanka 13Sericini Or.1 (13)
Maladera spp.    China 4Sericini Pal.1 (4)
Korea, Republic of 1Sericini Pal.1 (1)
Malaysia 2Sericini Or.2 (2)
Russian Federation 2Sericini Pal.1 (2)
Sri Lanka 5Sericini Or.2 (5)
Borneo, Indonesia / Malaysia 1Sericini Or.2 (1)
Maladera verticalis (Fairmaire,1888)   China 2Sericini Pal.1 (2)
Malagoniella punctatostriata Blanchard,1843   Unknown 2Scarabaeinae S.Am.9 (2) From Cordoba
Malagoniella spp.    Unknown 3Scarabaeinae S.Am.9 (3)
Malagoniella virens (Harold,1869)   Argentina 1Scarabaeinae S.Am.9 (1)
Unknown 1Scarabaeinae S.Am.9 (1)
Manopus biguttatus Laporte,1840   Colombia 1Melolonthinae S.Am.1 (1)
Marmylida impressa (Goldfuss,1805)   Mozambique 1Cetoniinae Afr.4 (1)
South Africa 2Cetoniinae Afr.4 (2)
Marmylida rufa (DeGeer,1778)   South Africa 1Cetoniinae Afr.3 (1)
Martineziella dutertrei (Chalumeau,1983)   United States 3Mississippi
Aphodiinae N.Am.5 (3)
Martineziella excavaticollis (Blanchard,1843)   United States 3Georgia
Aphodiinae N.Am.2 (3)
United States 2Florida
Aphodiinae N.Am.2 (2)
Mausoleopsis amabilis (Schaum,1844)   Mozambique 1Cetoniinae (1) Afr.26
Namibia 1Cetoniinae (1) Afr.15
South Africa 3Cetoniinae (3) Afr.15+26
Tanzania 21Cetoniinae Afr.6 (21)
Unknown 3Cetoniinae (3) Afr.26
Mazoe albonotata (Peringuey,1885)   South Africa 3Cetoniinae Afr.9 (3)
Mecinonota regia (Fabricius,1801)   Indonesia 1Java
Cetoniinae Or.4 (1)
Megaceras morpheus Burmeister,1847   Colombia 27Dynastinae (27) S.Am.1+2+3+4
Ecuador 1Dynastinae S.Am.3 (1)
Unknown 5Dynastinae (5) S.Am.2+3
Megaceras septentrionis Bates,1888   Unknown 1Dynastinae S.Am.1 (1)
Megaceras teucer Burmeister,1847   Colombia 1Dynastinae S.Am.1 (1)
Megalorrhina harrisi Westwood,1847   Tanzania 4Cetoniinae (4) Afr.19
Unknown 1Cetoniinae (1) Afr.19
Megapertha massageta (Kirsch,1880)   Afghanistan 1Rutelinae (1) Pal.3 (as Phyllopertha m)

Turkmenistan 2Rutelinae (2) Pal.3 (as Phyllopertha m)

Megasoma actaeon (Linnaeus,1758)   Colombia 2Dynastinae (2) S.Am.27
Unknown 2Dynastinae (2) S.Am.26+27
Megasoma anubis (Chevrolat,1836)   Colombia 2Dynastinae (2) S.Am.27
Unknown 1Dynastinae (1) S.Am.24
Megasoma elephas (Fabricius,1775)   Colombia 1Dynastinae (1) S.Am.26
Mexico 2Dynastinae N.Am.3 (2)
Unknown 3Dynastinae (3) S.Am.25+26
Megasoma gyas (Jablonsky,1785)   Unknown 3Dynastinae (3) S.Am.28
Melinesthes algoensis flavipennis Westwood,1843   South Africa 3Cetoniinae Afr.9 (3)
Melinesthes exilis Janson,1885   South Africa 3Cetoniinae Afr.9 (3)
Melinopterus abeillei (Sietti,1903)   Spain 3Aphodiinae Pal.11 (3)
Melinopterus consputus (Creutzer,1799)   Croatia 2Aphodiinae Pal.20 (2)
Greece 1Aphodiinae Pal.20 (1)
Hungary 3Aphodiinae Pal.20 (3)
Syria 1Aphodiinae Pal.20 (1)
Tunisia 1Aphodiinae Pal.10 (1)
Melinopterus prodromus (Brahm,1790)   Denmark 6Aphodiinae Main (6)
Sweden 642  Sk Bl Ha Sm Öl Go Ög Vg Sö Up Vs Vr Dr Hs Jä
Alan Dufberg 47 (21)
Aphodiinae Main (621)
Melinopterus punctatosulcatus (Sturm,1805)   Denmark 1Aphodiinae Main (1)
Russian Federation 5Aphodiinae Pal.6 (5)
Sweden 176  Sk Sm Öl Go Ög Up Vr Dr Jä Hs Nb Lu
Alan Dufberg 47 (29) as A. sabulicola Thomson
1868
Aphodiinae Main (142)
Type Coleoptera 4955 (5) Lectotype Paralectotype 04956:1-5 of Aphodius sabulicola Thomson 1868
  Aphodius sabulicola Thomson 1868

Melinopterus reyi (Reitter,1892)   Italy 4Aphodiinae Pal.15 (4)
Melinopterus sphacelatus (Panzer,1798)   Norway 1Aphodiinae Main (1)
Sweden 320  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up Vr Dr Gä Jä
Alan Dufberg 47 (6)
Aphodiinae Main (314)
Melinopterus spp.    Tunisia 1Aphodiinae Pal.28 (1)
Melinopterus tingens (Reitter,1892)   France 1Aphodiinae Pal.20 (1)
Spain 5Aphodiinae Pal.11 (1)
Pal.20 (1)
Pal.28 (3)
Melinopterus villarreali (Baraud,1975)   Spain 4Aphodiinae Pal.11 (4)
Melolontha albida Frivaldszky,1835   Syria 1Melolonthinae Pal.1 (1)
Turkey 2Melolonthinae Pal.15 (1)
Pal.16 (1)
Melolontha excisicauda Balthasar,1936   Turkey 1Melolonthinae Pal.16 (1)
Melolontha hippocastani Fabricius,1801   Denmark 2Melolonthinae Main (2)
Germany 5Melolonthinae (4) Pal.1+10
Pal.17 (1)
Russian Federation 2Melolonthinae Pal.8 (2)
Sweden 137  Sk Bl Ha Sm Go GS Ög Vg Bo Nä Sö Up Vr Dr Gä Ån
Alan Dufberg 48 (1)
Melolonthinae Main (136)
Turkey 3Melolonthinae Pal.16 (3)
Unknown 4Melolonthinae Pal.17 (4)
Melolontha kraatzi Reitter,1906   Iran 4Melolonthinae Pal.16 (4)
Melolontha melolontha (Linnaeus,1758)   France 4Melolonthinae (4) Pal.8+11
Germany 11Melolonthinae (10) Pal.1+8+10+11
Pal.17 (1)
Italy 1Melolonthinae Pal.11 (1)
Sweden 248  Skåne Blekinge Halland Småland Öland Östergötland Västergötland Södermanland
Alan Dufberg 48 (6)
Melolonthinae Main (242)
Turkey 1Melolonthinae Pal.1 (1)
United Kingdom 1Melolonthinae Pal.18 (1)
Unknown 1Melolonthinae Pal.17 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Melolonthinae Pal.1 (1)
Melolontha pectoralis Megerle von Muhlfeld,1812   Armenia 1Melolonthinae Pal.16 (1)
Germany 2Melolonthinae Pal.1 (2)
Turkey 2Melolonthinae Pal.16 (2)
Melolontha spp.    Greece 1Melolonthinae Pal.13 (1)
Uzbekistan 2Melolonthinae Pal.8 (2)
Mendidaphodius linearis (Reiche & Saulcy,1856)   Turkey 19Aphodiinae Pal.10 (4) as Aphodius
Pal.25 (1) as Aphodius
Pal.27 (7) as Aphodius
Pal.30 (7) as Aphodius
Unknown 2Aphodiinae Pal.22 (2) as Aphodius
Mendidaphodius rutilus (Klug,1845)   Ethiopia 1Aphodiinae Afr.29 (1)
Mendidius incommunis Fall,1932   United States 1Colorado
Aphodiinae N.Am.2 (1)
Mendidius multiplex Reitter,1897   Afghanistan 2Aphodiinae Pal.8 (2)
Mendidius palmenticolus Karsch,1881   Spain 1Canary Islands
Aphodiinae Pal.3 (1) as A. granulifrons Fairm.
Mendidius spp.    Tian Shan 1Aphodiinae Pal.28 (1)
Mericserica oberthuri (Brenske,1897)   India 1Sericini Or.2 (1)
Meroloba suturalis (Vollenhoven,11858)   Thailand 1Cetoniinae Or.4 (1)
Mesontoplatys arabicus (Harold,1875)   Oman 4Aphodiinae Pal.7 (4)
Mesontoplatys dorsalis (Klug,1855)   Botswana 4Aphodiinae Afr.9 (4)
Lesotho 2Aphodiinae Afr.24 (2)
Namibia 3Aphodiinae Afr.24 (3)
Sierra Leone 2Aphodiinae Afr.15 (2)
South Africa 2Aphodiinae Afr.23 (2)
Mesontoplatys effetus (Kolbe,1908)   Zambeze river 4Aphodiinae Afr.20 (4)
Mesontoplatys indoralis (Endrödi,1960)   Botswana 2Aphodiinae Afr.9 (2)
Mesontoplatys parvulus (Harold,1871)   Tanzania 1Aphodiinae Afr.9 (1)
Mesontoplatys probes (Péringuey,1901)   Ethiopia 6Aphodiinae Afr.29 (6)
Mesontoplatys rougoni (Petroviz,1974)   Gambia 1Aphodiinae Afr.9 (1)
Mesontoplatys sunantae (Stebnicka,1981)   Thailand 1Aphodiinae Or.3 (1)
Mesontoplatys triangularis (Schmidt,1907)   South Africa 1Aphodiinae Afr.25 (1)
Metacatharsius abortivus Fairmaire,1881   Congo 1Scarabaeinae Afr.15 (1)
Metacatharsius exiguiformis Ferreira,1964   South Africa 3Scarabaeinae Afr.9 (3) as Catharsius e.
Metacatharsius exiguus Boheman,1860   South Africa 2Scarabaeinae Afr.36 (2) as Catharisus e.
Metacatharsius inermis Castelnau,1840   India 1Scarabaeinae Or.5 (1)
Senegal 1Scarabaeinae Afr.46 (1)
Metacatharsius latifrons Harold,1868   South Africa 3Scarabaeinae Afr.36 (3) as Catharsius l.
Metacatharsius marani Balthasar, 1940   South Africa 2Scarabaeinae Afr.9 (2) as Catharsus m.
Metacatharsius opacus Waterhouse,1891   Namibia 1Scarabaeinae Afr.36 (1) as Catharsius o.
South Africa 1Scarabaeinae Afr.36 (1) as Catharsius o.
Metacatharsius spp.    Ethiopia 2Scarabaeinae Afr.6+12 (2) as Catharsius spp
Gambia 1Scarabaeinae Afr.12 (1) as Catharsius spp
Guinea-Bissau 4Scarabaeinae Afr.12+45 (4) as Catharsius spp
Guinea-Bissau 1Scarabaeinae Afr.11 (1) as Catharsius spp.
Kenya 2Scarabaeinae Afr.12 (2) as Catharsius spp
Senegal 1Scarabaeinae Afr.12 (1) as Catharsius spp
Tanzania 2Scarabaeinae Afr.12+45 (2) as Catharsius spp
Metacatharsius troglodytes Boheman,1857   South Africa 17Scarabaeinae Afr.35 (17) as Catharsius t.
Micraegialia pusilla (Horn,1887)   Canada 1British Columbia
Aphodiinae N.Am.4 (1)
Micranomala cingalensis Arrow,1911   Sri Lanka 1Rutelinae Or.8 (1)
Micranomala spp.    Sri Lanka 1Rutelinae Or.5 (1)
Micrelaphinis adspersula Gerstaecker,1884   Kenya 1Cetoniinae (1) Afr.25
Micrelaphinis irrorata (Fabricius,1798)   South Africa 2Cetoniinae (2) Afr.14+25
Unknown 3Cetoniinae (3) Afr.25
Micrelaphinis spp.    South Africa 2Cetoniinae (2) Afr.26
Unknown 1Cetoniinae (1) Afr.26
Microcopris reflexus (Fabricius,1787)   Viet Nam 1Scarabaeinae Or.3 (1)
Micropertha variabilis (Ballion,1871)   Turkey 1Rutelinae (1) Pal.6 (as Phyllopertha v.)

Uzbekistan 3Rutelinae (3) Pal.6 (as Phyllopertha v.)

Microserica compressipes (Wiedemann,1823)   Indonesia 4Java
Sericini Or.2 (4)
Microserica semperi (Brenske,1894)   Philippines 1Sericini Or.2 (1)
Microserica spp.    Malaysia 1Sericini Or.2 (1)
Microthopus castanopterus Burmeister,1855   Australia 1Western Australia
Melolonthinae Pac.1 (1)
Microvalgus spp.    Australia 1Cetoniinae Pac.1 (1)
Milichus spp.    Ethiopia 1Scarabaeinae Afr.3 (1)
Kenya 1Scarabaeinae (1) Afr.18
Tanzania 1Scarabaeinae (1) Afr.18
Mimela aurata (Fabricius,1801)   Austria 5Rutelinae (5) Pal 1+6
Germany 1Rutelinae Pal.9 (1)
Mimela chinensis Kirby,1823   China 3Rutelinae (3) Pal 6
Mimela confucius Hope,1836   China 1Rutelinae (1) Pal 6
India 15Rutelinae Or.7 (15)
Japan 1Rutelinae (1) Pal 6
Mimela costata (Hope,1839)   Japan 3Rutelinae (3) Pal 6
Japan 1Rutelinae (1) Pal.10 (as Anomala d.)
Mimela difficilis (Waterhouse,1875)   Japan 2Rutelinae (2) Pal.4 (as Anomala d.)
Japan 2Rutelinae Pal.10 (2)
Mimela flavilabris (Waterhouse,1875)   Japan 2Rutelinae Pal.10 (2)
Mimela heterochropus Blanchard,1851   India 4Rutelinae Or.7 (4)
Mimela holosericea (Fabricius,1787)   Japan 5Rutelinae (5) Pal 1+6
Mongolia 1Rutelinae (1) Pal 6
Russian Federation 2Rutelinae (2) Pal 6
Mimela horsfieldi Hope,1836   China 1Rutelinae Pal.6 (1)
Japan 1Rutelinae Pal.6 (1)
Mimela inscripta Nonfried,1891   Thailand 4Rutelinae Or.5 (4)
Mimela junii (Duftschmied,1805)   Germany 2Rutelinae Pal.1 (1)
Pal.9 (1)
Italy 5Rutelinae (5) Pal 1
Mimela junii calabrica (Duftschmied,1805)   Italy 1Rutelinae Pal.10 (1)
Mimela mundissima Walker,1859   Sri Lanka 1Rutelinae Or.6 (1)
Mimela ohausi Arrow,1908   India 1Rutelinae Or.7 (1)
Mimela peckinensis (Heyden,1886)   China 1Rutelinae (1) Pal 6
Japan 1Rutelinae (1) Pal 6
Unknown 3Rutelinae (3) Pal 6
Mimela pectoralis Blanchard,1851   India 1Rutelinae Or.7 (1)
Mimela rufoprasina Ohaus,1902   Cameroon 4Type Ohaus (4) Paratype 03250:2-5 of Mimela rufoprasina Ohaus 1902
  Mimela rufoprasina Ohaus 1902

Mimela splendens (Gyllenhal,1817)   Japan 2Rutelinae Pal.10 (2) as M. lucidula Hope
Korea, Republic of 2Rutelinae (2) Pal 6
Mimela testaceipes (Motschulsky,1860)   Japan 41Rutelinae (41) Pal 5
Mimela testaceoviridis Blanchard,1851   China 1Rutelinae (1) Pal 6
Japan 1Rutelinae (1) Pal 6
Korea, Republic of 1Rutelinae (1) Pal 6
Mimela vittaticollis Burmeister,1855   India 1Rutelinae Or.7 (1)
Miridiba formosana (Moser,1910)   China 2Melolonthinae Pal.5 (2)
Mitrophorus natalensis Peringuey,1902   South Africa 8Hopliini (8) Afr.12
Mucrencya mucronata (Klug,1832)   Madagascar 2Melolonthinae Afr.4 (2)
Musurgus stridens Vauloger,1898   Tunisia 1Dynastinae Pal.2 (1)
Unknown 1Dynastinae N.Am.2 (1)
Mycteristes rhinophyllus (Wiedemann,1823)   Indonesia 2Java
Cetoniinae Or.2 (2)
Mycterophallus spp.    Papua New Guinea 1Cetoniinae Or.7 (1)
Myrhessus afghanus (Balthasar,1939)   Afghanistan 2Aphodiinae Pal.9 (2) as Rhyssemus a.
Myrhessus samurai (Balthasar,1941)   Japan 1Aphodiinae Pal.28 (1)
Myrmecaphodius proseni Martinez,1952   Argentina 1Aphodiinae S.Am.2 (1)
Neateuchus proboscideus (Guerin,1884)   South Africa 8Scarabaeinae (8) Afr.25+26
Neocalaphodius moestus Fabricius,1801   Botswana 1Aphodiinae Afr.9 (1)
Kenya 2Aphodiinae Afr.9 (2)
Madagascar 1Aphodiinae Afr.9 (1)
Malaysia 8Aphodiinae Or.3 (8)
Namibia 2Aphodiinae Afr.23 (1)
Afr.35 (1)
South Africa 6Aphodiinae Afr.23 (4)
Afr.31 (2)
Sri Lanka 29Aphodiinae (29) Or.1+6
Neocolobopterus abyssinicus (Harold,1861)   Ethiopia 2Aphodiinae Afr.1 (1)
Afr.19 (1)
Neocolobopterus ballioni (Schmidt,1906)   Ethiopia 56Aphodiinae Afr.1 (53)
Afr.27 (2)
Afr.29 (1)
Neocolobopterus kivuanus (Endrödi,1956)   Kenya 9Aphodiinae Afr.1 (4)
Afr.35 (5)
Neocolobopterus maculicollis (Reiche,1847)   Cameroon 2Aphodiinae Afr.19 (2)
Ethiopia 7Aphodiinae Afr.1 (5)
Afr.27 (2)
Gambia 12Aphodiinae Afr.1 (12)
Kenya 1Aphodiinae Afr.1 (1)
Nigeria 3Aphodiinae Afr.28 (3)
South Africa 24Aphodiinae Afr.19 (11)
Afr.32 (13)
Tanzania 2Aphodiinae Afr.19 (2)
Unknown 1Aphodiinae Afr.19 (1)
Neocolobopterus stefenelli (Dellacasa,1984)   Kenya 22Aphodiinae Afr.35 (22)
Neodon pecuarius (Reiche,1860)   Australia 6Dynastinae Pac.2 (6)
Neoheptaulacus puberulus (Boheman,1857)   Tanzania 1Aphodiinae Afr.1 (1)
Neoserica bombycina (Karsch,1882)   Sri Lanka 1Sericini Or.1 (1)
Neoserica parursina Ahrens,2003   China 1Sericini Pal.1 (1)
Neoserica sexfoliata Moser,1815   Sri Lanka 3Sericini Or.2 (3)
Neosisyphus barbarossa (Wiedemann,1823)   Lesotho 1Scarabaeinae Afr.31 (1) as Sisyphus rugosus Gory
1833
South Africa 2Scarabaeinae Afr.31 (2) as Sisyphus rugosus Gory
1833
Neosisyphus calcaratus (Klug,1855)   Lesotho 1Scarabaeinae Afr.31 (1) as Sisyphus rubripes Boheman
1857
South Africa 1Scarabaeinae Afr.31 (1) as Sisyphus rubripes Boheman
1857
Neosisyphus infuscatus (Klug,1855)   Tanzania 2Scarabaeinae Afr.23 (2) as Sisyphus i.
Neosisyphus quadricollis Gory,1833   Tanzania 1Scarabaeinae Afr.23 (1) as Sisyphus q.
Neosisyphus regnardi Alluaud,1898   Mauritius 1Scarabaeinae Afr.57 (1)
Neosisyphus spinipes (Thunberg,1818)   Ethiopia 9Scarabaeinae Afr.6 (9)
South Africa 1Scarabaeinae Afr.32 (1) as S. atratus Klug
1855
Unknown 1Scarabaeinae Afr.23 (1) as Sisyphus s.
Neosisyphus spp.    Gambia 4Scarabaeinae (4) Afr.20
Neptunides polychrous J. Thomson,1879   Tanzania 1Cetoniinae Afr.32 (1)
Unknown 2Cetoniinae Afr.22 (1)
Afr.32 (1)
Nialaphodius nigrita (Fabricius,1801)   Botswana 2Aphodiinae Afr.3 (2)
Ethiopia 20Aphodiinae Afr.29 (20)
Gambia 43Aphodiinae Afr.3 (43)
Kenya 5Aphodiinae Afr.3 (5)
Namibia 6Aphodiinae Afr.23 (5)
Afr.35 (1)
Senegal 3Aphodiinae Afr.3 (3)
Sierra Leone 81Aphodiinae (81) Afr.15
South Africa 6Aphodiinae Afr.23 (2)
Afr.34 (4)
Nialus digittatus Harold,1871   Afghanistan 7Aphodiinae Pal.7 (7)
Nialus varians Duftschmid,1805   Afghanistan 2Aphodiinae Pal.13 (2)
Bulgaria 33Aphodiinae Pal.5 (15)
Pal.29 (18)
Germany 2Aphodiinae Pal.15 (2)
Turkey 8Aphodiinae (7) Pal.5+16
Pal.28 (1)
Niphetophora carneola (Burmeister,1847)   Tanzania 1Cetoniinae (1) Afr.18
Niphetophora hildebrandti (Harold,1878)   South Africa 13Cetoniinae (13) Afr.16+17
Niphetophora squamipes (Gerstaecker,1883)   Cameroon 3Cetoniinae Afr.7 (3)
Nipponoserica peregrina Chapin,1938   United States 1New York
Sericini N.Am.1 (1)
Nobius hepaticus (Roth,1851)   Ethiopia 5Aphodiinae Afr.4 (5) as Aphodius
Kenya 1Aphodiinae Afr.27 (1) as Aphodius
Lesotho 1Aphodiinae Afr.23 (1) as Aphodius
South Africa 2Aphodiinae Afr.23 (1) as Aphodius
Afr.25 (1) as Aphodius
Nobius serotinus (Panzer, 1799)   Austria 3Aphodiinae Pal.21 (3) as Aphodius
Finland 1Aphodiinae Main (1) as Aphodius
Nobius timidus (Boheman,1857)   Namibia 1Aphodiinae Afr.35 (1)
Zimbabwe 1Aphodiinae Afr.35 (1)
Nolicus veteranus (Petrovitz,1962)   Namibia 1Aphodiinae Afr.35 (1)
Notocaulus hurneri Machatschke,1955   Congo 2Aphodiinae Afr.21 (2) Incl. 1 paratype
Notocaulus laticollis Arrow,1906   Kenya 1Aphodiinae (1) Afr.25
Notocaulus muansicus Machatscke,1955   Tanzania 1Aphodiinae (1) 1 paratype Afr.20
Notocaulus muansicus kilimandjarensis Machatscke,1955   Tanzania 1Aphodiinae (1) 1 paratype Afr.20
Notocaulus neumanni Felsche,1909   Ethiopia 3Aphodiinae Afr.13 (3)
Tanzania 1Aphodiinae Afr.21 (1)
Notocaulus nigropiceus Quedenfeldt,1884   Gambia 2Aphodiinae (2) Afr.13
Guinea 2Aphodiinae (2) Afr.25
Nigeria 1Aphodiinae Afr.28 (1)
Notocaulus schoutedeni Boucomont,1928   Congo 2Aphodiinae (2 ) Afr.26 (incl. 1 paratype)

Novapus spp.    Australia 1Dynastinae Pac.2 (1)
Odochilus haroldi Racovic,1987   Borneo, Indonesia / Malaysia 1Aphodiinae Or.2 (1)
Odontolochus burgeoni Boucomont,1929   Chad 1Aphodiinae (1) Afr.24
Congo 2Aphodiinae (2 ) Afr.26 (incl. 1 paratype)

Odontolochus gestroi (Clouet,1900)   Ghana 1Aphodiinae (1) Afr.17
Sierra Leone 1Aphodiinae (1) Afr.14
Odontolochus oberthuri (Clouet,1900)   Congo 3Aphodiinae (3) Afr.26
Gambia 23Aphodiinae (23) Afr.13
Ghana 1Aphodiinae (1) Afr.17
Senegal 9Aphodiinae (9) Afr.26
Odontolochus paucus Schm.   Kenya 1Aphodiinae (1) Afr.24
Odontolochus raffrayi Paulian,1942   Côte D'ivoire 1Aphodiinae (1) Afr.24
Odontolochus spinicollis (Harold,1871)   Gambia 2Aphodiinae (2) Afr.13
Odontolochus spp.    Côte D'ivoire 1Aphodiinae (1) Afr.24
Odontolochus villiersi Petr.   Gambia 1Aphodiinae (1) Afr.13
Sierra Leone 1Aphodiinae (1) Afr.13
Odontoloma pauxillum Boheman,1857   South Africa 1Scarabaeinae (1) Afr.32
Odontoloma pygidiale Peringuey,1901   Lesotho 1Scarabaeinae (1) Afr.32
Odontorrhina pubescens (Olivier,1789)   South Africa 3Cetoniinae (3) Afr.14
Unknown 1Cetoniinae (1) Afr.22
Omaloplia diabolica Reitter,1887   Turkey 1Melolonthinae Pal.14 (1) as Homaloplia d.
Omaloplia erythroptera Frivaaldszky,1835   Bulgaria 18Sericini Pal.1 (18)
Omaloplia graeca (Reitter,1887)   Greece 1Sericini Pal.2 (1)
Omaloplia hericia (Chobaut,1907)   Greece 1Sericini Pal.2 (1)
Omaloplia nigromarginata (Herbst,1785)   Germany 2Sericini Pal.2 (2)
Omaloplia ottomana (Baraud,1965)   Turkey 11Melolonthinae Pal.13 (1)
Pal.15 (10)
Omaloplia ruricola (Fabricius,1775)   Czech Republic 1Sericini Pal.2 (1)
Germany 3Sericini Pal.1 (3)
Omocrates axillaris Burmeister,1844   South Africa 2Hopliini (2) Afr.12
Omocrates cylindricus Burmeister,1844   South Africa 1Hopliini (1) Afr.12
Omocrates depressus Blanchard,1850   South Africa 2Hopliini (2) Afr.12
Omocrates karrooanus Schein,1958   South Africa 2Hopliini (2) Types2032:2-3
Omocrates placens Peringuey,1902   South Africa 1Hopliini (1) Afr.12
Omocrates pygidialis Schein,1958   South Africa 2Hopliini (2) Types2033:2-3
Omocrates spatulipennis Blanchard,1850   South Africa 2Hopliini (2) Afr.12
Omocrates variabilis Burmeister,1844   South Africa 1Hopliini (1) Afr.12
Oniticellus cinctus (Fabricius,1775)   Thailand 13Scarabaeinae Or.1 (13)
Oniticellus formosus Chevrolat,1830   Gambia 2Scarabaeinae Afr.1 (2)
Kenya 1Scarabaeinae Afr.1 (1)
Oniticellus fulvus (Goeze ,1774)   Afghanistan 9Scarabaeinae Pal.5 (9)
Austria 2Scarabaeinae Pal.24 (2)
Iran 1Scarabaeinae Pal.34 (1)
Italy 7Scarabaeinae Pal.23 (7)
Spain 7Scarabaeinae Pal.27 (7)
Spain 4Mallorca
Scarabaeinae Pal.38 (4)
Oniticellus pallens (Olivier,1789)   Afghanistan 5Scarabaeinae Pal.5 (5)
Oniticellus pallipes (Fabricius,1781)   Afghanistan 18Scarabaeinae Pal.5 (18)
Spain 8Scarabaeinae (8) Pal.27
Oniticellus pictus (Hausman,1807)   South Africa 1Scarabaeinae (1) Afr.30
Oniticellus planatus Castelnau,1840   Cameroon 2Scarabaeinae Afr.1 (2)
Ethiopia 1Scarabaeinae (1) Afr.22
Kenya 6Scarabaeinae Afr.1 (6)
South Africa 8Scarabaeinae (8) Afr.1+30
Oniticellus tesselatus (Harold,1879)   Malaysia 3Scarabaeinae Or.1 (3)
Thailand 2Scarabaeinae Or.1 (2)
Onitis alexis Klug,1835   Ethiopia 2Scarabaeinae Afr.5 (2)
Gambia 176Scarabaeinae (176) Afr.13+14+15+16
Kenya 4Scarabaeinae (4) Afr.16
Namibia 18Scarabaeinae (18) Afr.17+38
Senegal 7Scarabaeinae (7) Afr.16
South Africa 14Scarabaeinae (14) Afr.17+38
Tanzania 4Scarabaeinae (4) Afr.17
Onitis alexis septentrionalis Balthasar,1942   Greece 14Scarabaeinae Pal.14 (14)
Tunisia 4Scarabaeinae Pal.14 (4)
Turkey 3Scarabaeinae Pal.14 (2)
Pal.47 (1)
Onitis belial (Fabricius,1798)   Spain 1Scarabaeinae (1) Pal.14
Tunisia 3Scarabaeinae (3) Pal.14+26
Onitis caffer Boheman,1857   Lesotho 2Scarabaeinae Afr.39 (2)
South Africa 4Scarabaeinae Afr.39 (3)
Type Coleoptera (1) Syntype 06357:x of Onitis caffer Boheman 1857
  Onitis caffer Boheman 1857

Onitis cf. niger ssp.? Lansberge,1875   Indonesia 1Java
Scarabaeinae Or.1 (1)
Onitis cf. subopacus ssp.? Arrow,1931   Thailand 1Scarabaeinae Or.1 (1)
Onitis crenatus Reiche,1850   South Africa 7Scarabaeinae (7) Afr.39
Onitis cupreus Laporte,1840   Guinea-Bissau 2Scarabaeinae Afr.5 (2)
Sierra Leone 3Scarabaeinae Afr.5 (3)
Onitis curvipes lansberge,1875   South Africa 1Scarabaeinae (1) Afr.39
Onitis damoetas Steven,1806   Armenia 2Scarabaeinae Pal.46 (2)
Greece 1Scarabaeinae Pal.50 (1)
Onitis ezechias Reiche & Saulcy,1856   Israel 2Scarabaeinae Pal.35 (1)
Pal.50 (1)
Onitis falcatus (Wulfen,1796)   Philippines 1Scarabaeinae Or.4 (1)
Onitis humerosus (Pallas,1771)   Armenia 1Scarabaeinae Pal.46 (1)
Greece 7Scarabaeinae Pal.14 (3)
Pal.51 (4)
Onitis inversidens Lansberge,1875   Tanzania 2Scarabaeinae (2) Afr.17
Onitis ion (Olivier,1789)   Italy 1Scarabaeinae (1) Pal.23
Spain 1Scarabaeinae (1) Pal.14
Tunisia 2Scarabaeinae (2) Pal.26
Onitis minutus Lansberge,1875   South Africa 3Scarabaeinae (3) Afr.39
Onitis niger Lansberge,1875   Thailand 2Scarabaeinae Pal.51 (2)
Onitis perbrincki Ferreira,1958   South Africa 1Type Coleoptera 2769 (1) Synonymized with O. pecuarius Lansb. by Ferreira 1976 Holotype 02770:1 of Onitis perbrincki Ferreira 1958
  Onitis perbrincki Ferreira 1958

Onitis phartopus Lansberge,1875   Indonesia 4Java
Scarabaeinae Or.1 (4)
Onitis philemon Fabricius,1801   Nepal 1Scarabaeinae Or.1 (1)
Onitis picticollis Boheman,1857   Kenya 2Scarabaeinae (2) Afr.12
South Africa 1Scarabaeinae (1) Afr.39
Onitis politus Lansberge,1875   Ethiopia 11Scarabaeinae Afr.5 (11)
Onitis reichei Lansberge,1875   Gambia 2Scarabaeinae (2) Afr.17
Senegal 3Scarabaeinae (3) Afr.17
Onitis robustus Boheman,1857   South Africa 2Type Coleoptera 2718 (2) Paralectotype 02771:2-3 of Onitis robustus Boheman 1857
  Onitis robustus Boheman 1857

Onitis spinipes (Drury,1770)   Thailand 12Scarabaeinae Or.1 (12)
Onitis spp.    Ethiopia 10Scarabaeinae Afr.5 (10)
Guinea-Bissau 1Scarabaeinae (1) Afr.18
Kenya 9Scarabaeinae (9) Afr.17+18
Namibia 1Scarabaeinae (1) Afr.18
South Africa 1Scarabaeinae (1) Afr.18
Tanzania 2Scarabaeinae (2) Afr.17
Onitis subopacus Arrow,1931   Thailand 2Scarabaeinae Pal.51 (2)
Onitis tortuosus Houston,1983   South Africa 16Scarabaeinae (16) Afr.17
Onitis uncinatus Klug,1855   Mozambique 4Scarabaeinae (4) Afr.39
South Africa 2Scarabaeinae (2) Afr.39
Onitis viridulus Boheman,1857   Ethiopia 2Scarabaeinae Afr.5 (2)
Mozambique 4Scarabaeinae (4) Afr.39
South Africa 5Scarabaeinae (5) Afr.39
Ontherus appendiculatus (Mannerheim,1829)   Brazil 2Scarabaeinae (2) S.Am.5
Paraguay 1Scarabaeinae S.Am.3 (1)
Ontherus azteca Harold,1869   Brazil 2Scarabaeinae (2) S.Am.5
Ontherus brevicollis Kirsch,1871   Brazil 2Scarabaeinae S.Am.3 (2)
Colombia 1Scarabaeinae S.Am.3 (1)
Ontherus diabolicus Genier,1996   Ecuador 2Scarabaeinae (2) S.Am.5
Ontherus lunicollis Genier,1996   Colombia 1Scarabaeinae (1) 1 paratype S.Am.3
Ontherus obliquus Genier,1996   Peru 1Scarabaeinae (1) 1 paratype S.Am.3
Ontherus pubens Genier,1996   Ecuador 4Scarabaeinae (4 ) S.Am.3+5 (incl. 2 paratypes)

Ontherus spp.    Guatemala 4Scarabaeinae S.Am.6 (4)
Ontherus sulcator (Fabricius,1775)   Brazil 9Scarabaeinae S.Am.3 (9)
Ontherus trituberculatus (Balthasar,1938)   Ecuador 2Scarabaeinae S.Am.5 (2)
Onthophagus acutus d’Orbigny,1908   Namibia 9Scarabaeinae Afr.40 (9)
South Africa 104Scarabaeinae (104) Afr.39+40
Onthophagus aeruginosus Roth,1851   South Africa 2Scarabaeinae Afr.40 (2)
Onthophagus afghanus Petrovitz,1961   Afghanistan 7Type Petrovitz (7) (= O.finschi Harold
1877?) Holotype Paratype 00925:1-7 of Onthophagus afghanus Petrovitz 1961
  Onthophagus afghanus Petrovitz 1961

Onthophagus alcyon Klug,1855   South Africa 2Type Coleoptera 2769 (2) Holotype Paratype 02772:1-2 of Onticellus splendens Wallengren 1881
  Onticellus splendens Wallengren 1881

Zimbabwe 1Scarabaeinae Afr.40 (1)
Onthophagus anatolicus Petrovitz,1962   Turkey 4Scarabaeinae (4) 4 paratypes Pal.26
Onthophagus andalusicus Waltl,1835   Tunisia 2Scarabaeinae Pal.49 (2)
Onthophagus angorensis Petrovitz,1963   Armenia 1Scarabaeinae Pal.45 (1)
Onthophagus antillarum Arow,1903   Guadeloupe 1Scarabaeinae S.Am.6 (1)
Onthophagus ardea Klug,1855   South Africa 1Scarabaeinae Afr.44 (1)
Onthophagus ater Waterhouse,1875   Japan 2Scarabaeinae (2) Pal.34
Onthophagus atripennis Waterhouse,1875   Japan 2Scarabaeinae (2) Pal.34
Onthophagus baraudi Nicolas,1964   France 14Scarabaeinae Pal.10 (14)
Onthophagus barbieri Paulian,1978   Viet Nam 1Scarabaeinae Or.5 (1)
Onthophagus basipustulatus Heyden,1889   Afghanistan 20Scarabaeinae Pal.7 (20)
Onthophagus bicallosus Klug,1855   Mozambique 2Scarabaeinae Afr.40 (2)
Onthophagus bidens (Olivier,1789)   Guinea-Bissau 11Scarabaeinae Afr.3 (11)
Sierra Leone 1Scarabaeinae Afr.3 (1)
Onthophagus binodis Thunberg,1818   Lesotho 17Scarabaeinae Afr.41 (17)
South Africa 10Scarabaeinae Afr.41 (10)
Onthophagus bituberculatus (Olivier,1789)   Benin 1Scarabaeinae Afr.45 (1)
Onthophagus bivertex Heyden,1887   Russian Federation 1Scarabaeinae (1) Pal.34
Onthophagus blanchardi Harold,1869 (?)   Ethiopia 4Scarabaeinae Afr.3 (4)
Onthophagus brucei Reiche,1849   Lesotho 1Scarabaeinae Afr.41 (1)
South Africa 8Scarabaeinae (8) Afr.9+41
Onthophagus bubalus Harold,1867   South Africa 15Scarabaeinae Afr.41 (15)
Onthophagus cameloides d’Orbigny,1900   South Africa 1Scarabaeinae Afr.41 (1)
Onthophagus carpanetoi Pittino,1982   Turkey 15Scarabaeinae Pal.25 (6)
Pal.48 (7)
Pal.53 (2)
Onthophagus cervenkai ?   China 1Scarabaeinae (1) Pal.34
Onthophagus cf. ovatus (Linnaeus,1767)   Armenia 4Scarabaeinae Pal.45 (4)
Turkey 1Scarabaeinae Pal.45 (1)
Onthophagus coenobita (Herbst,1783)   Austria 2Scarabaeinae (2) Pal.33
Bulgaria 3Scarabaeinae (3) Pal.25
Denmark 2Scarabaeinae Main (2)
France 1Scarabaeinae Pal.9 (1)
Germany 4Scarabaeinae (4) Pal.33+324
Romania 5Scarabaeinae Pal.9 (5)
Sweden 9 Skåne
Scarabaeinae Main (9)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 6Scarabaeinae (6) Pal.9+25
Unknown 1 Alan Dufberg 47 (1)
Onthophagus conspersus Reitter,1892   Afghanistan 4Scarabaeinae Pal.7 (4)
Onthophagus cornifrons J.Thomson,1858   Sierra Leone 9Scarabaeinae Afr.3 (9)
Onthophagus crassicollis Boucomont,1913   Viet Nam 2Scarabaeinae Or.3 (1)
Or.5 (1)
Onthophagus cuneus Boucomont,1924   Philippines 1Scarabaeinae Or.3 (1)
Onthophagus cupreus Harold,1880   Guinea-Bissau 8Scarabaeinae Afr.3 (8)
Onthophagus dama (Fabricius,1798)   Nepal 13Scarabaeinae Or.3 (1)
Pal.49 (12)
Onthophagus dapcauensis Boucomont,1921   Viet Nam 1Scarabaeinae Or.3 (1)
Onthophagus dedecor Wallengren,1881   South Africa 1Type Coleoptera 2769 (1) Typus Syntype 02773:1 of Onthophagus dedecor Wallengren 1881
  Onthophagus dedecor Wallengren 1881

Onthophagus dellacasai Pittino & Mariani,1981   Greece 1Scarabaeinae (1) Pal.25
Onthophagus depressus Harold,1871   South Africa 3Scarabaeinae Afr.41 (3)
Onthophagus dorsosignatus Orbigny,1898   Turkey 1Scarabaeinae Pal.53 (1)
Onthophagus dragei Harold,   Namibia 3Scarabaeinae Afr.44 (3)
Onthophagus falsus Gillet,1925   India 2Scarabaeinae Or.4 (2)
Onthophagus ferrugineus Fåhraeus,   Namibia 6Scarabaeinae Afr.41 (6)
Onthophagus fimetarius Roth,1851   South Africa 1Scarabaeinae Afr.41 (1)
Onthophagus fissicornis Steven,1809   Armenia 2Scarabaeinae Pal.45 (2)
Greece 7Scarabaeinae Pal.9 (7)
Romania 4Scarabaeinae Pal.9 (4)
Turkey 3Scarabaeinae Pal.8 (1)
Pal.48 (2)
Onthophagus flavocinctus Klug,1855   South Africa 7Scarabaeinae Afr.44 (7)
Onthophagus fodiens Waterhouse,1875   Japan 2Scarabaeinae (2) Pal.35
Onthophagus fracticornis (Preyssler,1790)   Algeria 1Scarabaeinae Pal.24 (1)
Armenia 9Scarabaeinae Pal.45 (9)
Bosnia and Herzegovina 1Bosnia
Scarabaeinae Pal.34 (1)
Denmark 9Scarabaeinae Main (9)
France 3Scarabaeinae Pal.10 (3)
Italy 12Scarabaeinae (12) Pal.9+24
Romania 3Scarabaeinae Pal.10 (3)
Spain 2Scarabaeinae Pal.9 (1)
Pal.24 (1)
Sweden 240  Skåne Halland Småland Öland Gotland Östergötland Västergötland Bohuslän Uppland
Alan Dufberg 47 (12)
Scarabaeinae Main (228)
Onthophagus furcatus (Fabricius,1781)   Bulgaria 57Scarabaeinae (54) Pal.10+25+30
Pal.48 (3)
France 1Scarabaeinae Pal.10 (1)
Greece 56Scarabaeinae Pal.10 (55)
Pal.52 (1)
Greece 2Crete
Scarabaeinae Pal.48 (2)
Spain 28Scarabaeinae (28) Pal.10+24
Turkey 29Scarabaeinae Pal.10 (5)
Pal.48 (18)
Pal.53 (6)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 8Scarabaeinae Pal.10 (8)
Onthophagus gazella (Fabricius,1787)   Namibia 11Scarabaeinae Afr.42 (11)
South Africa 27Scarabaeinae Afr.42 (27)
Tanzania 1Scarabaeinae Afr.57 (1)
Onthophagus gibberamus d’Orbigny,1902   Namibia 2Scarabaeinae Afr.44 (2)
Onthophagus gibbulus (Pallas,1781)   Czech Republic 2Scarabaeinae (2) Pal.34
Finland 1Reference Coleoptera (1)
Iraq 1Scarabaeinae (1) Pal.34
Russian Federation 15Scarabaeinae (15) Pal.11+34
Sweden 3 Skåne
Scarabaeinae Main (3)
Onthophagus glasunowi Koshantschikov,1895 (?)   Uzbekistan 1Scarabaeinae (1) Pal.35
Onthophagus graphicus Wallengren,1881   South Africa 1Type Coleoptera 2769 (1) Typus Holotype Syntype 02774:1 of Onthophagus graphicus Wallengren 1881
  Onthophagus graphicus Wallengren 1881

Onthophagus grossepunctatus Reitter,1905   Bulgaria 9Scarabaeinae (9) Pal.10+25
Greece 6Scarabaeinae Pal.52 (6)
Italy 3Scarabaeinae Pal.10 (3)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 3Scarabaeinae Pal.10 (3)
Onthophagus haroldi Ballion,1871 (?)   Afghanistan 2Scarabaeinae Pal.8 (2)
Onthophagus hecate (Panzer,1794)   Canada 2Scarabaeinae N.Am.1 (2)
Onthophagus hirtus (Illiger,1803)   Morocco 3Scarabaeinae (3) Pal.12
Tunisia 14Scarabaeinae (14) Pal.12
Onthophagus hoepfneri Harold,1869   Mexico 1Scarabaeinae N.Am.1 (1)
United States 1Arizona
Scarabaeinae N.Am.1 (1)
Onthophagus hyaena (Fabricius,1801)   Lesotho 4Scarabaeinae Afr.42 (4)
South Africa 8Scarabaeinae Afr.42 (8)
Onthophagus illyricus (Scopoli,1763)   Bulgaria 9Scarabaeinae (9) Pal.11+25+30
Romania 1Scarabaeinae Pal.11 (1)
Sweden 39   Gotland
Alan Dufberg 47 (6)
Scarabaeinae Main (33)
Turkey 2Scarabaeinae Pal.11 (2)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Scarabaeinae Pal.11 (1)
Unknown 1Scarabaeinae Pal.49 (1)
Onthophagus interstitialis Fåhraeus,1857   South Africa 14Scarabaeinae Afr.42 (14)
Onthophagus joannae Goljan,1953   Italy 15Scarabaeinae Pal.11 (15)
Norway 2Scarabaeinae (2)
Spain 2Scarabaeinae Pal.11 (2)
Sweden 119  Skåne Blekinge Gotland
Alan Dufberg 47 (9)
Scarabaeinae Main (110)
Onthophagus jugicola d ’Orbigny,1902 (?)   Ethiopia 4Scarabaeinae Afr.3 (4)
Onthophagus kindermanni Harold,1877   Cyprus 4Scarabaeinae Pal.52 (4)
Onthophagus knulli Howden & Cartwright,1963   United States 1Arizona
Scarabaeinae N.Am.1 (1)
Onthophagus kozlovi Kabakov,1990   China 1Scarabaeinae (1) Pal.35
Onthophagus lanista Castelnau,1840   South Africa 6Scarabaeinae Afr.41 (6)
Onthophagus lemur (Fabricius,1781)   Croatia 1Scarabaeinae Pal.34 (1)
France 11Scarabaeinae (11) Pal.10+24
Italy 7Scarabaeinae Pal.10 (7)
Romania 2Scarabaeinae Pal.10 (2)
Turkey 5Scarabaeinae Pal.48 (5)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 3Scarabaeinae Pal.10 (3)
Onthophagus lenzii Harold,1874   Japan 2Scarabaeinae (2) Pal.35
Onthophagus leucostigma (Stevens,1811)   Afghanistan 4Scarabaeinae (4) Pal.7 (as ssp. leucomelas Solsky)

Turkey 6Scarabaeinae (6) Pal.25
Onthophagus loceratus ?   Tanzania 1Scarabaeinae Afr.45 (1)
Onthophagus lucidus (Illiger,1800)   Armenia 1Scarabaeinae Pal.45 (1)
Greece 7Scarabaeinae Pal.10 (6)
Pal.51 (1)
Italy 1Scarabaeinae (1) Pal.34
Turkey 3Scarabaeinae Pal.48 (1)
Pal.53 (2)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Scarabaeinae Pal.10 (1)
Onthophagus lugubris Fåhraeus,1857   South Africa 10Scarabaeinae Afr.42 (10)
Onthophagus luridipennis Boheman,1858   China 1 Scarabaeinae Pal.34 (1)
Viet Nam 1Scarabaeinae Or.5 (1)
Onthophagus maki (Illiger,1803)   Tunisia 5Scarabaeinae Pal.25 (1)
Pal.52 (4)
Onthophagus marginalis (Gebler,1817)   Afghanistan 1Scarabaeinae Pal.8 (1)
Turkey 2Scarabaeinae (2) Pal.12
Uzbekistan 1Scarabaeinae (1) Pal.12
Onthophagus marginalis nigrimargo Goidanich,1926   Afghanistan 1Scarabaeinae Pal.8 (1)
Onthophagus massai Baraud,1975   Italy 1Scarabaeinae (1) Pal.35
Onthophagus medius (Kugelann,1792)   Bulgaria 1Scarabaeinae (1) Pal.36
France 5Scarabaeinae Pal.36 (5)
Germany 2Scarabaeinae Pal.36 (2)
Italy 7Sicilia
Scarabaeinae Pal.36 (7)
Sweden 7 Skåne
Scarabaeinae Main (7)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 5Scarabaeinae (5) Pal.36
Onthophagus melanocephalus (Klug,1845)   Tunisia 1Scarabaeinae (1) Pal.16
Onthophagus melitaeus (Fabricius,1798)   Spain 1Scarabaeinae (1) Pal.35
Onthophagus monodon Fåhraeus,1857   South Africa 2Scarabaeinae Afr.43 (2)
Onthophagus nebulosus Reiche,1864   Tunisia 1Scarabaeinae (1) Pal.35
Onthophagus neomirabilis Howden,1973   Honduras 16Scarabaeinae (16) S.Am.1+2
Onthophagus nigellus (Illiger,1803)   Tunisia 5Scarabaeinae (5) Pal.25
Onthophagus nuchicornis (Linnaeus,1758)   Bulgaria 3Scarabaeinae Pal.10 (3)
Denmark 4Scarabaeinae Main (4)
Finland 2Scarabaeinae (2)
Germany 1Scarabaeinae (1) Pal.34
Hungary 1Scarabaeinae (1) Pal.24
Italy 3Scarabaeinae Pal.23 (3)
Sweden 425  Skåne Blekinge Halland Småland Öland Gotland Östergötland Västergötland Bohuslän
Alan Dufberg 47 (17)
Scarabaeinae Main (408)
Tunisia 1Scarabaeinae Pal.52 (1)
Onthophagus nutans (Fabricius,1787)   Italy 5Scarabaeinae Pal.10 (5)
Romania 7Scarabaeinae Pal.10 (7)
Onthophagus obtusicornis Fåhraeus,1857   Namibia 1Scarabaeinae Afr.43 (1)
South Africa 13Scarabaeinae Afr.43 (13)
Onthophagus obtusus Peringuey,1901   South Africa 1Scarabaeinae Afr.42 (1)
Onthophagus ohbayashii Nomura,1939   Japan 1Scarabaeinae (1) Pal.35
Onthophagus opacicollis Reitter,1892   Algeria 1Scarabaeinae Pal.24 (1)
Bulgaria 4Scarabaeinae (4) Pal.10+25
Cyprus 3Scarabaeinae Pal.52 (3)
Greece 22Scarabaeinae Pal.10 (12)
Pal.52 (10)
Greece 1Crete
Scarabaeinae Pal.48 (1)
Italy 6Scarabaeinae Pal.10 (6)
Morocco 1Scarabaeinae (1) Pal.24
Spain 10Scarabaeinae (10) Pal.10+24
Turkey 19Scarabaeinae Pal.10 (13)
Pal.48 (4)
Pal.53 (2)
Onthophagus orpheus  (Panzer,1794)   United States 1Pennsylvania
Scarabaeinae N.Am.1 (1)
Onthophagus oshimanus Nakane,1960   Japan 1Scarabaeinae (1) Pal.35
Onthophagus ovatus (Linnaeus,1767)   Cyprus 4Scarabaeinae Pal.52 (4)
Germany 2Scarabaeinae Pal.34 (2)
Greece 3Scarabaeinae Pal.10 (1)
Pal.52 (2)
Italy 12Scarabaeinae Pal.23 (12)
Spain 6Scarabaeinae (6) Pal.10+24
Sweden 146  Skåne Blekinge Öland Gotland
Alan Dufberg 47 (1)
Scarabaeinae Main (145)
Turkey 4Scarabaeinae Pal.52 (4)
United Kingdom 1England
Scarabaeinae Pal.10 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Scarabaeinae Pal.10 (2)
Unknown 1Scarabaeinae Pal.35 (1)
Onthophagus panici Petrovitz,1964   Greece 2Scarabaeinae (2) Pal.25
Onthophagus papulatus Boucomont,1914   Viet Nam 2Scarabaeinae Or. (1)
Or.5 (1)
Onthophagus persianus Olsoufieff,1900   Armenia 1Scarabaeinae Pal.45 (1)
Onthophagus petrovitzi Pittino,2004   Turkey 4Scarabaeinae (4) Pal.25 (as O.vitulus ssp. citellorum Petrovitz)

Onthophagus pilosus Fåhraeus,1857   South Africa 1Scarabaeinae Afr.43 (1)
Onthophagus potanini Kabakov,1979   China 1Scarabaeinae (1) Pal.35
Onthophagus punctatus (Illiger,1803)   Spain 1Scarabaeinae (1) Pal.26
Onthophagus punneeae Masumoto,1989   Viet Nam 1Scarabaeinae Or.5 (1)
Onthophagus purpurascens Boucomont,1914   Viet Nam 2Scarabaeinae Or.4 (1)
Or.5 (1)
Onthophagus pygargus Motchulsky,1845   Afghanistan 28Scarabaeinae Pal.7 (28)
Kazakhstan 1Scarabaeinae (1) Pal.35
Onthophagus pyramidalis Klug,1855   South Africa 1Scarabaeinae Afr.42 (1)
Onthophagus ramosellus Bates,1891 (?)   Nepal 1Scarabaeinae Or.3 (1)
Onthophagus ruficapillus (Brulle,1832)   Bulgaria 14Scarabaeinae (14) Pal.12+25+30
Cyprus 19Scarabaeinae Pal.12 (18)
Pal.52 (1)
Greece 16Scarabaeinae Pal.12 (16)
Greece 1Crete
Scarabaeinae Pal.48 (1)
Italy 11Scarabaeinae (11) Pal.12+23
Romania 9Scarabaeinae Pal.12 (9)
Turkey 10Scarabaeinae Pal.12 (5)
Pal.48 (5)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 5Scarabaeinae Pal.12 (5)
Onthophagus sacharovskii Olsoufieff,1918   Turkey 5Scarabaeinae (5) Pal.25
Onthophagus saphirinus Fåhraeus,1857   Namibia 4Scarabaeinae Afr.43 (4)
South Africa 1Scarabaeinae Afr.43 (1)
Zimbabwe 1Scarabaeinae Afr.43 (1)
Onthophagus schaefernai Balthasar,1935   China 1Scarabaeinae Pal.35 (1)
Onthophagus schatzmayri Pierotti,1959   Italy 2Scarabaeinae (2) Pal.24
Onthophagus sericatus Reitter,1892   Greece 9Scarabaeinae (9) Pal.25
Onthophagus setosus Fåhraeus,1857   Lesotho 1Scarabaeinae Afr.43 (1)
South Africa 1Scarabaeinae Afr.43 (1)
Onthophagus sibuyanus ?   Philippines 1Scarabaeinae Or.4 (1)
Onthophagus similis (Scriba,1790)   Armenia 1Scarabaeinae Pal.45 (1)
Germany 1Scarabaeinae (1) Pal.30
Ireland 2Scarabaeinae (2) Pal.34
Italy 2Scarabaeinae (2) Pal.12+23
Spain 8Scarabaeinae (8) Pal.12
Sweden 181  Skåne Blekinge Halland Öland Gotland
Alan Dufberg 47 (16)
Scarabaeinae Main (165)
Onthophagus speculifer Solsky,1876   Afghanistan 5Scarabaeinae Pal.7 (5)
Onthophagus spp.    Afghanistan 1Scarabaeinae Pal.53 (1)
Botswana 1Scarabaeinae (1) Afr.23
Cameroon 1Scarabaeinae (1) Afr.18
China 1Scarabaeinae Pal.53 (1)
Ecuador 3Scarabaeinae S.Am.2 (3)
Ethiopia 14Scarabaeinae (14) Afr.4+18+23
Gambia 8Scarabaeinae (8) Afr.18
Gambia 1Scarabaeinae (1) Afr.22
Greece 1Scarabaeinae Pal.53 (1)
Guatemala 2Scarabaeinae S.Am.1 (2)
Guinea-Bissau 20Scarabaeinae (20) Afr.4+18
Honduras 4Scarabaeinae (4) S.Am.1+5
Indonesia 5Java
Scarabaeinae Or.2 (5)
Iran 1Scarabaeinae Pal.53 (1)
Kenya 11Scarabaeinae (11) Afr.18
Malaysia 1Scarabaeinae Or.2 (1)
Mexico 4Scarabaeinae N.Am.1 (4)
Namibia 1Scarabaeinae (1) Afr.18
Sierra Leone 9Scarabaeinae Afr.5 (9)
South Africa 11Scarabaeinae (11) Afr.4+18+44
Spain 5Mallorca
Scarabaeinae Pal.53 (5)
Sri Lanka 1Scarabaeinae Or. (1)
Sweden 1 Skåne
Scarabaeinae Main (1) Marked as misslabelled (Vomb 1972)
Sweden 4Scarabaeinae Main (4)
Tanzania 2Scarabaeinae (2) Afr.18
Tanzania 3Scarabaeinae (3) Afr.22
Onthophagus suermelii Petrovitz,1963   Turkey 1Scarabaeinae Pal.49 (1)
Onthophagus suffusus Klug,1855   South Africa 4Scarabaeinae Afr.43 (4)
Zimbabwe 2Scarabaeinae Afr.43 (2)
Onthophagus sugillatus Klug,1855   Lesotho 21Scarabaeinae Afr.43 (21)
South Africa 16Scarabaeinae Afr.43 (16)
Onthophagus suturellus Brulle,1832   Armenia 1Scarabaeinae Pal.45 (1)
Greece 5Scarabaeinae (5) Pal.12+25+26
Morocco 2Scarabaeinae Pal.10 (2)
Tunisia 6Scarabaeinae Pal.10 (6)
Onthophagus talpa Boheman,1857   Namibia 2Scarabaeinae Afr.43 (2)
South Africa 2Scarabaeinae Afr.43 (2)
Onthophagus taurus (Schreber,1759)   Afghanistan 30Scarabaeinae (30) Pal.6+7
Algeria 3Scarabaeinae Pal.11 (3)
Austria 1Scarabaeinae Pal.34 (1)
Bulgaria 8Scarabaeinae Pal.11 (7)
Pal.53 (1)
Germany 9Scarabaeinae (9) Pal.34
Greece 26Scarabaeinae Pal.11 (24)
Pal.49 (2)
Greece 2Crete
Scarabaeinae Pal.43 (2)
Hungary 2Scarabaeinae Pal.35 (2)
Italy 12 Alan Dufberg 47 (1)
Scarabaeinae (11) Pal.11+30
Morocco 1Scarabaeinae Pal.11 (1)
Portugal 1Scarabaeinae (1) Pal.23
Portugal 83 Azores
Alan Dufberg 47 (2)
Scarabaeinae (37) Pal.23+30
Scarabaeinae Pal.48 (29)
Scarabaeinae Pal.49 (14)
Scarabaeinae Pal.53 (1)
Spain 53Scarabaeinae (52) Pal.11+23
Pal.53 (1)
Spain 15Mallorca
Scarabaeinae Pal.38 (14)
Pal.53 (1)
Tunisia 9Scarabaeinae (7) Pal.11+25
Pal.49 (2)
Turkey 9Scarabaeinae Pal.11 (9)
Unknown 4Scarabaeinae Pal.49 (4)
Onthophagus tesquorum Semenov & Medvedv,1927   Greece 1Scarabaeinae Pal.11 (1)
Greece 3Crete
Scarabaeinae Pal.48 (3) as O. andalusicus ssp. t.
Onthophagus tragus (Fabricius,1792)   China 1Scarabaeinae (1) Pal.35
Onthophagus transcaspicus König, 1889   Afghanistan 18Scarabaeinae Pal.8 (18)
Onthophagus tricuspis Semenov,1899   Turkey 1Scarabaeinae (1) Pal.25
Onthophagus trigibber Reitter,1892   Tunisia 12Scarabaeinae (12) Pal.25
Onthophagus trituber (Wiedemann,1823)   Japan 1Scarabaeinae (1) Pal.35
Onthophagus truchmenus Kolenati,1846   Afghanistan 1Scarabaeinae (1) Pal.7 (as ssp. araxicola Reitter)

Armenia 8Scarabaeinae Pal.45 (8)
Iran 1Scarabaeinae Pal.45 (1)
Onthophagus turpidus Reitter,1887   Afghanistan 9Scarabaeinae Pal.7 (9)
Onthophagus uniformis Heyden,1886   Russian Federation 1Scarabaeinae (1) Pal.35
Onthophagus vacca (Linnaeus,1767)   Armenia 3Scarabaeinae Pal.45 (3)
Denmark 1Scarabaeinae Main (1)
Germany 5Scarabaeinae (5) Pal.36
Greece 36Scarabaeinae Pal.36 (32)
Pal.52 (4)
Greece 2Rhodos
Scarabaeinae Pal.43 (2)
Greece 5Scarabaeinae Pal.36 (5)
Hungary 1Scarabaeinae Pal.36 (1)
Italy 2Scarabaeinae Pal.36 (2)
Italy 35Sardinia
Scarabaeinae Pal.36 (35)
Morocco 1Scarabaeinae Pal.36 (1)
Spain 2Scarabaeinae Pal.36 (2)
Sweden 3 Skåne
Scarabaeinae Main (3)
Turkey 4Scarabaeinae (4) Pal.36
Turkey 2Scarabaeinae Pal.36 (2)
Onthophagus vanellus Lansberge,1883   Namibia 1Scarabaeinae Afr.44 (1)
Onthophagus variegatus (Fabricius,1798)   Namibia 1Scarabaeinae Afr.44 (1)
Onthophagus venustulus Erichson,   Namibia 2Scarabaeinae Afr.44 (2)
Onthophagus verticalis Fåhraeus,1857   South Africa 4Scarabaeinae Afr.44 (4)
Onthophagus verticicornis (Laicharting,1781)   Armenia 4Scarabaeinae Pal.45 (4)
Germany 2Scarabaeinae (2) Pal.34
Italy 17Scarabaeinae (17) Pal.23
Romania 1Scarabaeinae (1) Pal.12
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 10Scarabaeinae (10) Pal.12
Onthophagus vinctus Erichson,1843   Ethiopia 1Scarabaeinae Afr.4 (1)
Lesotho 1Scarabaeinae Afr.43 (1)
Namibia 6Scarabaeinae Afr.43 (6)
Sierra Leone 94Scarabaeinae Afr.4 (94)
South Africa 11Scarabaeinae Afr.43 (11)
Onthophagus vitulus (Fabricius,1777)   Germany 1Scarabaeinae (1) Pal.34
Onthophagus wittei Harold,1867   Namibia 1Scarabaeinae Afr.44 (1)
Onthophagus yakuinsulanus Nakane,1984   Japan 2Scarabaeinae (2) Pal.35
Onthophagus zapotecus Zunino & Haffner,1988   Mexico 13Scarabaeinae N.Am.1 (13)
Oogenius castilloi Martinez & Pena,1990   Chile 1Rutelinae (1) S.Am 2
Oplostomus fuligineus (Olivier,1789)   Namibia 5Cetoniinae (5) Afr.5+29
South Africa 3Cetoniinae (3) Afr.17+29
Zimbabwe 1Cetoniinae Afr.29 (1)
Oplostomus rotundiceps Mos.,1905   Malawi 1Cetoniinae (1) Afr.5 (as Goniochilus r)

Oplostomus rufiventris Harold,1878   Kenya 1Cetoniinae (1) Afr.5 (as Goniochilus r)


Orizabus clunalis (LeConte,1856)   United States 2Arizona
Dynastinae N.Am.5 (2)
Orizabus isodonoides Fairmaire,1878   Mexico 2Dynastinae N.Am.2 (2)
Unknown 1Dynastinae N.Am.2 (1)
Orizabus pyriformis (LeConte,1848)   United States 1New Mexico
Dynastinae N.Am.5 (1)
Orodaliscus zangi Schmidt,1906   Russian Federation 1Aphodiinae Pal.13 (1) Paratype of Aphodius jureceki Balthasar 1932
Orphnus bicolor Fabricius,1801   Unknown 2Orphninae Afr.1 (2)
Orphnus macleayi zambezianus Péringuey,1896   Central African Republic 4Orphninae Afr.1 (4)
Orphnus parentalis Péringuey,1908   South Africa 2Orphninae Afr.1 (2)
Orphnus picinus Westwood,1845   Unknown 2Orphninae Afr.1 (2)
Orphnus rufulus Boheman,1857   South Africa 3Orphninae Afr.1 (3)
Orsilochus cornutus (Thunberg,1818)   South Africa 1Dynastinae (1) Afr.12
Orubesa linnavuori Endrödi,1970   Sudan 1Type Coleoptera 3955 (1) Paratype 03959:2 of Dynamopus linnavuori Endrödi 1970
  Dynamopus linnavuori Endrödi 1970

Oruscatus davus (Erichson,1847)   Peru 7Scarabaeinae S.Am.1 (7)
Oryctes agamennon arabicus Fairmaire,1896   Oman 3Dynastinae Pal.6 (3)
Oryctes augias (Olivier,1789)   Madagascar 1Dynastinae Afr.4 (1)
Oryctes boas (Fabricius,1775)   Eritrea 1Dynastinae Afr.2 (1)
Gambia 18Dynastinae (18) Afr.1+2+17+18
Guinea 2Dynastinae Afr.2+3 (2)
Guinea-Bissau 1Dynastinae Afr.17 (1)
Kenya 16Dynastinae (16) Afr.2+3+4+17+18
Lesotho 2Dynastinae (2) Afr.13
Namibia 10Dynastinae (10) Afr.13+14
Senegal 2Dynastinae Afr.2 (2)
Sierra Leone 1Dynastinae (1) Afr.17
South Africa 10Dynastinae (10) Afr.2+3+13+14
Tanzania 10Dynastinae (10) Afr.2+3+4+18
Unknown 3Dynastinae (3) Afr.2+3
Oryctes chevrolati Guerin-Meneville,1844   Reunion 1Dynastinae Afr.4 (1)
Oryctes erebus Burmeister,1847   Cameroon 2Dynastinae Afr.4 (2)
Oryctes gigas Castelnau,1840   Cameroon 1Dynastinae Afr.1 (1)
Oryctes gnu Mohnike,1874   Unknown 1Dynastinae Or.1 (1)
Oryctes monocerus (Olivier,1789)   Comoros 2Dynastinae Afr.19 (2)
Gambia 5Dynastinae (5) Afr.17
Kenya 6Dynastinae (6) Afr.17
Mozambique 3Dynastinae Afr.5 (3)
Namibia 1Dynastinae Afr.5 (1)
Senegal 1Dynastinae Afr.5 (1)
Sri Lanka 1Dynastinae Or.1 (1)
Tanzania 2Dynastinae Afr.5 (2)
Unknown 2Dynastinae Afr.5 (2)
Unknown 2Dynastinae Or.1 (2)
Oryctes nasicornis (Linnaeus,1758)   Afghanistan 1Dynastinae Pal.10 (1)
Armenia 2Dynastinae Pal.12 (2)
Croatia 1Dynastinae Pal.19 (1)
Germany 3Dynastinae Pal.1 (3)
Sweden 549  Sk Bl Ha Sm Öl Go Ög Vg Bo Nä Sö Up Vr Dr Gä Hs
Alan Dufberg 48 (4)
Dynastinae Main (545)
Oryctes nasicornis corniculatus Villa & Villa,1833   Austria 2Dynastinae Pal.1 (2) as ssp. laevigatus Heer.
France 2Dynastinae Pal.1 (2) as ssp. laevigatus Heer
Italy 4Dynastinae Pal.7 (4) as ssp. laevigatus Heer
Oryctes nasicornis grypus (Illiger,1803)   Algeria 1Dynastinae Pal.9 (1)
Italy 2Sardinia
Dynastinae Pal.9 (2)
Italy 4Dynastinae Pal.5 (2)
Pal.9 (2)
Morocco 2Dynastinae Pal.1 (2)
Spain 4Dynastinae Pal.9 (1)
Pal.1+6 (3)
Tunisia 1Dynastinae Pal.1 (1)
Unknown 1Dynastinae Pal.1 (1)
Oryctes nasicornis holdhausi Minck,1914   Bulgaria 1Dynastinae Pal.9 (1)
Croatia 1Dynastinae Pal.9 (1)
Greece 9Crete
Dynastinae Pal.9 (9)
Oryctes nasicornis kuntzeni Minck,1914   Romania 1Dynastinae Pal.1 (1)
Turkey 1Dynastinae Pal.1 (1)
Oryctes nasicornis turkestanicus    Russian Federation 1Dynastinae Pal.1 (1)
Oryctes owariensis Beauvois,1805   Gabon 1Dynastinae Afr.4 (1)
Unknown 1Dynastinae Afr.4 (1)
Oryctes prolixus Wollaston,1864   Spain 1Canary Islands
Dynastinae Pal.19 (1)
Oryctes rhinoceros (Linnaeus,1758) (Rykanes)   India 1Dynastinae Or.1 (1)
Sri Lanka 1Dynastinae Or.3 (1)
Thailand 3Dynastinae Or.10 (3)
Unknown 9Dynastinae Or.1 (9)
Oryctes sjöstedti Kolbe,1905   Cameroon 2Dynastinae Afr.4 (2)
Senegal 1Dynastinae Afr.4 (1)
Oryctes spp.    France 2Corsica
Dynastinae Pal.8 (2)
Oryctoderus latitarsis Boisduval,1835   Indonesia 1Java
Dynastinae Or.3 (1)
Indonesia 7New Guinea
Dynastinae Or.3 (7)
Unknown 1Dynastinae Or.3 (1)
Oryctomorphus bimaculatus (Guerin-Meneville,1830)   Argentina 2Dynastinae S.Am.36 (2)
Chile 4Dynastinae S.Am.36 (4)
Sweden 1 Imported to Småland
Dynastinae S.Am.36 (1) Imported
Osmoderma barnabita Motschulsky,1845   Finland 2Cetoniinae Main (2)
Osmoderma eremicola (Knoch,1801)   Canada 1Cetoniinae N.Am.3 (1)
Canada 1Quebec
Cetoniinae N.Am.1 (1)
Unknown 2Cetoniinae N.Am.3 (2)
Osmoderma eremita (Scopoli,1763)   Austria 1Cetoniinae Pal.22 (1)
Czech Republic 1Cetoniinae Pal.22 (1)
Denmark 2Cetoniinae Main (2)
Germany 1Cetoniinae Pal.34 (1)
Sweden 156   Skåne Blekinge Halland Småland Öland Östergötland Västergötland Uppland
Alan Dufberg 49 (4)
Dufberg 71 (1)
Cetoniinae Main (151)
Unknown 3Cetoniinae Pal.25 (3)
Osmoderma scabrum Palisot de Beauvois,1805   Canada 2Cetoniinae N.Am.3 (2)
Canada 1Ontario
Cetoniinae N.Am.1 (1)
Canada 6Quebec
Cetoniinae N.Am.1 (6)
United States 1West Virginia
Cetoniinae N.Am.1 (1)
United States 1Pennsylvania
Cetoniinae N.Am.4 (1)
United States 1Illinois
Cetoniinae N.Am.3 (1)
Unknown 13Cetoniinae N.Am.3 (13)
Outenikwanus tomentosus Scholtz & Howden,1987   South Africa 1Scarabaeinae (1) Afr.22
Outeniqua festiva Peringuey,1885   South Africa 3Hopliini Afr.12 (3)
Oxygrylius ruginosus (LeConte,1856)   Mexico 5Dynastinae N.Am.5 (5)
United States 7Arizona
Dynastinae N.Am.6 (7)
United States 6Texas
Dynastinae N.Am.6 (6)
Oxyligyrus zoilus (Olivier,1789)   Unknown 1Dynastinae S.Am.15 (1)
Oxyomus inaequalis Lansberge,1886   Congo 1Aphodiinae (1) Afr.25
Oxyomus sylvestris (Scopoli,1763)   Austria 6Aphodiinae (6) Pal.7+11
Czech Republic 4Aphodiinae (4) Pal.15
Germany 2Aphodiinae (2) Pal.15+16
Greece 26Aphodiinae Pal.7 (26)
Poland 4Aphodiinae (4) Pal.13
Romania 1Aphodiinae Pal.7 (1)
Spain 4Aphodiinae Pal.7 (4)
Sweden 172  Skåne Småland Öland Gotland Västergötland
Alan Dufberg 48 (24)
Aphodiinae Main (148)
Switzerland 2Aphodiinae Pal.16 (2)
Turkey 16Aphodiinae Pal.26 (14)
Pal.28 (1)
Pal.30 (1)
United States 5New York
Aphodiinae N.Am.5 (5)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 4Aphodiinae Pal.7 (4)
Oxysternon conspicillatum (Weber,1801)   Ecuador 3Scarabaeinae (3) S.Am.1+5
Peru 3Scarabaeinae S.Am.1 (3)
Oxysternon durantoni Arnaud,   Brazil 2Scarabaeinae (2) S.Am.5
Oxysternon festivum viridanum d’Olsoufieff,   Brazil 2Scarabaeinae (2) S.Am.5
Oxysternon palaemon Castelnau,1840,   Brazil 2Scarabaeinae (2) S.Am.4
Oxythyrea abigail Reiche & Saulcy,1856   Greece 1Cetoniinae Pal.33 (1)
Turkey 2Cetoniinae Pal.8 (1)
Pal.30 (1)
Oxythyrea albopicta Motschulsky,1845   Armenia 11 ()
Oxythyrea albosignata Boheman,1860   Unknown 1Cetoniinae (1) Afr.27 (possible type)
Oxythyrea amethystina (Macleay,1838)   Namibia 4Cetoniinae (4) Afr.15
South Africa 17Cetoniinae (17) Afr.15
Oxythyrea cinctella (Schaum,1841)   Afghanistan 38Cetoniinae (38) Pal.9+10
Cyprus 9Cetoniinae Pal.8 (7)
Pal.34 (2)
Greece 258Cetoniinae (246) Pal.2+3+5+11+19
Pal.30 (5)
Pal.34 (7)
Turkey 69Cetoniinae (47) Pal.6+8+19
Pal.28 (17)
Pal.30 (3)
Pal.35 (2)
Unknown 2Cetoniinae Pal.19 (1)
Pal.22 (1)
Oxythyrea funesta (Poda,1761)   Bulgaria 4 Alan Dufberg 72 (2)
Cetoniinae Pal.1 (1)
Cetoniinae Pal.35 (1)
Croatia 3Cetoniinae Pal.33 (3)
Cyprus 6Cetoniinae Pal.30 (6)
France 10Cetoniinae Pal.1 (5)
Pal.35 (5)
Germany 2Cetoniinae Pal.19 (1)
Pal.28 (1)
Greece 36Cetoniinae (36) Pal.5+17
Greece 3Crete
Cetoniinae Pal.32 (3)
Hungary 2Cetoniinae Pal.33 (2)
Italy 47Cetoniinae (45) Pal.1+10+18+19+20+22
Pal.28 (1)
Pal.35 (1)
Portugal 20Cetoniinae (20) Pal.1+2
Romania 1Cetoniinae Pal.9 (1)
Russian Federation 4Cetoniinae Pal.32 (4)
Spain 8Mallorca
Cetoniinae Pal.28 (8)
Spain 14Cetoniinae (14) Pal.1+9+10+18
Sweden 2 Östergötland
Cetoniinae Main (2)
Switzerland 9Cetoniinae (5) Pal.20+22
Pal.22 (1)
Pal.28 (3)
Turkey 5Cetoniinae Pal.1 (3)
Pal.28 (2)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 29Cetoniinae Pal.1 (29)
Unknown 3Cetoniinae Pal.22 (2)
Pal.25 (1)
Oxythyrea haemorrhoidalis (Fabricius,1775)   Lesotho 1Cetoniinae (1) Afr.15
South Africa 8Cetoniinae (8) Afr.15
Oxythyrea haemorrhoidalis vitticollis (Fabricius,1775)   Lesotho 4Cetoniinae (4) Afr.16
South Africa 7Cetoniinae (7) Afr.16
Oxythyrea lucens Janson,1877   Namibia 4Cetoniinae (4) Afr.16
Oxythyrea noemi Reiche & Saulcy,1856   Armenia 5Cetoniinae Pal.30 (5)
Cyprus 6Cetoniinae Pal.28 (1)
Pal.30 (2)
Pal.33 (1)
Pal.35 (2)
Israel 4Cetoniinae Pal.30 (4)
Oxythyrea pantherina (Gory & Percheron,1833)   Iran 9Cetoniinae Pal.30 (9)
Tunisia 18Cetoniinae Pal.9 (8) as ssp. of pantherina
Pal.30 (3)
Pal.33 (7)
Turkey 2Cetoniinae Pal.30 (2)
Oxythyrea rudebecki Schein,1960   South Africa 2Type Schein (2) Paratype 00004:2-3 of Oxythyrea rudebecki Schein 1960
  Oxythyrea rudebecki Schein 1960

Oxythyrea spp.    Spain 4Mallorca
Cetoniinae Pal.27 (4)
Oxythyrea transvaalensis Moser,1913   South Africa 11Cetoniinae (11) Afr.16
Pachnoda aemula (Bourgoin,1919)   Kenya 1Cetoniinae Afr.2 (1)
Pachnoda arborescens Vigors,   Namibia 1Cetoniinae Afr.9 (1)
Pachnoda basilewskyi Ruter,1953   Gambia 3Cetoniinae Afr.3 (3)
Pachnoda bella Kraatz,1898   Namibia 1Cetoniinae Afr.9 (1)
Pachnoda ephippiata Gerstaecker,1867   Kenya 15Cetoniinae Afr.3 (15)
Tanzania 2Cetoniinae Afr.3 (1)
Afr.32 (1)
Uganda 1Cetoniinae Afr.32 (1)
Pachnoda fasciata Fabricius,1775   Unknown 1Cetoniinae Pal.24 (1)
Pachnoda flaviventris (Gory-Pecheron,1833)   South Africa 6Cetoniinae (6) Afr.9+10+21
Pachnoda impressa (Goldfuss,1805)   Cameroon 1Cetoniinae (1) Afr.24
Namibia 5Cetoniinae (5) Afr.10+11
South Africa 11Cetoniinae (11) Afr.10+11+21
Tanzania 1Cetoniinae (1) Afr.24
Unknown 3Cetoniinae (3) Afr.24
Zambia 1Cetoniinae (1) Afr.24
Zimbabwe 1Cetoniinae (1) Afr.24
Pachnoda interrupta (Olivier,1789)   Gambia 10Cetoniinae Afr.2 (10)
Senegal 3Cetoniinae (3) Afr.23
Unknown 4Cetoniinae (4) Afr.23
Pachnoda marginata (Drury,1773)   China 1Cetoniinae (1) Afr.23
Gambia 2Cetoniinae Afr.3 (1)
Afr.32 (1)
Ghana 3Cetoniinae Afr.3 (3)
Guinea 1Cetoniinae (1) Afr.23
Sierra Leone 1Cetoniinae (1) Afr.23
Togo 1Cetoniinae (1) Afr.23
Unknown 9Cetoniinae (9) Afr.23
Pachnoda marginata monacha Gory & Percheron,1833   Gambia 1Cetoniinae Afr.3 (1)
Ghana 2Cetoniinae Afr.3 (2)
Unknown 9Cetoniinae (9) Afr.23
Pachnoda marginella Fabricius,1775   Cameroon 1Cetoniinae (1) Afr.23
Pachnoda picturata Boheman,1860   Namibia 4Cetoniinae (4) Afr.3+9+23
Unknown 3Cetoniinae (3) Afr.23
Pachnoda rufa (DeGeer,1778)   Lesotho 3Cetoniinae Afr.10 (3)
Namibia 1Cetoniinae Afr.10 (1)
South Africa 2Cetoniinae Afr.10 (2)
Unknown 4Cetoniinae (4) Afr.23
Pachnoda rufa frenata (DeGeer,1778)   South Africa 7Cetoniinae Afr.10 (7)
Unknown 1Cetoniinae (1) Afr.23
Pachnoda savignyi Gory & Percheron,1833   Sudan 2Cetoniinae (2) Afr.21
Unknown 4Cetoniinae (4) Afr.21
Pachnoda sinuata Harold,1878   Unknown 4Cetoniinae (4) Afr.21
Pachnoda sinuata calceata Harold,1878   Namibia 12Cetoniinae Afr.4 (12)
Pachnoda sinuata flaviventris Harold,1878   Congo, the democratic republic of the 1Cetoniinae (1) Afr.22
South Africa 1Cetoniinae (1) Afr.22
Zambia 2Cetoniinae (2) Afr.3+22
Pachnoda spp.    Ethiopia 2Cetoniinae (2) Afr.24
Kenya 2Cetoniinae (2) Afr.22
Tanzania 3Cetoniinae (3) Afr.22
Uganda 3Cetoniinae Afr.22 (2)
Afr.32 (1)
Unknown 10Cetoniinae (10) Afr.22+23+24
Pachnoda tridentata Olivier,1789   South Africa 1Cetoniinae (1) Afr.23
Pachnoda vitticollis Moser,1914   Zimbabwe 1Cetoniinae (1) Afr.21
Pachnodella marginella (Fabricius,1775)   Kenya 1Cetoniinae (1) Afr.18
Pachycnema abdominalis Burmeister,1844 (Pachycnemula)   South Africa 2Hopliini Afr.7 (2)
Pachycnema alternans Burmeister,1844 (s.str)   South Africa 3Hopliini (3) Afr.7+19
Pachycnema calcarata Burmeister,1844 (Physocnema)   South Africa 3Hopliini Afr.6 (3)
Pachycnema calviniana Schein,1959 (Physocnema)   South Africa 1Hopliini Afr.6 (1)
Pachycnema crassipes (Fabricius,1775) (s.str)   South Africa 5Hopliini (5) Afr.6+19
Pachycnema flavolineata Burmeister,1844 (Physocnema)   South Africa 2Hopliini Afr.6 (2)
Pachycnema lineola Burmeister,1844 (Pachycnemula)   South Africa 2Hopliini Afr.7 (2)
Pachycnema marginella (Fabricius,1792) (Pachycnemula)   South Africa 2Hopliini Afr.7 (2)
Pachycnema melanospila Burmeister,1844 (s.str)   South Africa 1Hopliini Afr.6 (1)
Pachycnema moerens Peringuey,1902 (Pachycnemula)   South Africa 1Hopliini (1) 1 paratype Afr.7
Pachycnema murina Burmeister,1844 (s.str)   South Africa 2Hopliini Afr.6 (2)
Pachycnema namaqua Peringuey,1902 (Pachycnemula)   South Africa 2Hopliini (2) 2 paratypes Afr.7
Pachycnema pubibunda (Burmeister,1844) (Stenocnema)   South Africa 2Hopliini Afr.7 (2)
Pachycnema pulverulenta Burmeister,1844 (s.str)   South Africa 2Hopliini Afr.7 (2)
Pachycnema rostrata Burmeister,1844 (Pachycnemula)   South Africa 2Hopliini Afr.7 (2)
Pachycnema saga Peringuey,1902 (Pachycnemula)   South Africa 2Hopliini Afr.7 (2)
Pachycnema tibialis (Olivier,1789) (Pachycnemula)   South Africa 1Hopliini Afr.7 (1)
Pachydema anthracina Fairmaire,1860   Morocco 1Melolonthinae Pal.3 (1)
Pachydema bipartita (Brullé,1838)   Spain 2Canary Islands
Melolonthinae (2) Pal.
Spain 5Canary Islands
Melolonthinae Pal.15 (5) as Ootoma b.
Pachydema castanea (Brullé,1838)   Spain 1Canary Islands
Melolonthinae (1) Pal.
Spain 1Canary Islands
Melolonthinae (1) Pal.
Pachydema fuscipennis (Brullé,1838)   Spain 4Canary Islands
Melolonthinae Pal. (3)
Pal.14 (1) as Ootoma f.
Pachydema hirticollis (Fabricius,1787)   Tunisia 3Melolonthinae (3) Pal.3+8
Pachydema integra (Wollaston,1864)   Spain 6Canary Islands
Melolonthinae (6) Pal. (as Ootoma i.)
Spain 1Canary Islands
Melolonthinae Pal.15 (1) as Ootoma i.
Pachydema obscurella (Wollaston,1864)   Spain 3Canary Islands
Melolonthinae Pal.2 (2) as Ootoma o.
Pal.15 (1) as Ootoma o.
Spain 3Canary Islands
Melolonthinae (3) Pal. (as Ootoma o.)
Pachydema rubripennis (Lucas,1846)   Algeria 1Melolonthinae Pal.3 (1)
Pachydema xanthochroa Fairmaire,1879   Algeria 1Melolonthinae Pal.3 (1)
Tunisia 3Melolonthinae Pal.3 (3)
Pachylomera femoralis Kirby,1828   Namibia 4Scarabaeinae (4) Afr.18+26
South Africa 1Scarabaeinae (1) Afr.26
Zimbabwe 1Scarabaeinae (1) Afr.26
Pachypus caesus Erichson,1840   Italy 1Pachypodinae Pal.1 (1)
Unknown 2Pachypodinae Pal.1 (2)
Pachypus candidae (Petagna,1787)   Italy 1Pachypodinae Pal.1 (1)
Unknown 2Pachypodinae Pal.1 (2)
Pachyserica ambiversa Ahrens,   Nepal 2Sericini Or.2 (2)
Pachysoma femoralis Kirby,   Namibia 3Scarabaeinae Afr.45 (3)
Pachysoma hippocrates MacLeay,1821   South Africa 4Scarabaeinae (4) Afr.23
Pachysoma spp.    Namibia 1Scarabaeinae Afr.45 (1)
Pachytricha castanea Hope,1841   Australia 1Western Australia
Melolonthinae Pac.1 (1)
Pacuvia castanea Curtis,1845   Chile 1Melolonthinae S.Am.1 (1)
Paleira femorata (Illiger,1803)   Morocco 1Cetoniinae Pal.30 (1)
Spain 24Canary Islands
Cetoniinae Pal.18 (18)
Pal.27 (6)
Paleopragma petersi Harold,1878   Tanzania 1Cetoniinae Afr.5 (1)
Panelus rufulus Nomura,1973   Japan 3Scarabaeinae (3) Pal.25
Panelus spp.    Borneo, Indonesia / Malaysia 1Scarabaeinae Or.1 (1)
Panotrogus myschenkovi (Ballion,1871)   Afghanistan 1Melolonthinae Pal.5 (1)
Russian Federation 3Melolonthinae Pal.5 (3)
Papuana fortepunctata Heller,1913   Indonesia 1Dynastinae Or.9 (1)
Papuana japenensis Arrow,1941   Indonesia 1Dynastinae Or.9 (1)
Papuana oryctoides Dechambre,2005   Indonesia 1Dynastinae Or.9 (1)
Parabodilus wollastoni (Harold,1862)   Spain 24Canary Islands
Aphodiinae Pal.3 (13)
Pal.27 (7)
Pal.28 (3)
Pal.30 (1)
Parachrysina truquii (J.Thomson,1857)   Unknown 3Rutelinae N.Am.1 (3)
Unknown 1Rutelinae S.Am.1 (1)
Paracotalpa ursina (G.Horn,1867)   United States 3California
Rutelinae (3) N.Am 1+2
Paragymnopleurus melanarius (Harold,1867)   China 1Scarabaeinae Pal.35 (1)
Paragymnopleurus planus (Sharp,1875)   Indonesia 1Java
Scarabaeinae Or.2 (1)
Borneo, Indonesia / Malaysia 4Scarabaeinae Or.2 (4)
Paraphytus spp.    Gambia 2Scarabaeinae (2) Afr.22
Parapucaya amazonica Prell,1934   Peru 1Dynastinae S.Am.15 (1)
Pararhyssemus coluber (Mayet,1887)   Afghanistan 1Aphodiinae Pal.9 (1)
Arabia 1Aphodiinae (1) Pal.15
Paraschizognathus prasinus (Boisduval,1835)   Australia 2Rutelinae Pac.3 (2)
Paraserica grisea (Motschulsky,1866)   Japan 2Sericini Pal.2 (2)
Parastasia confluens Westwood,   Indonesia 1Celebes
Rutelinae Or.8 (1)
Parataenius derbesis (Solier,1851)   Chile 8Aphodiinae S.Am.1 (8)
Parataenius simulator (Harold,1868)   Argentina 1Aphodiinae S.Am.1 (1)
Paratrichius doenitzi (Harold,1879)   Japan 12Cetoniinae (12) Pal.21+22+25
Paratrichius itoi Tagawa,1938   Japan 1Cetoniinae Pal.22 (1)
Paratrichius septemdecimguttatus (Vollenhoven,1864)   Japan 1Cetoniinae Pal.22 (1)
Paratrichius spp.    Japan 2Cetoniinae Pal.22 (2)
Paratriodonta alicantina (Reitter,1890)   Spain 1Sericini Pal.2 (1)
Paratriodonta cinctipennis (Lucas,1846)   Tunisia 5Sericini Pal.1 (5)
Paratriodonta normandi (Baraud,1961)   Tunisia 7Melolonthinae Pal.18 (7)
Parelaphinis moesta (Gory & Percheron,1833)   South Africa 1Type Gory (1) Holotype 00003:1 of Micrelaphinis brincki Schein 1960
  Micrelaphinis brincki Schein 1960

Paroniticellus festivus Steven,1809   Armenia 2Scarabaeinae Pal.46 (2) as Oniticellus f.
Turkey 1Scarabaeinae (1) Pal.26
Paulianellus decoctor (Balthasar,1965)   Nepal 2Aphodiinae Or.7 (2)
Paulianellus nepalensis (Balthasar,1965)   Nepal 1Aphodiinae Or.7 (1)
Pectinichelus rhizotrogoides Ballion,1871   Russian Federation 1Melolonthinae Pal.5 (1)
Pedaria criberrima Waterhouse,1890   Guinea-Bissau 9Scarabaeinae Afr.2 (9)
Pedaria ouangoensis Cambefort,   Guinea-Bissau 15Scarabaeinae Afr.2 (15)
Pedaria picea Fåhraeus,1857   South Africa 7Scarabaeinae (7) Afr.32
Pedaria spp.    Gambia 3Scarabaeinae (3) Afr.18
Kenya 1Scarabaeinae (1) Afr.22
Senegal 1Scarabaeinae (1) Afr.22
South Africa 1Scarabaeinae (1) Afr.18
Unknown 1Scarabaeinae Afr.45 (1)
Pedaria tuberculigera Waterhouse,1890   Guinea-Bissau 2Scarabaeinae Afr.2 (2)
Pedaridium spp.    Paraguay 1Scarabaeinae (1) S.Am.5
Pedinorrhina mediana Westwood,1942   Cameroon 1Cetoniinae (1) Afr.18 (as P. septa Harlold)

Pedinorrhina picturata (Harold,1878)   Malawi 1Cetoniinae (1) Afr.5 (as Chondrorrhina p.)

Zambia 1Cetoniinae (1) Afr.5 (as Chondrorrhina p.)

Pedinorrhina trivittata (Schaum,1841)   Malawi 5Cetoniinae (5) Afr.5 (as Dyspilophora t.)

Namibia 3Cetoniinae Afr.8 (3) as Dyspilophora t.ssp. nigricans Kraatz
South Africa 8Cetoniinae (8) Afr.5+(8+?) (as Dyspilophora t.)

South Africa 3Cetoniinae Afr.8 (3) as Dyspilophora t.ssp. nigricans Kraatz
Pegylis sommeri Burmeister,1855   South Africa 1Melolonthinae Afr.4 (1)
Unknown 1Melolonthinae Afr.4 (1)
Pelidnota aeruginosa (Linnaeus,1758)   Brazil 3Rutelinae (3) S.Am 2
Pelidnota alliacea (Germar,184)   Colombia 1Rutelinae (1) S.Am 2
Pelidnota chalcoptera And. ??   Brazil 1Rutelinae (1) S.Am 3
Pelidnota cyanipes (Kirby,1818)   Brazil 1Rutelinae (1) S.Am 3
Unknown 1Rutelinae (1) S.Am 3
Pelidnota fulva Blanchard,1850   Brazil 1Rutelinae (1) S.Am 3
Unknown 1Rutelinae (1) S.Am 3
Pelidnota gracilis (Gory,1834)   Brazil 1Rutelinae (1) S.Am 3
Pelidnota kirbyi (Gray,1832)   Brazil 1Rutelinae (1) S.Am 3
Pelidnota kirschi Bates,1904   Unknown 1Rutelinae (1) S.Am 2
Pelidnota liturella (Kirby,1818)   Brazil 1Rutelinae (1) S.Am 2
Pelidnota lucida Burmeister,1844   Brazil 1Rutelinae (1) S.Am 2
Pelidnota notata Blanchard,1850   Colombia 2Rutelinae (2) S.Am 2
Unknown 1Rutelinae (1) S.Am 2
Pelidnota prasina (Burmeister,1844)   Brazil 1Rutelinae (1) S.Am 1
Colombia 1Rutelinae (1) S.Am 1
Venezuela 1Rutelinae (1) S.Am 1
Pelidnota pulchella (Kirby,1818)   Brazil 1Rutelinae (1) S.Am 2
Pelidnota punctata (Linnaeus,1758)   Canada 1Rutelinae N.Am.1 (1)
United States 1Connecticut
Rutelinae N.Am.1 (1)
United States 1District of Columbia
Rutelinae N.Am.1 (1)
United States 9Rutelinae N.Am.1 (9)
Pelidnota rugulosa Burmeister,1844   Brazil 1Rutelinae (1) S.Am 3
Pelidnota sanctijacobi Ohaus,1905 (Chalcoplethis)   Unknown 1Rutelinae (1) S.Am 2
Pelidnota sordida (Germar,1824)   Brazil 2Rutelinae (2) S.Am 3
Paraguay 1Rutelinae (1) S.Am 3
Unknown 1Rutelinae (1) S.Am 3
Pelidnota spp.    Argentina 1Rutelinae (1) S.Am 4
Pelidnota sumptuosa Vigors,1825   Unknown 1Rutelinae (1) S.Am 2
Pelidnota unicolor (Drury,1778)   Brazil 3Rutelinae (3) S.Am 1
Unknown 3Rutelinae (3) S.Am 1
Pelidnota virescens Burmeister,1844   Brazil 3Rutelinae (3) S.Am 2
Colombia 1Rutelinae (1) S.Am 2
Costa Rica 1Rutelinae (1) S.Am 2
Mexico 1Rutelinae (1) S.Am 2
Pentodon algerinus (Fuessly,1788)   Cyprus 1Dynastinae Pal.10 (1)
Egypt 1Dynastinae Pal.2 (1)
Greece 3Dynastinae Pal.2+4 (3)
Greece 2Rhodos
Dynastinae Pal.10 (2)
Israel 2Dynastinae Pal.12 (2) as P.algerinum Herbst
Morocco 1Dynastinae Pal.2 (1)
Spain 1Dynastinae Pal.4 (1)
Tunisia 1Dynastinae Pal.10 (1)
Turkey 12Dynastinae Pal.11 (6)
Pal.1+2 (6)
Unknown 2Dynastinae Pal.2+4 (2)
Pentodon algerinus bispinifrons Reitter,1894   Afghanistan 3Dynastinae Pal.3 (2)
Pal.10 (1)
Cyprus 2Dynastinae Pal.12 (2)
Greece 2Dynastinae Pal.11 (2)
Uzbekistan 5Dynastinae Pal.3 (5)
Pentodon algerinus dispar Baudi,1870   Egypt 1Dynastinae Pal.5 (1)
Oman 1Dynastinae Pal.5 (1)
Pentodon bengalensis Arr.   Afghanistan 1Dynastinae Pal.10 (1)
Pentodon bidens (Pallas,1771)   Cyprus 1Dynastinae Pal.12 (1)
Greece 1Dynastinae Pal.12 (1)
Turkey 1Dynastinae Pal.12 (1)
Pentodon bidens bidens (Pallas,1771)   Greece 1Crete
Dynastinae Pal.11 (1)
Russian Federation 2Dynastinae Pal.2 (2)
Pentodon bidens punctatus (Villers,1789)   Cyprus 3Dynastinae Pal.12 (3) as Pentodon punctatus
Italy 1Sardinia
Dynastinae Pal.10 (1)
Italy 20Dynastinae Pal.10 (2)
Pal.2+5+7 (18)
Malta 4Dynastinae Pal.10 (4)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Dynastinae Pal.2+6 (2)
Unknown 7Dynastinae Pal.2 (7)
Pentodon bidens sulcifrons Kuster,1848   Bulgaria 2Dynastinae Pal.10 (2)
Russian Federation 1Dynastinae Pal.2 (1)
Pentodon caminarius (Faldermann,1835)   Turkey 4Dynastinae Pal.11 (2)
Pal.12 (2)
Pentodon dubius Ballion,1871   Afghanistan 1Dynastinae Pal.10 (1)
Pentodon idiota (Herbst,1789)   Bulgaria 1Dynastinae Pal.6 (1)
Cyprus 1Dynastinae Pal.10 (1)
Greece 2Crete
Dynastinae Pal.11 (2)
Greece 3Dynastinae Pal.2 (2)
Pal.10 (1)
Morocco 1Dynastinae Pal.2 (1)
Turkey 3Dynastinae Pal.11 (3)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Dynastinae Pal.5 (1)
Unknown 6Dynastinae Pal.2 (6)
Pentodon puncticollis Burmeister,1847   Afghanistan 1Dynastinae Pal.10 (1)
Pentodon quadridens mongolicus  Motschulsky,1849   China 1Dynastinae Pal.2 (1)
Pentodon spp.    Cyprus 1Dynastinae Pal.12 (1)
Greece 2Crete
Dynastinae Pal.12 (2) as "Pentodon eremita Herbst"
Pentodon variolopunctatus Fairmaire,1879   Algeria 1Dynastinae Pal.9 (1)
Turkey 1Dynastinae Pal.12 (1)
Pentodontoschema aries Peringuey,1901   South Africa 1Dynastinae (1) Afr.12
Pentodontoschema capicola Peringuey,1901   Lesotho 1Dynastinae (1) Afr.12
South Africa 2Dynastinae (2) Afr.12
Pentodontoschema deceptor Peringuey,1901   Lesotho 1Dynastinae (1) Afr.12
Pentodontoschema nireum (Burmeister,1847)   Lesotho 2Dynastinae (2) Afr.12
Periserica interrupta (Walker,1859)   Sri Lanka 4Sericini Or.1 (4)
Peritrichia abdominalis Blanchard,1850   South Africa 2Hopliini Afr.2 (2)
Peritrichia albovillosa Schein,1959   South Africa 2Hopliini Afr.2 (2)
Peritrichia andreaei Schein,1959   South Africa 2Hopliini (2) Types 2006:2-3
Peritrichia antennata Schein,1959   South Africa 2Hopliini Afr.3 (2)
Peritrichia badiipennis Schein,1959   South Africa 2Hopliini Afr.2 (2)
Peritrichia bella Moser,1918   Namibia 3Hopliini Afr.2 (3)
Peritrichia bicolor Schein,1959   South Africa 1Hopliini Afr.3 (1)
Peritrichia capicola Fabricius,1781   South Africa 59Hopliini Afr.3 (59)
Peritrichia cinerea Olivier,1789   South Africa 4Hopliini Afr.2 (4)
Peritrichia dimidiata Burmeister,1844   South Africa 2Hopliini Afr.3 (2)
Peritrichia ditissima Peringuey,1902   South Africa 2Hopliini Afr.2 (2)
Peritrichia dubia Schein,1959   South Africa 3Hopliini (3) Types 2010:2-4
Peritrichia fasciata Burmeister,1844   South Africa 1Hopliini Afr.2 (1)
Peritrichia flabellata Schein,1959   South Africa 1Hopliini Afr.2 (1)
Peritrichia flavoornata Moser,1918   Namibia 3Hopliini Afr.2 (3)
Peritrichia guttata Burmeister,1844   South Africa 2Hopliini Afr.2 (2)
Peritrichia hessei Schein,1959   South Africa 2Hopliini Afr.2 (2)
Peritrichia nigrita Blanchard,1850   South Africa 2Hopliini Afr.2 (2)
Peritrichia nitidipennis Blanchard,1850   South Africa 4Hopliini Afr.2 (4)
Peritrichia nitidipennis transiens Blanchard,1850   South Africa 2Hopliini Afr.2 (2)
Peritrichia nitidipennis tulbaghina Blanchard,1850   South Africa 7Hopliini Afr.2 (7)
Peritrichia pistrinaria Peringuey,1888   South Africa 2Hopliini Afr.3 (2)
Peritrichia proboscidea Burmeister,1844   South Africa 2Hopliini Afr.3 (2)
Peritrichia pseudopistrinaria Schein,1959   South Africa 7Hopliini Afr.3 (7)
Peritrichia pseudopistrinaria meridionalis Schein,1959   South Africa 2Hopliini (2) Types 2011:2-3
Peritrichia pseudoplebeja Schein,1959   South Africa 1Hopliini (1) Type 2007:2
Peritrichia pseudursa Schein,1959   South Africa 3Hopliini (3) Afr.2+19
Peritrichia pulchella Peringuey,1902   South Africa 2Hopliini Afr.2 (2)
Peritrichia pygidialis Peringuey,1902   South Africa 2Hopliini Afr.3 (2)
Peritrichia rigida Schein,1959   South Africa 2Hopliini Afr.3 (2)
Peritrichia rufotibialis Schein,1959   South Africa 1Hopliini Afr.2 (1)
Peritrichia saga Peringuey,1902   South Africa 2Hopliini Afr.2 (2)
Peritrichia spuria Peringuey,1902   South Africa 2Hopliini Afr.3 (2)
Peritrichia spuria pseudospuriella Peringuey,1902   South Africa 2Hopliini Afr.3 (2)
Peritrichia spuria spuriella Peringuey,1902   South Africa 6Hopliini (6) Types 2009:2-7
Peritrichia subsquamosa Schein,1959   South Africa 44Hopliini Afr.19 (42)
Type Schein (2) Paratype 02008:2-3 of Peritrichia subsquamosa Schein 1959
  Peritrichia subsquamosa Schein 1959

Peritrichia tristis Burmeister,1844   South Africa 1Hopliini Afr.2 (1)
Peritrichia ursus Olivier,1789   South Africa 2Hopliini Afr.2 (2)
Peritrichia vansoni Schein,1959   South Africa 4Hopliini (4) Types 2005:2-3
+2-Afr.2
Peritrichia ypsilon Schein,1959   South Africa 2Hopliini Afr.3 (2)
Phaeadoretus comptus (Menetries,1849)   Russian Federation 2Rutelinae Pal.9 (2)
Phaeaphodius cavatus (Boucomont,1932)   Congo 1Aphodiinae Afr.20 (1)
Phaeaphodius himalorectus (Ahrens & Stebnicka,1997)   Nepal 3Aphodiinae Or.2 (3) Paratypes
Phaeaphodius motoi (Paulian,1939)   Congo 1Aphodiinae Afr.24 (1)
Phaeaphodius rectus (Motschulsky,1866)   Japan 7Aphodiinae (5) Pal.18+24
Pal.28 (2)
Turkey 1Aphodiinae Pal.28 (1)
Phaeaphodius tauricola (Hrubant,1961)   Turkey 1Aphodiinae Pal.28 (1)
Phaedimus lumawigi Arnaud,1989   Philippines 1Cetoniinae Or.7 (1)
Phaenomeris besckei Mannerheim,1838   Mozambique 2Phaenomerinae Afr.3 (2)
Tanzania 6Phaenomerinae Afr.3 (6)
Unknown 2Phaenomerinae Afr.3 (2)
Phaenomeris splendida Ohaus,1901   Kenya 1Phaenomerinae Afr.3 (1)
Phaenomeris spp.    Unknown 2Phaenomerinae Afr.3 (2)
Phalacronothus avunculus Balthasar,1946   Sri Lanka 3Aphodiinae Or.1 (3) as Aphodius
Phalacronothus biguttatus (Germar,1823)   Italy 1Aphodiinae Pal.30 (1)
Phalacronothus botulus Balthasar,1945   Japan 1Aphodiinae Pal.17 (1) as Aphodius
Phalacronothus carinulatus Motschulsky,1864   Sri Lanka 6Aphodiinae Or.1 (6) as Aphodius
Phalacronothus ceylonensis Schmidt,1912   Sri Lanka 4Aphodiinae Or.1 (4) as Aphodius
Phalacronothus quadrimaculatus (Linnaeus,1761)   France 2Aphodiinae Pal.16 (2) as Aphodius
Germany 1Aphodiinae Pal.16 (1) as Aphodius
Italy 1Aphodiinae Pal.15 (1) as Aphodius
United Kingdom 4England
Aphodiinae Pal.15 (4) as Aphodius
Caucasus, Ryssland / Georgien / Azerbajdzjan / Armenien / Iran / Turkiet 1Aphodiinae Pal.20 (1) as Aphodius
Unknown 3Aphodiinae Pal.20 (3) as Aphodius
Phalangonyx bibatillatus (Petrovitz,1955)   Afghanistan 1Melolonthinae (1) Pal.5 (as Tanyproctus b.)

Phalangonyx coniceps Reitter,1889   Iran 1Melolonthinae Pal.9 (1)
Phalops spp.    Guinea-Bissau 1Scarabaeinae Afr.3 (1)
Phanaeus chalcomelas (Perty,1830)   Brazil 2Scarabaeinae S.Am.5 (2)
Peru 1Scarabaeinae S.Am.1 (1)
Phanaeus daphnis Harold,1863   Mexico 1Scarabaeinae N.Am.1 (1)
Phanaeus kirbyi Vigors,1825   Brazil 2Scarabaeinae S.Am.5 (2)
Phanaeus splendidulus (Fabricius,1781)   Brazil 3Scarabaeinae (3) S.Am.5
Phanaeus spp.    Brazil 1Scarabaeinae S.Am.7 (1)
Pharaonus caucasicus (Reitter,1888)   Portugal 7Azores
Rutelinae Pal.11 (7)
Pharaonus lederi (Reitter,1888)   Turkmenistan 1Rutelinae Pal.8 (1)
Pharaonus semenovi (Reitter,1887)   Turkmenistan 1Rutelinae Pal.8 (1)
Pharaphodius acutidens (Petrovitz,1964)   Gambia 7Aphodiinae Afr.2 (7)
Senegal 1Aphodiinae Afr.2 (1)
Tanzania 20Aphodiinae Afr.2 (20)
Pharaphodius argutus (Schmidt,1911)   India 1Aphodiinae Pal.18 (1)
Sierra Leone 1Aphodiinae (1) Afr.15
Unknown 3Aphodiinae Pal.18 (3)
Zimbabwe 1Aphodiinae (1) Afr.23
Pharaphodius bechuanus (Petrovitz,1958)   Botswana 3Aphodiinae (3) Afr.2+16 (inc.1 paratype)

Kenya 2Aphodiinae Afr.2 (2)
South Africa 3Aphodiinae (3) Afr.16
Pharaphodius corax (Balthasar,1937)   Namibia 10Aphodiinae Afr.35 (10)
South Africa 2Aphodiinae Afr.16 (1)
Afr.23 (1)
Pharaphodius costatulus (Schidt,1908)   Nepal 51Aphodiinae Or.3 (51)
Pharaphodius crenatus (Harold,1862)   Thailand 6Aphodiinae Or.3 (6)
Pharaphodius curvodilatatus (Schmidt,1909)   Tanzania 1Aphodiinae Afr.2 (1)
Pharaphodius desertus (Klug,1845)   Gambia 1Aphodiinae Afr.2 (1)
Pharaphodius difficilis (Endrödi,1955)   Rwanda 1Aphodiinae Afr.26 (1) paratype
Pharaphodius discoidalis (Boheman,1857)   Kenya 1Aphodiinae Afr.2 (1) as ab. neghellinus Balthasar
Namibia 1Aphodiinae Afr.35 (1)
South Africa 3Aphodiinae Afr.23 (1)
Afr.31 (1)
Afr.35 (1)
Pharaphodius discolor (Erichson,1842)   Chad 2Aphodiinae (2) Afr.20
Gambia 2Aphodiinae Afr.2 (2)
Ghana 1Aphodiinae (1) Afr.18
Nigeria 2Aphodiinae Afr.28 (2)
Senegal 4Aphodiinae Afr.2 (4)
Sierra Leone 8Aphodiinae (8) Afr.15
Pharaphodius dubiosus Péringuey,1901   South Africa 9Aphodiinae Afr.31 (9)
Pharaphodius evae (Endrödi,1973)   Gambia 1Aphodiinae Afr.2 (1)
Pharaphodius guineensis (Klug,1835)   Ethiopia 1Aphodiinae Afr.20 (1)
Gambia 45Aphodiinae Afr.2 (45)
Mozambique 1Aphodiinae (1) Afr.20
Nigeria 2Aphodiinae Afr.28 (2)
Senegal 23Aphodiinae Afr.2 (23)
Sierra Leone 1Aphodiinae (1) Afr.15
South Africa 4Aphodiinae Afr.34 (4)
Zimbabwe 1Aphodiinae (1) Afr.16
Pharaphodius impurus (Roth,1851)   Ethiopia 2Aphodiinae Afr.29 (2)
Kenya 9Aphodiinae Afr.2 (9)
Namibia 1Aphodiinae Afr.27 (1)
South Africa 39Aphodiinae (34) Afr.22+25
Afr.31 (5)
Zimbabwe 9Aphodiinae (9) Afr.16
Pharaphodius intercalaris (Peringuey,1901)   Namibia 2Aphodiinae Afr.22 (2)
South Africa 57Aphodiinae Afr.22 (56)
Afr.35 (1)
Pharaphodius marginellus (Fabricius,1781)   Indonesia 5Java
Aphodiinae Or.2 (5)
Pharaphodius massaicus (Gerstaecker,1884)   Kenya 1Aphodiinae (1) Afr.24
Pharaphodius nigripennis (Petrovitz,1958)   Guinea 2Aphodiinae (2) Afr.16 (incl. 2 paratypes)

Pharaphodius posticus (Boheman,1857)   Botswana 1Aphodiinae Afr.2 (1)
Namibia 1Aphodiinae Afr.35 (1)
South Africa 2Aphodiinae Afr.2 (2)
Pharaphodius priscus (Motschulsky,1858)   Malaysia 6Aphodiinae Or.3 (6)
Thailand 1Aphodiinae Or.3 (1)
Pharaphodius pseudodesertus (Bordat,1996)   South Africa 3Aphodiinae Afr.2 (3) Paratypes
Pharaphodius robustus (Walker,1858)   Sri Lanka 2Aphodiinae Or.1 (2)
Pharaphodius rubricosus (Boheman,1857)   Namibia 3Aphodiinae (3) Afr.22
South Africa 3Aphodiinae (3) Afr.22
Pharaphodius rufomaculatus (Petrovitz,1958)   Ghana 3Aphodiinae (3) Afr.17
Guinea 2Aphodiinae (2) Afr.16 (incl. 2 paratypes)

Pharaphodius rugosostriatus (Waterhouse,1875)   Japan 3Aphodiinae Pal.18 (2)
Pal.28 (1)
Pharaphodius russatus (Erichson,1842)   Gambia 1Aphodiinae Afr.2 (1)
Lesotho 4Aphodiinae (4) Afr.22
Nigeria 2Aphodiinae (2) Afr.20
South Africa 8Aphodiinae (8) Afr.22+25
Pharaphodius spp.    Namibia 5Aphodiinae Afr.30 (1)
Afr.35 (4)
Pharaphodius subcostatus (Kolbe,1886)   China 1Aphodiinae Pal.18 (1)
Pharaphodius ugandanus (Balthasar,1963)   Tanzania 1Aphodiinae Afr.4 (1)
Pharaphodius zavattarii (Müller,1941)   Kenya 4Aphodiinae Afr.35 (4)
Unknown 1Aphodiinae Afr.25 (1) as Macrotetrus
Phileurus didymus (Linnaeus,1758)   Colombia 1Dynastinae (1) S.Am.21
Peru 1Dynastinae (1) S.Am.31
Unknown 3Dynastinae (3) S.Am.21
Phileurus exavatus Prell,1911   Brazil 2Dynastinae (2) S.Am.21
Unknown 1Dynastinae (1) S.Am.21
Phileurus hospes Burmeister,1847   Brazil 2Dynastinae (2) S.Am.21
Phileurus truncatus (Beauvois,1805)   United States 1Georgia
Dynastinae N.Am.2 (1)
Phileurus valgus (Olivier,1789)   Brazil 1Dynastinae (1) S.Am.21
Phileurus valgus meridionalis (Olivier,1789)   Unknown 1Dynastinae (1) S.Am.21
Phileurus youngi Ratcliffe,1988   Costa Rica 1Dynastinae (1) S.Am.36
Phonotaenia aequinoctialis Olivier,1789   Unknown 5Cetoniinae (5) Afr.26
Phonotaenia balteata De Geer,1778   Ethiopia 1Cetoniinae (1) Afr.26
Unknown 2Cetoniinae (2) Afr.26
Phonotaenia scalaris Gory & Percheron,1833   Cameroon 1Cetoniinae (1) Afr.26
Phoxomela umbrosa (Gory-Percheron,1833)   South Africa 10Cetoniinae (10) Afr.15
Unknown 2Cetoniinae (2) Afr.26
Phyllognathus burmeisteri Arrow,1911   Senegal 1Dynastinae (1) Afr.19
Phyllognathus degener Fairmaire,1891   Guinea-Bissau 4Dynastinae (4) Afr.19
Phyllognathus dionysius (Fabricius,1792)   Sri Lanka 1Dynastinae Or.3 (1)
Phyllognathus excavatus (Forster,1771)   Algeria 4Dynastinae Pal.3 (3)
Pal.9 (1)
Egypt 2Dynastinae Pal.3 (2)
France 3Dynastinae Pal. (3) Pal.3+5
Greece 6Dynastinae Pal.9 (1)
Pal.3+5 (5)
Italy 1Dynastinae Pal.6 (1)
Morocco 1Dynastinae Pal.3 (1)
Spain 24Canary Islands
Dynastinae Pal.11 (6)
Pal.1+4 (18)
Spain 4Dynastinae Pal. (4) Pal.3+5
Spain 2Canary Islands
Dynastinae Pal.19 (2)
Spain 1Canary Islands
Dynastinae Pal.11 (1)
Tunisia 6Dynastinae Pal.9 (1)
Pal.3+5+6 (5)
Unknown 10Dynastinae Pal.3 (10)
Phyllognathus orion (Olivier,1789)   Senegal 2Dynastinae Afr.5 (2)
Phyllopertha arenaria Brulle,1832   Turkey 1Rutelinae Pal.6 (1)
Phyllopertha diversa Waterhouse,1875   Japan 2Rutelinae Pal.10 (2)
Phyllopertha fuscata Niijima & Kinoshita,1923   China 1Rutelinae Pal.6 (1)
Phyllopertha hirtella Brulle,1832   Greece 1Rutelinae Pal.6 (1)
Phyllopertha horticola (Linnaeus,1758)   Belgium 1Rutelinae Pal.10 (1)
France 1Rutelinae Pal.6 (1)
Germany 8Rutelinae (8) Pal.6+10
Italy 1Rutelinae Pal.6 (1)
Romania 2Rutelinae Pal.6 (2)
Spain 1Rutelinae Pal.11 (1)
Sweden 533  Sk Bl Ha Sm Öl Go GS Ög Vg Bo Ds Nä Sö Up Vr Dr
Alan Dufberg 48 (12)
Rutelinae Main (521)
Switzerland 1Rutelinae Pal.10 (1)
Phyllopertha horticoloides Lin,1965   China 1Rutelinae Pal.10 (1)
Phyllopertha irregularis Waterhouse,1875   Japan 3Rutelinae Pal.10 (3)
Phyllopertha orientalis Waterhouse,1875   Japan 2Rutelinae Pal.10 (2)
Phyllopertha puncticollis Reitter,1888   China 1Rutelinae Pal.6 (1)
Phyllophaga futilis (LeConte,1850)   United States 2Indiana
Melolonthinae N.Am.3 (2)
Phyllophaga hirticula (Knoch,1801)   United States 1Illinois
Melolonthinae (1) N.Am.1 (as Lachnost. hirticula Knoch)
Phyllophaga latifrons LeConte,1856   United States 1Florida
Melolonthinae N.Am.1 (1)
Phyllophaga pleei (Blanchard,1850)   Guadeloupe 1Melolonthinae S.Am.1 (1)
Phyllophaga spp.    Canada 3Ontario
Melolonthinae N.Am.1 (3)
Phyllotocus australis (Boisduval,1835)   Australia 1Victoria
Melolonthinae Pac.1 (1)
Phyllotocus erythropterus Blanchard,1850   Australia 1Melolonthinae Pac.1 (1)
Phyllotocus macleayi Fischer,1823   Australia 1Melolonthinae Pac.1 (1)
Unknown 1Melolonthinae Pac.1 (1)
Phyllotocus navicularis Blanchard,1850   Australia 3Queensland
Melolonthinae Pac.1 (3)
Australia 1Melolonthinae Pac.1 (1)
Phyllotocus rufipennis (Boisduval,1835)   Australia 4Melolonthinae Pac.1 (4)
Australia 1Victoria
Melolonthinae Pac.1 (1)
Unknown 1Melolonthinae Pac.1 (1)
Pimelopus nothus Burmeister,1847   Australia 2Dynastinae Pac.2 (2)
Pinacotarsus dohrni Harold,1875   Guinea-Bissau 2Scarabaeinae Afr.3 (2)
Placodidus compransor Peringuey,1900   South Africa 1Cetoniinae (1) Afr.17
Plaesiorrhina ochreata    Madagascar 1Cetoniinae (1) Afr.18
Plaesiorrhina plana (Wiedemann,1821)   Mozambique 1Cetoniinae Afr.8 (1)
South Africa 7Cetoniinae Afr.8 (7)
Plaesiorrhina watkinsiana    Cameroon 1Cetoniinae (1) Afr.18
Planolinus borealis (Gyllenhal,1827)   Sweden 193  Sk Bl Ha Sm Öl Go Ög Vg Bo Sö Up Vs Vr Dr Gä Hr Jä Ån Vb Nb Ås Ly Lu To
Alan Dufberg 48 (14)
Aphodiinae Main (179)
Planolinus fasciatus (A.G. Olivier,1789)   Norway 4Aphodiinae Main (4)
Sweden 165  Sk Bl Ha Sm Öl Ög Vg Sö Up Vs Vr Dr Gä Hs Hr Jä Vb Nb Ås Lu To
Alan Dufberg 48 (32)
Aphodiinae Main (133)
Planolinus tenellus (Say,1823)   Canada 1Ontario
Aphodiinae N.Am.2 (1)
United States 1Washington
Aphodiinae N.Am.2 (1)
United States 4Colorado
Aphodiinae N.Am.4 (4)
Planolinus vittatus (Say,1825)   Greece 2Aphodiinae Pal.6 (2)
Russian Federation 1Aphodiinae (1) Pal.18 (as ssp. mundus Reitter)

Tajikistan 2Aphodiinae (2) Pal.18 (as ssp. tjanshanicus Balthasar)

Turkey 1Aphodiinae Pal.27 (1)
United States 7Arizona
Aphodiinae N.Am.1 (7)
Platychelus fallax Schein,1959   South Africa 30Type Schein (30) Holotype Paratype 02036:1-30 of Platychelus fallax Schein 1959
  Platychelus fallax Schein 1959

Platychelus gravidus Burmeister,1844   South Africa 1Hopliini Afr.12 (1)
Platychelus lutosus Schein,1959   South Africa 9Type Schein (9) Holotype Paratype 02038:1-9 of Platychelus lutosus Schein 1959
  Platychelus lutosus Schein 1959

Platychelus melanurus Burmeister,1844   South Africa 2Hopliini Afr.12 (2)
Platychelus pyropygus Burmeister,1844   Unknown 2Hopliini Afr.20 (2) Old spms
partly poor labeling.
Platychelus semihirtuss Burmeister,1844   South Africa 1Hopliini Afr.12 (1)
Platychelus spp.    South Africa 1Hopliini Afr.12 (1)
Platychelus transvaalensis Schein,1959   South Africa 3Type Schein (3) Holotype Paratype 02037:1-3 of Platychelus transvaalensis Schein 1959
  Platychelus transvaalensis Schein 1959

Platychelus virgatus Burmeister,1844   South Africa 22Hopliini Afr.13 (22)
Platycoelia humeralis Bates,1888   Ecuador 3Rutelinae (3) S.Am.7
Platycoelia spp.    Ecuador 1Rutelinae (1) S.Am.7
Unknown 1Rutelinae (1) S.Am.7
Platygenia barbata Afzelius,1817   Cameroon 2Cetoniinae Afr.31 (2)
Kenya 1Cetoniinae Afr.31 (1)
Unknown 3Cetoniinae Afr.31 (3)
Platytomus laevistriatus (Perris,1870)   Spain 1Aphodiinae Pal.12 (1)
Tunisia 17Aphodiinae (17) Pal.12
Platytomus longulus Cartwright,1948   Guatemala 1Aphodiinae S.Am.1 (1)
United States 15South Carolina
Aphodiinae N.Am.3 (15) as Pleurophorus l.
Platytomus micros (Bates,1887)   United States 3California
Aphodiinae (3) N.Am.2 (as Pleurophorus m.)

Platytomus spp.    United States 1Arizona
Aphodiinae N.Am.4 (1)
Platytomus tibialis (Fabricius,1798)   Spain 4Canary Islands
Aphodiinae Pal.3 (4)
Spain 5Aphodiinae (5) Pal.2+11+12+24
Tunisia 5Aphodiinae Pal.12 (1)
Pal.30 (4)
Platytomus variolosus (Kolenati,1846)   Afghanistan 15Aphodiinae Pal.9 (15)
Kyrgyzstan 1Aphodiinae (1) Pal.16 (as Pleurophorus v.)

Pleuraphodius anceps (Schmidt,1911)   Gambia 21Aphodiinae Afr.8 (21)
Ghana 2Aphodiinae Afr.18 (2)
Senegal 2Aphodiinae Afr.8 (1)
Afr.19 (1)
Pleuraphodius hanstroemi Landin,1956   South Africa 1Aphodiinae Afr.19 (1) Holotype 03484:1 of Aphodius hanstroemi Landin 1956
  Aphodius hanstroemi Landin 1956

Pleuraphodius landini (Endrödi,1956)   Angola 1Aphodiinae (1) Afr.19
Congo, Democratic Republic of the 1Aphodiinae Afr.26 (1) type
Gambia 44Aphodiinae Afr.9 (44)
Senegal 7Aphodiinae Afr.9 (7)
Sierra Leone 2Aphodiinae (2) Afr.15
Pleuraphodius letabus Landin,1956   South Africa 2Type Landin (2) Holotype Paratype 03485:1-2 of Aphodius letabus Landin 1956
  Aphodius letabus Landin 1956

Pleuraphodius lineatosulcatus (Harold,1859)   South Africa 15Aphodiinae Afr.32 (15)
Pleuraphodius mutilus Schmidt,1911   Angola 1Aphodiinae Afr.18 (1) Paratype of Aphodius chaboti Paulian
1942
South Africa 15Aphodiinae Afr.24 (1)
Afr.31 (14)
Pleuraphodius ratnapurensis Stebnicka,1988   Sri Lanka 3Type Stebnicka (3) Holotype Paratype 00924:1-3 of Aphodius ratnapurensis Stebnicka 1988
  Aphodius ratnapurensis Stebnicka 1988

Pleuraphodius teter (Roth,1851)   Kenya 2Aphodiinae Afr.9 (2)
Lesotho 12Aphodiinae (12) Afr.18+19
Namibia 1Aphodiinae (1) Afr.18
South Africa 10Aphodiinae (10) Afr.18
Tanzania 6Aphodiinae Afr.9 (6)
Pleurophorus africanus Pittino,1984   South Africa 1Aphodiinae Afr.31 (1)
Pleurophorus anatolicus Petrovitz,1961   Azerbaijan 1Aphodiinae Pal.15 (1)
Cyprus 29Aphodiinae Pal.12 (29)
Lebanon 1Aphodiinae Pal.15 (1)
Syria 1Aphodiinae Pal.15 (1)
Turkey 53Aphodiinae (16) Pal.10+15
Pal.26 (36)
Pal.30 (1)
Pleurophorus apicipennis Reitter,1892   Afghanistan 209Aphodiinae (209) Pal.8 +5
Algeria 2Aphodiinae (2) Pal.15
Turkey 2Aphodiinae (2) Pal.15
Uzbekistan 4Aphodiinae (4) Pal.12+15
Pleurophorus caesus (Creutzer,1796)   Bulgaria 16Aphodiinae (16) Pal.11+12
Croatia 5Aphodiinae Pal.30 (5)
Denmark 1Aphodiinae (1)
Egypt 2Aphodiinae (2) Afr.13
France 7Aphodiinae (7) Pal.2+16
Germany 5Aphodiinae (5) Pal.16
Greece 42Aphodiinae (41) Pal.2+12
Pal.30 (1)
Greece 2Rhodos
Aphodiinae Pal.30 (2)
Italy 23Aphodiinae (23) Pal.3+11+12
Portugal 2Azores
Aphodiinae Pal.13 (2)
Portugal 1Madeira
Aphodiinae Pal.4 (1)
Romania 1Aphodiinae (1) Pal.12
Spain 13Aphodiinae (13) Pal.2+12+15
Sweden 2 Skåne Halland
Aphodiinae Main (2)
Tunisia 5Aphodiinae (5) Pal.12
Turkey 6 Alan Dufberg 48 (3)
Aphodiinae Pal.12 (1)
Aphodiinae Pal.30 (2)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 4Aphodiinae (4) Pal.12
Pleurophorus cracens Motschulsky,1863   Sri Lanka 21Aphodiinae Or.2 (21)
Pleurophorus mediterranicus Pittino & Mariani,1986   Italy 1Aphodiinae (1) Pal.12
Spain 2Aphodiinae (2) Pal.12
Tunisia 1Aphodiinae (1) Pal.12
Pleurophorus pannonicus Petrovitz,1961   Austria 1Aphodiinae Pal.15 (1)
Cyprus 1Aphodiinae Pal.16 (1)
Greece 1Corfu
Aphodiinae Pal.24 (1)
Hungary 1Aphodiinae Pal.15 (1) Pal.15
Romania 1Aphodiinae Pal.15 (1) Pal.15
Pleurophorus spp.    Bulgaria 3Aphodiinae (3) Pal.13
Mexico 2Aphodiinae N.Am.3 (2)
Spain 3Aphodiinae (3) Pal.13
Podalgus cuniculus arabicus Fairmaire,1895   Oman 45Dynastinae (45) Pal.7+8
Podischnus agenor (Olivier,1789)   Colombia 9Dynastinae S.Am.5 (9)
Unknown 2Dynastinae S.Am.5 (2)
Poecilophila hebraea (Olivier,1789)   South Africa 11Cetoniinae (11) Afr.17+?+27
Poecilophila maculatissima (Boheman,1860)   Malawi 1Cetoniinae (1) Afr.18
Tanzania 3Cetoniinae Afr.29 (3)
Unknown 2Cetoniinae Afr.29 (2)
Polybaphes abyssinica Schurhoff,1935   Gambia 9Cetoniinae Afr.6 (9)
Polybaphes balteata (DeGeer,1778)   Gambia 3Cetoniinae Afr.6 (3)
Polybaphes sanguinolenta (Olivier,1789)   South Africa 1Cetoniinae (1) Afr.26
Unknown 2Cetoniinae (2) Afr.26
Polybaphes spp.    Unknown 1Cetoniinae (1) Afr.26
Polybaphes subfasciata (Swederus,1787)   South Africa 2Cetoniinae (2) Afr.14 (as Phonotaenia s.)
Unknown 2Cetoniinae Afr.26 (2)
Polyphylla adspersa Motschulsky,1854   Afghanistan 23Melolonthinae Pal.10 (23)
Russian Federation 3Melolonthinae Pal.2 (3)
Polyphylla alba (Pallas,1773)   Russian Federation 4Melolonthinae Pal.2 (4)
Polyphylla asiatica Desbr.   Turkey 5Melolonthinae Pal.6 (5)
Polyphylla boyi Brulle,1832   Greece 1Melolonthinae Pal.5 (1)
Polyphylla fullo (Linnaeus,1758)   Germany 6Melolonthinae Pal.2 (1)
Pal.5 (1)
Pal.11 (1)
Pal.17 (3)
Greece 1 Crete
Alan Dufberg 48 (1)
Hungary 1Melolonthinae Pal.2 (1)
Italy 1Melolonthinae Pal.11 (1)
Romania 4Melolonthinae (3) Pal.5+11
Pal.18 (1)
Spain 1Melolonthinae Pal.5 (1)
Sweden 2 Skåne Västergötland
Melolonthinae Main (2)
Unknown 5Melolonthinae Pal.2 (2)
Pal.17 (3)
Polyphylla gracilicornis Blanchard,1871   Spain 1Melolonthinae Pal.2 (1)
Unknown 1Melolonthinae Pal.2 (1)
Polyphylla irrorata (Gebler,1841)   Russian Federation 3Melolonthinae Pal.2 (3)
Polyphylla laticollis Lewis,1887   Japan 3Melolonthinae Pal.11 (3)
Polyphylla olivieri (Castelnau,1840)   Iran 4Melolonthinae Pal.2 (4)
Turkey 2Melolonthinae Pal.6 (1)
Pal.16 (1)
Polyphylla sobrina Casey,1914   United States 1California
Melolonthinae N.Am.1 (1)
Polyphylla variolosa (Hentz,1830)   United States 3Massachusetts
Melolonthinae N.Am.1 (3)
Unknown 5Melolonthinae N.Am.3 (5)
Polystalactica furfurosa (Burmeister,1847)   South Africa 1Cetoniinae (1) Afr.14
Polystalactica spp.    Unknown 3Cetoniinae (3) Afr.26
Popillia adamas Newman,1838   India 1Rutelinae Or.8 (1)
Popillia amabilis Arrow,1913   China 1Rutelinae Pal.7 (1)
Popillia biguttata (Wiedemann,1821)   Indonesia 4Java
Rutelinae Or.8 (4)
Popillia bipunctata (Fabricius,1787)   Kenya 2Rutelinae Afr.2 (2)
South Africa 7Rutelinae (7) Afr.1+2
South Africa 16Rutelinae Afr.1 (16)
Tanzania 4Rutelinae Afr.2 (4)
Unknown 1Africa
Rutelinae Afr.3 (1)
Zambia 4Rutelinae Afr.2 (4)
Popillia callipyga Dohrn,1879   Cameroon 12Rutelinae (12) Afr.2+3+?
Popillia complanata Newman,1838   India 3Rutelinae Or.8 (3)
Popillia cupricollis Hope,1831   India 5Rutelinae Or.8 (5)
Popillia cyanea Hope,1931   Taiwan 1Rutelinae Pal.7 (1)
Popillia discalis Walker,1859   Sri Lanka 7Rutelinae Or.5 (7)
Popillia distinguenda Fairmaire,1887   Tanzania 2Rutelinae Afr.2 (2)
Unknown 1Rutelinae Afr.2 (1)
Popillia dorsofasciata Fairmaire,1887   Tanzania 1Rutelinae Afr.3 (1)
Popillia ertli Ohaus,1911   Tanzania 2Rutelinae Afr.3 (2)
Popillia fallaciosa Fairmaire,1889   China 1Rutelinae Pal.10 (1)
Popillia flavosellata Fairmaire,1886   China 2Rutelinae Pal.7 (2)
China 9Rutelinae (9) Pal.6+7 (as Popillia atrocoerulea Bates)
China 1Rutelinae (1) Pal.7 (as P. inconstans Fairmaire)
Korea, Republic of 1Rutelinae (1) Pal.7 (as Popillia atrocoerulea Bates)
Popillia gemma Newman,1839   India 5Rutelinae Or.8 (5)
Popillia hilaris Kraatz,1892   Tanzania 8Rutelinae (8) Afr.2+3+?
Unknown 8Rutelinae Afr.3 (8)
Zambia 1Rutelinae Afr.2 (1)
Popillia histeroidea (Gyllenhal,1817)   China 1Rutelinae Pal.7 (1)
Viet Nam 1Rutelinae (1) Or.8 (as P.caerulea)

Popillia insularis Lewis,1895   China 5Rutelinae Pal.7 (5)
Unknown 5Rutelinae Pal.7 (5)
Popillia japonica Newman,1838   China 6Rutelinae Pal.7 (6)
Japan 19Rutelinae Pal.7 (19)
United States 3Massachusetts
Rutelinae N.Am.2 (3)
United States 3Minnesota
Rutelinae N.Am.2 (3)
Popillia lewisi Arrow,1913   China 3Rutelinae Pal.7 (3)
Popillia maclellandi Hope,1843   China 1Rutelinae Pal.7 (1)
Popillia meinhardti Kolbe,1894   Cameroon 23Rutelinae Afr.2 (23)
Popillia mongolica Arrow,1913   China 1Rutelinae Pal.7 (1)
Popillia mutans Newman,1838   China 2Rutelinae Pal.7 (2)
Russian Federation 1Rutelinae Pal.10 (1)
Unknown 3Rutelinae Pal.7 (3)
Popillia ohausi Kolbe,1897   Tanzania 5Rutelinae Afr.3 (5)
Unknown 2Rutelinae Afr.3 (2)
Popillia opacicollis Kraatz,1892   India 1Rutelinae Or.8 (1)
Popillia parvula Burmeister,1844   Borneo, Indonesia / Malaysia 4Rutelinae Or.8 (4)
Popillia pustulata Fairmaire,1887   Unknown 1Rutelinae Pal.7 (1)
Popillia quadriguttata (Fabricius,1787)   China 6Rutelinae (6) Pal.7+8
Japan 3Rutelinae (3) Pal.7 (as Popillia bogdanowi Ballion)
Japan 2Rutelinae Pal.8 (2)
United States 3New York
Rutelinae N.Am.2 (3)
Popillia rufipes (Fabricius,1787)   Sierra Leone 2Rutelinae Afr.2 (2)
Popillia semiaenea Kraatz,1892   China 1Rutelinae Pal.7 (1)
Unknown 3Rutelinae Pal.7 (3)
Popillia tandallae Kolbe,1903   Tanzania 2Rutelinae Afr.3 (2)
Porphyronota carnifex (Fabricius,1781)   South Africa 2Cetoniinae (2) Afr.17+28
Unknown 2Cetoniinae Afr.28 (2)
Prionoryctes capreolus (Quedenfeldt, 1884)   Cameroon 1Dynastinae Afr.7 (1)
Pristerophora paulseni Smith,2008   Chile 1Melolonthinae S.Am.1 (1) Paratype
Proagoderus chrysopes (Bates,1888)   Tanzania 1Scarabaeinae Afr.45 (1)
Proagopertha lucidula (Falderman,1835)   China 1Rutelinae (1) Pal.6 (as Phyllopertha l.)

Proagopertha pubicollis (Waterhouse,1875)   Japan 2Rutelinae Pal.10 (2)
Propomacrus bimucronatus (Pallas,1781)   Turkey 5Euchirinae Pal.1 (5)
Prosphileurus liberianus (Dohrn,1875)   Sierra Leone 1Dynastinae (1) Afr.19
Protaetia acuminata (Fabricius,1775)   Unknown 1Cetoniinae Or.4 (1)
Protaetia aeruginosa (Drury,1770)   Croatia 7Cetoniinae Pal.30 (2) as "Cetonia auriginosa"
Pal.16+25 (5)
Germany 1Cetoniinae Pal.9 (1)
Greece 8Cetoniinae Pal.6 (8)
Hungary 2Cetoniinae (2) Pal.16
Syria 1Cetoniinae Pal.25 (1)
Protaetia affinis (Andersch,1797)   Austria 3Cetoniinae Pal.15 (1)
Pal.27 (2)
France 1Cetoniinae Pal.15 (1)
Germany 1Cetoniinae Pal.9 (1)
Portugal 1Cetoniinae Pal.31 (1) as Eupotosia affinis
Protaetia affinis pyrodera Reitter,1891   Syria 9Cetoniinae Pal.15 (9)
Protaetia afflicta Gory & Percheron,1833   Cyprus 9Cetoniinae Pal.31 (1) as Potosia afflicta
Pal.33 (4) as Cetonia a.
Pal.35 (4)
Greece 1Cetoniinae Pal.33 (1) as Cetonia a.
Syrian Arab Republic 5Cetoniinae Pal.14 (5)
Turkey 2Cetoniinae (2) Pal.14
Unknown 3Cetoniinae Pal.24 (3)
Protaetia afghana (Petrovitz,1955)   Afghanistan 1Cetoniinae Pal.10 (1)
Protaetia agglomerata (Solsky,1876)   Turkestan, Kazakhstan / Uzbekistan / Kyrgyzstan / Tajikistan / Turkmenistan / Uzbekistan 2Cetoniinae (2) Pal.15
Protaetia alboguttata (Vigors.,1826)   India 4Cetoniinae Or.3 (3)
Or.7 (1)
Unknown 1Cetoniinae Or.5 (1)
Protaetia angustata (Germar,1817)   Austria 1Cetoniinae Pal.9 (1)
Croatia 1Cetoniinae Pal.33 (1) as Cetonia a.
Greece 9Cetoniinae (8) Pal.4+16
Pal.33 (1) as Cetonia a.
Israel 3Cetoniinae Pal.31 (3) as Netocia angustata
Italy 2Cetoniinae (2) Pal.11+16
Syria 3Cetoniinae (3) Pal.16
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 5Cetoniinae Pal.7 (5)
Unknown 4Cetoniinae Pal.16 (3)
Pal.22 (1)
Protaetia angustata angustula (Germar,1817)   Syria 11Cetoniinae (11) Pal.13 (as var. eibesensis Reitter)

Protaetia anovittata Chevrolat,1841   Unknown 2Cetoniinae (2) Or.5
Protaetia aurichalcea (Fabricius,1775)   India 3Cetoniinae (3) Or.3+5
Unknown 6Cetoniinae (6) Or.5
Protaetia aurichalcea madagassae (Fabricius,1775)   Unknown 1Cetoniinae Or.3 (1)
Protaetia bogdanoffi (Solsky,1876)   Afghanistan 1Cetoniinae Pal.10 (1)
Protaetia brevitarsis (Lewis,1879)   China 9Cetoniinae (9) Pal.17
China 1Cetoniinae Pal.12 (1) as ssp. fairmairei Kraatz
Protaetia bucharica Curti,1915   Afghanistan 1Cetoniinae Pal.10 (1)
Protaetia cretica (Kraatz,1880)   Greece 6Cetoniinae Pal.6 (2)
Pal.33 (4)
Protaetia cuprea (Fabricius,1775)   Denmark 1Cetoniinae (1)
Finland 1Cetoniinae (1)
Germany 1Cetoniinae Pal.2 (1)
Greece 14Cetoniinae Pal.31 (1) as Cetonia cuprea
Pal.3+7 (13)
Italy 7Cetoniinae Pal.11 (7)
Norway 1Cetoniinae (1)
Portugal 5Cetoniinae Pal.7 (5)
Sweden 594   Sk Bl Ha Sm Öl Go GS Ög Vg Bo Ds Nä Sö Up Vs Vr Dr Gä Hs Me Hr Jä Ån Vb Nb Ås Ly Pi Lu To
Alan Dufberg 48 (6) as Potosia c.
Dufberg 49 (3)
Cetoniinae Main (585)
Turkey 11Cetoniinae Pal.29 (8)
Pal.31 (3) as Netocia cuprea
Protaetia cuprea daurica (Motschulsky,1860)   Russian Federation 1Cetoniinae Pal.20 (1)
Protaetia cuprea hypocrita (Ragusa,1905)   Greece 1Cetoniinae Pal.33 (1) as Cetonia c.
Protaetia cuprea ignicollis (Gory & Pecheron,1833)   Greece 1Cetoniinae (1) Pal.13
Syria 1Cetoniinae (1) Pal.13
Protaetia cuprea metallica (Herbst,1782)   Germany 2Cetoniinae Pal.20 (2) as Protaetia metallica (Herbst
1782)
Italy 1Cetoniinae Pal.20 (1) as Protaetia metallica (Herbst
1782)
Russian Federation 1Cetoniinae Pal.13 (1) as Protaetia metallica (Herbst
1782)
Russian Federation 3Cetoniinae Pal.32 (3)
Protaetia cuprea obscura (Andersch,1797)   Austria 1Cetoniinae (1) Pal.14
Italy 3Cetoniinae (3) Pal.14
Protaetia cuprea phoebe (Reitter,1899)   Syria 18Cetoniinae (18) Pal.13
Protaetia cupreola Kraatz,1899   Russian Federation 1Cetoniinae Pal.27 (1)
Protaetia cuprina (Motschulsky,1849)   Cyprus 2Cetoniinae Pal.31 (2)
Turkey 11Cetoniinae Pal.13 (7)
Pal.31 (1)
Pal.31 (3) as rhodensis Rataj
Protaetia cupripes Wiedemann,1821   Unknown 1Cetoniinae (1) Or.5
Protaetia cyanea (Kraatz,1886)   Uzbekistan 2Cetoniinae Pal.15 (2)
Protaetia excavata (Gory & Percheron,1833   Afghanistan 2Cetoniinae Pal.10 (2)
Protaetia funesta (Menetries,1836)   Israel 3Cetoniinae (3) Pal.14
Syria 13Cetoniinae (13) Pal.14
Protaetia fusca (Herbst,1790)   Kashmir: Pakistan / India / China 3Cetoniinae Or.3 (3)
Unknown 6Cetoniinae (6) Or.5
Protaetia impavida (Janson,1878)   Kashmir: Pakistan / India / China 2Cetoniinae Or.3 (2)
Protaetia incerta (Costa,1852)   Italy 1Cetoniinae (1) Pal.13 (as ssp. of cuprea)

Protaetia judith (Reiche,1871)   Greece 1Cetoniinae (1) Pal.13
Protaetia karelini (Zoubkov,1829)   Mongolia 2Cetoniinae (2) Or.5
Turkestan, Kazakhstan / Uzbekistan / Kyrgyzstan / Tajikistan / Turkmenistan / Uzbekistan 1Cetoniinae (1) Pal.14
Protaetia marmorata (Fabricius,1792)   Germany 1Cetoniinae Pal.34 (1)
Protaetia marmorata marmorata (Fabricius,1792)   Austria 1Cetoniinae Pal.17 (1) as P. lugubris
Hungary 1Cetoniinae Pal.12 (1) as P. lugubris
Sweden 115  Skåne Småland Öland Östergötland Västergötland Södermanland Uppland
Alan Dufberg 48 (1) as Liocola lugubris Hbst.
Cetoniinae Main (114)
Protaetia mayeti LeComte,1906   Israel 1Cetoniinae Pal.31 (1)
Protaetia mirifica mirifica (Reitter,1894)   Syria 2Cetoniinae (2) Pal.15 (as Protaetia koenigi Reitter)

Protaetia morio (Fabricius,1781)   Austria 1Cetoniinae Pal.9 (1)
France 7Cetoniinae (6) Pal.9+10
Pal.35 (1)
Italy 12Cetoniinae (12) Pal.9+15+24
Portugal 1Cetoniinae Pal.9 (1)
Spain 15Cetoniinae Pal.31 (2) as Netocia morio
Pal.9+15 (13)
Syria 1Cetoniinae (1) Pal.15
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 1Cetoniinae Pal.9 (1)
Unknown 2Cetoniinae Pal.24 (2)
Protaetia oblonga (Gory & Percheron, 1833)   Portugal 3Cetoniinae Pal.6 (3)
South Africa 1Cetoniinae (1) Afr.21 (as Cetonia o.)
Protaetia opaca (Fabricius,1787)   Algeria 6Cetoniinae Pal.14 (6)
France 5Cetoniinae Pal.9 (1)
Pal.31 (4) as Netocia opaca
Greece 8Cetoniinae Pal.17 (8) as Netocia opaca
Israel 1Cetoniinae Pal.31 (1)
Portugal 4Cetoniinae (4) Pal.7+9
Spain 1Cetoniinae Pal.14 (1)
Turkey 6Cetoniinae Pal.29 (3)
Pal.35 (3)
Protaetia opaca cretica (Kraatz,1880)   Greece 7Crete
Cetoniinae Pal.29 (2) as Netocia o.
Pal.31 (5) as Netocia o.o.
Protaetia orientalis (Gory & Percheron,1833)   China 5Cetoniinae (5) Pal.15
Japan 6Cetoniinae Pal.12 (6)
Protaetia orientalis submarmorea (Gory & Percheron,1833)   Japan 3Cetoniinae (3) Pal.16+22 (as Cetonia m.)
Unknown 1Cetoniinae (1) Pal.16
Protaetia palawanica Miksic,    Philippines 2Cetoniinae Or.3 (2)
Protaetia philippensis Fabricius, 1775   Philippines 1Cetoniinae (1) Or.5
Unknown 5Cetoniinae (5) Or.5
Protaetia proctotricha (Fischer von Waldheim,1842)   Kyrgyzstan 1Cetoniinae Pal.20 (1)
Protaetia prolongata Gory & Percheron,1833   Indonesia 6Cetoniinae (6) Or.3+5
Protaetia pryeri oschimana (Nonfried,1895)   Japan 2Cetoniinae (2) Pal.17
Protaetia sardea (Gory & Percheron,1833)   France 1Corsica
Cetoniinae (1) Pal.
Italy 1Sardinia
Cetoniinae (1) Pal.14
Protaetia scepsia Dohrn,1872   Unknown 1Cetoniinae (1) Or.5
Protaetia sibirica Gebler,1829   Unknown 1Cetoniinae Pal.24 (1)
Protaetia solorensis sumbavana Kraatz,1900   Indonesia 9Sumatra
Cetoniinae Or.3 (9)
Protaetia speciosa jousselini (Gory & Percheron,1833)   Syria 2Cetoniinae (2) Pal.16
Protaetia speculifera (Swartz,1817)   China 3Cetoniinae (3) Pal.12
Protaetia taciturna Guérin-Méneville,1830   Indonesia 1New Guinea
Cetoniinae (1) Or.5
Unknown 2Cetoniinae (2) Or.5
Protaetia trojana (Gory & Percheron,1833)   Unknown 1Cetoniinae (1) Pal.14
Protaetia turkestanica (Kraatz,1886)   Afghanistan 1Cetoniinae (1) Pal.19
Protaetia ungarica (Herbst,1790)   Austria 1Cetoniinae Pal.9 (1)
Hungary 1Cetoniinae Pal.24 (1)
Israel 1Cetoniinae Pal.31 (1) as "Potosia hungarica Scop."
Unknown 8Cetoniinae Pal.24 (8)
Protaetia ungarica armeniaca (Menetries,1832)   Syria 1Cetoniinae Pal.27 (1)
Unknown 1Cetoniinae Pal.24 (1)
Protaetia ungarica auliensis (Reitter,1901)   Unknown 5Cetoniinae Pal.24 (5)
Protaetia vidua (Gory & Percheron,1833)   Greece 1Cetoniinae Pal.35 (1)
Syria 3Cetoniinae (3) Pal.15
Turkey 2Cetoniinae Pal.15 (2)
Psacadoptera leucomelona Gory & Percheron,1833   Namibia 1Cetoniinae Afr.5 (1)
South Africa 2Cetoniinae (2) Afr.21 (as Cetonia l.)
Psacadoptera simonsi Janson,1877   South Africa 1Cetoniinae (1) Afr.11
Psammodius asper (Fabricius,1775)   Denmark 1Aphodiinae Main (1)
France 1Aphodiinae Pal.16 (1)
Germany 1Aphodiinae Pal.15 (1)
Romania 3Aphodiinae Pal.7+11 (3)
Sweden 186   Skåne Halland Gotland Uppland Värmland
Alan Dufberg 48 (11)
Aphodiinae Main (175)
Psammodius basalis (Mulsant & Rey,1871)   Greece 1Aphodiinae (1) Pal.12
Psammodius laevipennis Costa,1844   Greece 1Aphodiinae Pal.30 (1)
Italy 1Aphodiinae Pal.12 (1)
Turkey 20Aphodiinae Pal.12 (10)
Pal.26 (9)
Pal.30 (1)
Psammodius pierottii Pittino,1979   Italy 3Aphodiinae (3) Pal.11+12
Psammoporus criddlei (Brown,1931)   Canada 1British Columbia
Aphodiinae N.Am.3 (1)
United States 6Alaska
Aphodiinae N.Am.4 (6)
Psammoporus cylindrica (Eschscholtz,1822)   Canada 9British Columbia
Aphodiinae N.Am.4 (1)
N.Am.7 (8)
Canada 34Yukon
Aphodiinae N.Am.7 (34)
Psammoporus lacustris (LeConte,1850)   Canada 7Ontario
Aphodiinae N.Am.3 (7)
Canada 6British Columbia
Aphodiinae (6) N.Am.3+4
United States 3Alaska
Aphodiinae N.Am.3 (3)
United States 1Colorado
Aphodiinae N.Am.3 (1)
Psammoporus opaca (Brown,1931)   Canada 4British Columbia
Aphodiinae (4) N.Am.3+4
Psammoporus sabuleti (Panzer,1797)   Norway 16Aphodiinae Main (16)
Sweden 293  Sk Ha Sm Ög Vg Vr Dr Me Hr Jä Ån Vb Nb Ly Lu To
Alan Dufberg 47 (14)
Aphodiinae Main (279)
Psammoporus terminalis (Brown,1931)   Canada 1British Columbia
Aphodiinae N.Am.4 (1)
Canada 31Yukon
Aphodiinae N.Am.4 (19)
N.Am.7 (2)
N.Am.8 (10)
United States 1Alaska
Aphodiinae N.Am.4 (1)
Pseudacrossus balangensis Petrovitz,1958   China 1Aphodiinae Pal.13 (1) incl. 1 paratype
Pseudacrossus grombczewskyi Koshantschikov,1891   Kazakhstan 1Aphodiinae Pal.28 (1)
Kyrgyzstan 1Aphodiinae Pal.17 (1)
Russian Federation 3Aphodiinae Pal.21 (3)
Pseudadoretus phthisicus (Dohrn,1882)   Russian Federation 4Rutelinae Pal.9 (4)
Pseudesymus lucidus (Klug,1845)   Egypt 14Aphodiinae Afr.9 (14)
Spain 3Canary Islands
Aphodiinae Pal.27 (2)
Pal.28 (1)
Pseudinca admixtus Hope,1842   Cameroon 1Cetoniinae Afr.28 (1)
Pseudocanthon spp.    Peru 1Scarabaeinae S.Am.2 (1)
Pseudoclinteria cincticollis Burmeister,1847   South Africa 1Cetoniinae Afr.9 (1) as Clinteria c.
Pseudoclinteria infuscata (Gory & Percheron,1833)   Zambia 1Cetoniinae (1) Afr.7 (as Clinteria i.)

Pseudodrepanocanthus nasutus (Harold,1869)   South Africa 7Aphodiinae Afr.32 (7)
Pseudoheterochelus wallekraalensis Schein,1959   South Africa 2Hopliini Afr.7 (2)
Pseudohomonyx javanus (Burmeister,1847)   Indonesia 1Java
Dynastinae Or.9 (1)
Pseudomacraspis affinis (Laporte,1840))   Ecuador 2Rutelinae (2) S.Am.4
Pseudopharaphodius anthrax (Gerstaecker,1871)   Ghana 1Aphodiinae Afr.18 (1) as Aphodius
Kenya 1Aphodiinae Afr.3 (1) as Aphodius
Pseudopharaphodius anthrax Gerstaecker,1871   Nigeria 2Aphodiinae Afr.28 (2) as Aphodius
Pseudopharaphodius apicesetosus (Clement,1969)   Gambia 18Aphodiinae Afr.2 (18) as Aphodius
Senegal 4Aphodiinae Afr.2 (4) as Aphodius
Sierra Leone 11Aphodiinae Afr.15 (11) as Aphodius
Pseudopharaphodius kaszabi (Endrödi,1957)   South Africa 1Aphodiinae Afr.32 (1)
Pseudopharaphodius tumulosus (Schmidt,1910)   Kenya 2Aphodiinae Afr.35 (2)
Pseudoprotaetia stolata Olivier,1789   Senegal 1Cetoniinae (1) Afr.26
Unknown 1Cetoniinae (1) Afr.26
Pseudoptilophorus plagosus (Boheman,1857)   South Africa 2Cetoniinae Afr.17 (2)
Pseudosaproecius spp.    Guinea-Bissau 1Scarabaeinae Afr.5 (1)
Sierra Leone 1Scarabaeinae Afr.5 (1)
Pseudosinghala dalmannii (Gyllenhal,1817)   China 3Rutelinae Pal.6 (3)
Pseudosinghala transversa (Burmeister,1855)   China 3Rutelinae Pal.6 (3)
Pseudosinghala vorstmanni Hell,1891   Borneo, Indonesia / Malaysia 1Rutelinae Or.8 (1)
Pseudostereomera mirabilis Bordat & Howden,1995   Malaysia 1Type Coleoptera 3036 (1) Holotype 03042:1 of Pseudostereomera mirabilis Bordat & Howden 1995
  Pseudostereomera mirabilis Bordat & Howden 1995

Pseudosyrichthus senegalensis (Castelnau,1840)   Guinea-Bissau 1Dynastinae (1) Afr.16
Tanzania 1Dynastinae Afr.9 (1)
Pseudoteuchestes schirparakensis (Petrovitz.1955)   Afghanistan 13Aphodiinae Pal.8 (6) as Aphodius sch.
Pal.8 (7) as Aphodius sch. ab. postpositus Balt.
Pseudotorynorrhina japonica (Hope,1841)   China 2Cetoniinae (2) Pal.12
Pseudotrematodes frivaldszkyi (Menetries,1836)   Greece 2Melolonthinae Pal.3 (1)
Pal.13 (1)
Turkey 4Melolonthinae Pal.14 (1)
Pal.3+12 (3)
Unknown 1Melolonthinae Pal.11 (1)
Pseudoxyomus geminata (Schmidt,1911)   South Africa 2Aphodiinae Afr.32 (2)
Pseudoxyomus rubescens Petrovitz,1962   South Africa 1Aphodiinae Afr.32 (1)
Ptychodesthes gratiosa Ancey,1881   Tanzania 1Cetoniinae (1) Afr.22
Pychnoschema antiorii (Gestro,1881)   Ethiopia 2Dynastinae (2) Afr.16
Pychnoschema corydon (Olivier,1789)   South Africa 1Dynastinae (1) Afr.12
Pychnoschema diversum cavifrons Lesne,1906   Ethiopia 17Dynastinae (17) Afr.15
Pychnoschema diversum diversum Lesne,1906   Ethiopia 1Dynastinae (1) Afr.15
Pychnoschema subulatum Quedenfeldt, 1884   Cameroon 1Dynastinae Afr.7 (1)
Pygora ignita Westwood,1879   Madagascar 1Cetoniinae (1) Afr.18
Raceloma jansoni Distant,1897   Lesotho 2Cetoniinae Afr.9 (2)
Ranzania bertolonii ?   Unknown 1Cetoniinae (1) Afr.22
Rectoscutaria peringueyi Schein,1958   South Africa 2Hopliini (2) Afr.12
Repsimus aeneus (Fabricius,1775)   Australia 2Rutelinae (2) Pac.1
Unknown 2Rutelinae (2) Pac.1
Repsimus manicatus (Swartz,1817)   Australia 3Rutelinae (3) Pac.1
Rhabdotis aulica (Fabricius,1781)   Lesotho 2Cetoniinae Afr.11 (2)
South Africa 15Cetoniinae (15) Afr.11+21 (as Cetonia a.)
Tanzania 1Cetoniinae (1) Afr.24
Unknown 4Cetoniinae (4) Afr.24
Rhabdotis chalcea transvaalica Peringuey,   South Africa 4Cetoniinae Afr.11 (4)
Rhabdotis semipunctata (Fabricius,1787)   Namibia 12Cetoniinae (12) Afr.5+11+25
Unknown 1Cetoniinae (1) Afr.25
Rhabdotis sobrina (Gory & Percheron,1833)   Central African Republic 1Cetoniinae Afr.25 (1)
Ethiopia 1Cetoniinae Afr.25 (1)
Tanzania 3Cetoniinae Afr.25 (2)
Afr.32 (1)
Uganda 1Cetoniinae Afr.25 (1)
Unknown 8Cetoniinae Afr.25 (8)
Rhabdotis sobrina virginea Klug,1855   South Africa 2Cetoniinae Afr.11 (2)
Tanzania 1Cetoniinae Afr.32 (1)
Rhabdotis spp.    Unknown 1Africa
Cetoniinae (1) Afr.22
Rhinocoeta cornuta (Fabricius,1781)   South Africa 1Cetoniinae Afr.9 (1)
Rhinyptia spp.    Aschantes 1Rutelinae Pal.8 (1)
Rhizoplatys auriculatus (Burmeister,1847)   Namibia 1Dynastinae (1) Afr.15
Rhizoplatys bituberculatus Klug,1855   Namibia 1Dynastinae (1) Afr.15
South Africa 1Dynastinae (1) Afr.15
Rhizoplatys mucronatus (Beauvois,1905)   Cameroon 1Dynastinae Afr.9 (1)
Ghana 1Dynastinae Afr.9 (1)
Rhizoplatys mucronatus trituberculatus (Beauvois,1905)   South Africa 1Dynastinae Afr.9 (1)
Rhizotrogus aestivus (Olivier,1789)   France 3Melolonthinae (3) Pal.4+9
Germany 1Melolonthinae Pal.9 (1)
Rhizotrogus cicatricosus Mulsant,1842   France 2Melolonthinae Pal.4 (2)
Rhizotrogus genei Blanchard,1850   France 1Melolonthinae Pal.5 (1)
Rhizotrogus insubricus Burmeister,1855   Italy 2Melolonthinae Pal.4 (2)
Rhizotrogus latiusculus Schaufuss,1864   Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Melolonthinae Pal.5 (2)
Rhizotrogus maculicollis Villa & Villa,1833   France 2Melolonthinae Pal.5 (2)
Rhizotrogus marginipes Mulsant,1842   France 2Melolonthinae Pal.4 (2)
Spain 2Melolonthinae Pal.4 (2)
Rhizotrogus mascarauxi Desbroches,1896   France 1Melolonthinae Pal.4 (1)
Rhizotrogus pallidipennis Blanchard,1851   Spain 2Melolonthinae Pal.4 (2)
Rhizotrogus punicus Burmeister,1855   Algeria 5Melolonthinae Pal.4 (5)
Rhizotrogus subemarginatus Reiche,1862   Spain 1Melolonthinae Pal.5 (1)
Rhizotrogus vicinus Mulsant,1842   France 1Melolonthinae Pal.4 (1)
Rhomborrhina mellyi knirschi Schürhoff,1933   China 1Cetoniinae (1) Pal.12
Rhomborrhina unicolor Motschulsky,1862   Japan 5Cetoniinae (5) Pal.12
Rhyparus adebratti Bordat,1996   Malaysia 24Type Coleoptera 3036 (24) Holotype Paratype 03036:1-24 of Rhyparus adebratti Bordat 1996
  Rhyparus adebratti Bordat 1996

Rhyparus approximatus Fairmaire,1893   Borneo, Indonesia / Malaysia 117Aphodiinae (117) Or.4+5
Rhyparus burckhardti Paulian,1989   Borneo, Indonesia / Malaysia 19Aphodiinae Or.4 (19)
Rhyparus comorianus Fairmaire,1896   Benin 1Aphodiinae (1) Afr.24
Rhyparus danielssoni Bordat,1996   Borneo, Indonesia / Malaysia 7Type Coleoptera 3036 (7) Holotype Paratype 03037:1-7 of Rhyparus danielssoni Bordat 1996
  Rhyparus danielssoni Bordat 1996

Rhyparus denticollis Fairmaire,1893   Borneo, Indonesia / Malaysia 14Aphodiinae Or.5 (14)
Rhyparus desjardinsi Westwood,1845   Reunion 1Aphodiinae (1) Afr.24
Rhyparus helophoroides Fairmaire,1893   Borneo, Indonesia / Malaysia 8Aphodiinae (8) Or.5+6
Rhyparus magnus Schmidt,1911   Borneo, Indonesia / Malaysia 10Aphodiinae Or.6 (10)
Rhyparus micros Bordat,1996   Malaysia 16Type Coleoptera 3036 (16) Holotype Paratype 03038:1-16 of Rhyparus micros Bordat 1996
  Rhyparus micros Bordat 1996

Rhyparus n.sp.    Borneo, Indonesia / Malaysia 1Aphodiinae Or.6 (1)
Rhyparus occidentalis Paulian,1990   Senegal 1Aphodiinae (1) Afr.13
Rhyparus pseudominor Bordat,1996   Malaysia 8Type Coleoptera 3036 (8) Holotype Paratype 03039:1-8 of Rhyparus pseudominor Bordat 1996
  Rhyparus pseudominor Bordat 1996

Rhyparus spp.    Malaysia 5Borneo
Aphodiinae Or.6 (5)
Mexico 1Aphodiinae N.Am.3 (1)
Rhyparus suturalis Schmidt,1911   Kenya 1Aphodiinae (1) Afr.24
Rhyparus xanthi (Frivaldszky,1883)   Borneo, Indonesia / Malaysia 16Aphodiinae Or.4 (16)
Rhyssemodes orientalis (Mulsant & Godart,1874),   Afghanistan 107Aphodiinae Pal.9 (107)
Egypt 1Aphodiinae (1) Afr.12
Kyrgyzstan 9Aphodiinae (9) Pal.16
Tunisia 1Aphodiinae (1) Pal.12
Turkey 1Aphodiinae (1) Pal.12
Rhyssemodes sindicus Pittino,1984   Thailand 17Aphodiinae Or.2 (17)
Rhyssemus algiricus Lucas,1846   Spain 7Aphodiinae (7) Pal.12
Rhyssemus annaedicatus Pierotti,1980   Italy 2Aphodiinae (2) Pal.12
Rhyssemus archambaulti Benard,1929   Gambia 176Aphodiinae (176) Afr.10+11
Guinea-Bissau 1Aphodiinae Afr.14 (1)
Senegal 16Aphodiinae (16) Afr.11
Rhyssemus aspericollis Petrovitz,1965   Kenya 7Aphodiinae (7) Afr.12
Rhyssemus atramentarius Peringuey,1903   South Africa 2Aphodiinae Afr.16 (2)
Zimbabwe 2Aphodiinae Afr.16 (2)
Rhyssemus aurivilliusi Clouet,1901   Congo 1Aphodiinae (1 ) Afr.25
Rhyssemus bechuanus  Petrovitz,1956   Botswana 2Aphodiinae (2) Afr.16
Rhyssemus brevitarsis Pittino,1910   Oman 2Aphodiinae (2) Pal.7.
Rhyssemus carinatipennis Peringuey,1901   Congo 3Aphodiinae (3) Afr.25
Kenya 2Aphodiinae (2) Afr.12
Sierra Leone 2Aphodiinae (2) Afr.14
Rhyssemus congolanus Clouet,1901   Congo 4Aphodiinae (4) Afr.25
Rhyssemus germanus (Linnaeus,1767)   Afghanistan 12Aphodiinae Pal.9 (12)
Austria 1Aphodiinae (1) Pal.13
Germany 1Aphodiinae (1) Pal.15
Ghana 2Aphodiinae Afr.17 (2)
Greece 1Rhodos
Aphodiinae Pal.30 (1)
Hungary 1Aphodiinae (1) Pal.16
Italy 24Aphodiinae (24) Pal.3+12+15
Mozambique 1Aphodiinae Afr.35 (1)
Spain 1Aphodiinae Pal.16 (1)
Sweden 1  Skåne
Aphodiinae Main (1)
Turkey 3Aphodiinae Pal.30 (3)
Rhyssemus granosus (Klug,1842),   Congo 3Aphodiinae Afr.26 (3 )
Gambia 22Aphodiinae (22) Afr.11+12
Rhyssemus hauseri Balthasar,1933   Kenya 5Aphodiinae (5) Afr.12
Rhyssemus inscitus (Walker,1858)   Sri Lanka 22Aphodiinae Or.2 (22)
Rhyssemus keisseri Benard,1910   Gambia 4Aphodiinae (4) Afr.12
Rhyssemus laesifrons Fairmaire,1892   Ethiopia 1Aphodiinae (1) Afr.20
Rhyssemus limbolarius Petrovitz,1963   Italy 1Aphodiinae (1) Pal.13
Rhyssemus macedonicus Bénard,1923   Greece 16Aphodiinae Pal.12 (16)
Rhyssemus mayeti Clouet,1901   Congo 1Aphodiinae Afr.26 (1 )
Egypt 1Aphodiinae (1) Afr.12
Rhyssemus mimus Balthasar,1961   Namibia 2Aphodiinae (2) Afr.169
Rhyssemus mourgliai in lit.   Sierra Leone 17Aphodiinae Afr.14 (17)
Rhyssemus parallelus Reitter,1892   Spain 7Aphodiinae (7) Pal.12
Rhyssemus pectoralis Clouet,1901   Congo 3Aphodiinae Afr.25 (3)
Rhyssemus plicatus (Germar,1817)   Greece 17Aphodiinae Pal.12 (17)
Italy 2Aphodiinae (2) Pal.3 (as R.arenarius Costa)

Rhyssemus propinquus Peringuey,1964   Gambia 7Aphodiinae (7) Afr.12
Rhyssemus puncticollis Brown,1929   Russian Federation 1Kurgan oblast
Aphodiinae Pal.24 (1)
Rhyssemus reitteri Clouet,1901   Congo 3Aphodiinae (3 ) Afr.26
Rhyssemus ritsemae Clouet,1901   Gambia 3Aphodiinae (3) Afr.12
Senegal 14Aphodiinae (14) Afr.12
Sierra Leone 62Aphodiinae (62) Afr.15
Rhyssemus sculptipennis Schmidt,1922   Kenya 11Aphodiinae (11) Afr.12
South Africa 1Aphodiinae (1) Afr.16
Rhyssemus senegalensis Petrovitz,1972   Gambia 3Aphodiinae (3) Afr.11
Guinea-Bissau 2Aphodiinae (2) Afr.14
Nigeria 1Aphodiinae (1) Afr.20
Senegal 1Aphodiinae (1) Afr.20
Rhyssemus severini Clouet,1901   Congo 3Aphodiinae Afr.25 (3)
Rhyssemus sexcostatus Schmidt,1909   Congo 4Aphodiinae (4 ) Afr.26
Gambia 1Aphodiinae (1) Afr.11
Senegal 42Aphodiinae (42) Afr.11
Rhyssemus similis Petrovitz,1963   Sierra Leone 1Aphodiinae (1) Afr.14
Rhyssemus spp.    Botswana 1Aphodiinae Afr.16 (1)
Greece 1Aphodiinae Pal.30 (1)
Indonesia 2Aphodiinae Or.7 (2)
Kenya 27Aphodiinae Afr.16 (27)
Madagascar 1Aphodiinae Afr.30 (1)
Oman 1Aphodiinae Pal.13 (1)
Sierra Leone 6Aphodiinae Afr.16 (6)
South Africa 2Aphodiinae Afr.16 (1)
Afr.34 (1)
Spain 1Aphodiinae Pal.13 (1)
Turkey 5Aphodiinae Pal.26 (5)
Rhyssemus sulcatus (Olivier,1789)   Spain 41Aphodiinae Pal.12 (41)
Rhyssemus verrucosus Mulsant,1842   Spain 3Aphodiinae (3) Pal.12+13
Rhyssemus zumpti Petrovitz,1956   Botswana 1Aphodiinae (1) Afr.16 (incl. 1 paratype)

Rutela coerulea Perty,1832   Unknown 1Rutelinae S.Am.3 (1)
Rutela formosa Burmeister,1844   Cuba 2Rutelinae S.Am.3 (2)
Rutela histrio Sahlberg,1823   Argentina 1Rutelinae S.Am.3 (1)
Brazil 5Rutelinae S.Am.3 (5)
Unknown 1Rutelinae S.Am.3 (1)
Rutela laeta (Weber,1801)   Unknown 1Rutelinae (1) S.Am.4
Rutela lineola (Linnaeus,1767)   Brazil 8Rutelinae S.Am.3 (8)
Trinidad and Tobago 1Trinidad
Rutelinae S.Am.3 (1)
Unknown 5Rutelinae S.Am.3 (5)
Rutela striata (Olivier,1789)   Guadeloupe 1Rutelinae S.Am.3 (1)
Ruteloryctes morio (Fabricius,1798)   Cameroon 1Dynastinae Afr.9 (1)
Guinea-Bissau 1Dynastinae Afr.16 (1)
Saprosites basilewski Endrödi,1955   Burundi 1Aphodiinae Afr.26 (1 ) Paratype
Rwanda 1Aphodiinae Afr.26 (1 ) Paratype
Saprosites burgeoni Paulian,1940   Congo 2Aphodiinae Afr.26 (2 ) 2 syntypes
Saprosites calvus Schmidt,1911   Ghana 11Aphodiinae (11) Afr.16
Kenya 2Aphodiinae (2) Afr.12
Saprosites camerounensis Paulian,1940   Cameroon 1Aphodiinae (1) 1 syntype Afr.20
Saprosites carinatus Bordat,1992   Kenya 1Aphodiinae Afr.12 (1)
Saprosites cf. rufopolitus Gillet,1924   Malaysia 1Borneo
Aphodiinae Or.7 (1)
Saprosites consonus Schmidt,1911   Ghana 1Aphodiinae (1) Afr.17
Sierra Leone 1Aphodiinae (1) Afr.14
Saprosites falcatus Schmidt,1908   Madagascar 1Aphodiinae (1) Afr.24
Saprosites laticeps Fairmaire,1871   Madagascar 1Aphodiinae (1) Afr.24
Seychelles 1Aphodiinae Or.6 (1)
Saprosites lepersonnei Paulian,1942   Kenya 3Aphodiinae (3) Afr.12
Saprosites lodoicea Scott,1912   Seychelles 1Aphodiinae Or.6 (1) Paratype
Saprosites malaisei Paulian,1944   Borneo, Indonesia / Malaysia 10Aphodiinae Or.6 (10)
Saprosites mendax Blackburn,1892   United Kingdom 1England
Aphodiinae (1) Pal.15
Saprosites monteverdeae Stebnicka,2001   Costa Rica 1Aphodiinae S.Am.2 (1)
Saprosites palmarum (Scott,1912)   Seychelles 1Aphodiinae Or.6 (1)
Saprosites perbrevitarsis (Schmidt,1909)   Gabon 1Aphodiinae (1) Afr.24
Saprosites perssoni Bordat,1996   Ethiopia 6Type Coleoptera 4759 (6) Holotype Paratype 04762:1-6 of Saprosites perssoni Bordat 1996
  Saprosites perssoni Bordat 1996

Saprosites schoutedeni Paulian,1939   Congo 2Aphodiinae (2 ) Afr.20
Saprosites spp.    Sierra Leone 2Aphodiinae Afr.16 (2)
South Africa 1Aphodiinae Afr.34 (1)
Saprosites vosseleri Petrovitz,1965   Kenya 2Aphodiinae (2) Afr.13
Sarophorus costatus (Fåhraeus,1857)   South Africa 9Scarabaeinae Afr.33 (9)
Sarophorus tuberculatus (Castelnau,1840)   South Africa 2Scarabaeinae Afr.47 (2)
Scapanes australis (Boisduval,1832)   Unknown 1Dynastinae Pac.1 (1)
Scapanes australis solomonensis (Boisduval,1832)   Kiribati 1Dynastinae Pac.1 (1)
Scapanes spp.    Unknown 1Dynastinae Pac.1 (1)
Scaphorhinadoretus bimaculatus Ohaus,1912   Japan 2Rutelinae Pal.9 (2)
Scaptobius capensis (Gory-Percheron,1833)   South Africa 7Cetoniinae (6) Afr.17+27
Melolonthinae Afr.3 (1) (probably missid.)
Scaptobius natalensis Boheman,1857   South Africa 1Cetoniinae (1) Afr.17
Scarabaeus acuticollis (Motschulsky,1849)   Afghanistan 14Scarabaeinae (14) Pal.1 ( as S. sacer ssp. a.)
Uzbekistan 2Scarabaeinae (2) Pal.19 ( as S. sacer ssp. a.)
Scarabaeus aegyptiorum (Latreille,1827)   Turkey 1Scarabaeinae (1) Pal.33
Scarabaeus aegyptiorum purpurascens Gerstaecker,1871   Kenya 3Scarabaeinae Afr.50 (3)
Unknown 1Scarabaeinae Afr.50 (1)
Scarabaeus aff. puncticollis (Latreille,1819)   Unknown 1Scarabaeinae Pal.40 (1)
Scarabaeus aff. sennaariensis (Castelnau,1840)   Turkey 1Scarabaeinae Pal.40 (1)
Scarabaeus ambiguus (Boheman,1857)   South Africa 1Scarabaeinae (1) Afr.23
Scarabaeus ambiguus deceptor Peringuey,1901   Lesotho 1Scarabaeinae (1) Afr.23
Scarabaeus andrewesi Felsche,1907   Afghanistan 1Scarabaeinae Pal.2 (1)
Scarabaeus armeniacus (Menetries,1832)   Armenia 1Scarabaeinae Pal.46 (1)
Iran 1Scarabaeinae (1) Pal.35
Scarabaeus babori Balthasar,1934   Afghanistan 1Scarabaeinae (1) Pal.31
Scarabaeus brahminus (Laporte 1840)   Unknown 2Scarabaeinae Or.3 (2)
Scarabaeus caffer (Boheman,1857)   South Africa 1Scarabaeinae (1) Afr.23
Swaziland 2Scarabaeinae Afr.8 (2)
Scarabaeus carinatus (Gebler,1841)   Afghanistan 6Scarabaeinae (6) Pal.1 (as S. sacer ssp. c)

Scarabaeus catenatus (Gerstaecker,1871)   Kenya 1Scarabaeinae Afr.50 (1)
Scarabaeus cristatus Fabricius,1775   Unknown 2Scarabaeinae (2) Pal.33
Scarabaeus cupreus Castelnau,   Unknown 1Scarabaeinae Afr.45 (1)
Scarabaeus flavicornis (Boheman,1860)   South Africa 9Scarabaeinae (9) Afr.23
Scarabaeus funebris (Boheman,1857)   South Africa 6Scarabaeinae (6) Afr.24
Scarabaeus gangeticus Castelnau,   Indonesia 1Java
Scarabaeinae Or.3 (1)
Sri Lanka 3Scarabaeinae Or.3 (3)
Unknown 3Scarabaeinae Or.3 (3)
Scarabaeus goryi (Castelnau,1840)   Guinea 2Scarabaeinae Afr.50 (2)
Scarabaeus isidis    Tanzania 1Scarabaeinae (1) Afr.23
Scarabaeus laticollis Linnaeus,1767   France 3Scarabaeinae Pal.24 (1)
Pal.40 (2)
Morocco 2Scarabaeinae Pal.40 (2)
Spain 1Mallorca
Scarabaeinae Pal.38 (1)
Spain 11Scarabaeinae (11) Pal.20+31
Europa meridionalis 1Scarabaeinae Pal.40 (1)
Unknown 5Scarabaeinae Pal.40 (5)
Scarabaeus multidentatum (Klug,1845)   Egypt 1Scarabaeinae (1) Pal.33 (as Mnematium m.)

Scarabaeus pabulator Peringuey,1908   South Africa 1Scarabaeinae (1) Afr.24
Scarabaeus palemo Olivier,1789   Botswana 1Scarabaeinae Afr.50 (1)
Lesotho 1Scarabaeinae Afr.24 (1)
Namibia 5Scarabaeinae Afr.24 (5)
South Africa 2Scarabaeinae Afr.24 (2)
Scarabaeus parvulus Boheman,1860   South Africa 2Scarabaeinae (2) Afr.24
Scarabaeus pius Illiger,1803   Afghanistan 19Scarabaeinae Pal.2 (19)
Armenia 3Scarabaeinae Pal.46 (3)
Greece 1Scarabaeinae Pal.20 (1)
Italy 1Scarabaeinae Pal.34 (1)
Unknown 4Scarabaeinae Pal.24 (1)
Pal.40 (3)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Scarabaeinae Pal.20 (2)
Scarabaeus prodigiosus (Erichson,1843)   Kenya 1Scarabaeinae Afr.50 (1) Tungu
Tanzania 2Scarabaeinae Afr.50 (2)
Scarabaeus puncticollis (Latreille,1819)   Algeria 3Scarabaeinae Pal.20 (3)
Armenia 2Scarabaeinae Pal.46 (2)
Morocco 3Scarabaeinae Pal.20 (3)
Syria 1Scarabaeinae Pal.40 (1)
Tunisia 1Scarabaeinae Pal.20 (1)
Unknown 2Scarabaeinae Pal.40 (2)
Scarabaeus pustulosus (Gerstaecker,1871)   Tanzania 1Scarabaeinae Afr.57 (1)
Africa orientalis 1Scarabaeinae Afr.52 (1)
Unknown 2Scarabaeinae Afr.52 (2)
Scarabaeus ritchiei MacLeay,1821   Unknown 3Scarabaeinae (3) Pal.33 (as Mnematium r.)

Scarabaeus rugosus Hausman,1807   South Africa 1Scarabaeinae (1) Afr.24
Scarabaeus rusticus (Boheman,1857)   South Africa 23Scarabaeinae (23) Afr.8+9+24
Scarabaeus sacer Linnaeus,1758   Algeria 2Scarabaeinae Pal.19 (2)
Egypt 3Scarabaeinae Pal.31 (2)
Pal.39 (1)
Hungary 1Scarabaeinae Pal.39 (1)
Italy 6Scarabaeinae (6) Pal.19+31
Spain 2Scarabaeinae Pal.19 (2)
Tunisia 9Scarabaeinae Pal. (8) Pal.19+27+31+46
Pal.35 (1)
Turkey 6Scarabaeinae Pal.19 (1)
Pal.39 (3)
Pal.49 (2)
Unknown 7Scarabaeinae Pal.24 (1)
Pal.35 (1)
Pal.39 (5)
Scarabaeus satyrus (Boheman,1860)   South Africa 5Scarabaeinae (5) Afr.24
Scarabaeus savignyi Macleay,1821   South Africa 2Scarabaeinae Afr.50 (2)
Scarabaeus semipunctatus Fabricius,1792   France 2Scarabaeinae Pal.35 (1)
Pal.39 (1)
Italy 1Sicily
Scarabaeinae Pal.39 (1)
Italy 22Scarabaeinae Pal.21 (14)
Pal.39 (8)
Spain 5Scarabaeinae (5) Pal.21
Tunisia 2Scarabaeinae (2) Pal.21
Unknown 7Scarabaeinae Pal.39 (7)
Scarabaeus spp.    Afghanistan 33Scarabaeinae (33) Pal.31+32+33
Ethiopia 4Scarabaeinae Afr.8 (4)
India 1Scarabaeinae Or.10 (1)
Indonesia 1Java
Scarabaeinae Or.10 (1)
Kenya 2Scarabaeinae Afr.18 (1)
Afr.57 (1)
Namibia 3Scarabaeinae (3) Afr.18
South Africa 2Scarabaeinae Afr.50 (1) as S. bonelli
Afr.51 (1)
Sri Lanka 2Scarabaeinae Or.10 (2)
Tanzania 1Zanzibar
Scarabaeinae Afr.51 (1)
Turkey 1Scarabaeinae Pal.46 (1)
Unknown 18Scarabaeinae Afr.51 (18)
Zimbabwe 1Scarabaeinae Afr.51 (1)
Scarabaeus suri Hausman,1807   South Africa 22Scarabaeinae (18) Afr.24+25
Afr.50 (4)
Scarabaeus transcaspicus Stolfa,1938   Afghanistan 2Scarabaeinae (2) Pal.1 (as S. sacer ssp
t.)

Uzbekistan 2Scarabaeinae (2) Pal.22 (as S. sacer ssp
t.)

Scarabaeus typhon (Fischer,1823)   Bulgaria 1Scarabaeinae Pal.21 (1)
Greece 1Scarabaeinae Pal.53 (1)
Israel 1Scarabaeinae Pal.51 (1) as S. affinis Sturm
1800
Italy 2Scarabaeinae Pal.20 (1) as S. affinis Brullé
Pal.21 (1)
Spain 1Scarabaeinae Pal.20 (1) as S. affinis Brullé
Tunisia 1Scarabaeinae Pal.51 (1) as S. affinis Sturm
1800
Turkey 3Scarabaeinae Pal.20 (2) as S. affinis Brullé
Pal.53 (1)
Turkmenistan 1Scarabaeinae Pal.40 (1)
Scarabaeus variolosus Fabricius,1787   Algeria 12Scarabaeinae Pal.22 (4)
Pal.41 (8)
Croatia 1Scarabaeinae Pal.41 (1)
Greece 1Rhodos
Scarabaeinae Pal.41 (1)
Greece 5Scarabaeinae Pal.22 (1)
Pal.27 (1)
Pal.41 (3)
Greece 2Crete
Scarabaeinae Pal.49 (2)
Italy 1Sicily
Scarabaeinae Pal.41 (1)
Italy 13Scarabaeinae (13) Pal.22+23
Malta 1Scarabaeinae Pal.22 (1)
Morocco 3Scarabaeinae Pal.22 (3)
Spain 4Scarabaeinae Pal.22 (3)
Pal.41 (1)
Tunisia 1Scarabaeinae Pal.41 (1)
Illyria, Balkans 1Scarabaeinae Pal.41 (1)
Europa meridionalis 2Scarabaeinae Pal.41 (2)
Unknown 4Scarabaeinae Pal.35 (3)
Pal.41 (1)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Scarabaeinae Pal.22 (2)
Scarabaeus westwoodi (Harold,1869)   South Africa 2Scarabaeinae Afr.9 (2)
Scarabaeus wilsoni Waterhouse,1890   Turkey 2Scarabaeinae Pal.40 (2)
Unknown 1Scarabaeinae Pal.40 (1)
Scarabaeus winkleri Stolfa,1938   Afghanistan 1Scarabaeinae Pal.2 (1)
Sceliages adamastor Serville,1828   South Africa 2Scarabaeinae Afr.9 (2)
Sceliages curvipes Gillet,1911   South Africa 1Scarabaeinae (1) Afr.28
Sceliages hippias Westwood,1837   South Africa 3Scarabaeinae (3) Afr.29
Sceliages spp.    South Africa 3Scarabaeinae (3) Afr.18
Schizognathus spp.    Australia 1Rutelinae Pac.3 (1)
Schizonycha affinis Boheman,1857   South Africa 1Melolonthinae Afr.3 (1) (originaly as S. oblonga Boh.)
Schizonycha africana (Castelnau,1840)   Algeria 1Melolonthinae Afr.4 (1)
Gambia 9Melolonthinae Afr.1 (9)
Ghana 9Melolonthinae Afr.1 (9)
Senegal 4Melolonthinae Afr.1 (4)
Schizonycha algirina Fairmaire,1876   Tunisia 3Melolonthinae Pal.18 (3) as S. alegrinea
Schizonycha armipes Moser   Gambia 2Melolonthinae Afr.1 (2)
Senegal 14Melolonthinae Afr.1 (14)
Schizonycha aspera Brenske,1898   Cameroon 1Melolonthinae Afr.1 (1)
Schizonycha bottegoi Brenske,1895   Kenya 1Melolonthinae Afr.2 (1)
Schizonycha carbonaria Boheman,1847   South Africa 4Melolonthinae Afr.3 (4)
Schizonycha cf. fartula Per.   South Africa 1Melolonthinae Afr.2 (1)
Schizonycha crenata (Gyllenhal,1817)   Gambia 1Melolonthinae Afr.1 (1)
Senegal 1Melolonthinae Afr.1 (1)
Togo 1Melolonthinae Afr.4 (1)
Schizonycha damarina Péringuey,1908   Botswana 1Melolonthinae Afr.4 (1)
Guatemala 1Melolonthinae S.Am.1 (1)
Mexico 1Melolonthinae S.Am.1 (1)
Schizonycha flavicornis Brenske,1898   Eritrea 1Melolonthinae Afr.4 (1)
Sudan 1Melolonthinae Afr.2 (1)
Schizonycha insulicola Dalla Torre ,1912   Kenya 4Melolonthinae Afr.1 (4)
Schizonycha keniensis Moser   Kenya 2Melolonthinae Afr.1 (2)
Schizonycha propinqua Kolbe,1891   Tanzania 4Melolonthinae Afr.1 (4)
Schizonycha puncticollis Boheman,1857   Botswana 1Melolonthinae Afr.4 (1)
South Africa 2Melolonthinae Afr.4 (2)
Schizonycha rubricollis Moser ?   Tanzania 1Melolonthinae Afr.2 (1)
Schizonycha ruficollis (Fabricius,1781)   Sri Lanka 64Melolonthinae Or. (64) Or.3+4
Schizonycha rufina Boheman,1857   South Africa 1Melolonthinae Afr.3 (1)
Schizonycha salaama Brenske,1898   Mozambique 1Melolonthinae Afr.1 (1)
Schizonycha spp.    Cameroon 2Melolonthinae Afr.2 (2)
Eritrea 1Melolonthinae Afr.4 (1) as S. pilosa n.sp. Hultgren
Kenya 3Melolonthinae Afr.2 (3)
Sudan 1Melolonthinae Afr.2 (1)
Tanzania 2Melolonthinae Afr.2 (2)
Togo 1Melolonthinae Afr.4 (1) as S. pilosa n.sp. Hultgren
Tanganyika 4Melolonthinae Afr.2 (4)
Zambia 2Melolonthinae Afr.2 (2)
Schizonycha squamifera Wallengren,1881   South Africa 4Type Coleoptera 1650 (4) Syntype: 42 of Schizonycha squamifera Wallengren 1881
  Schizonycha squamifera Wallengren 1881

Schoenherria argus Burmeister,1855   Indonesia 1Java
Melolonthinae Or.10 (1)
Unknown 1Melolonthinae Pal.1 (1)
Schoenherria borneensis Brenske,1894   Borneo, Indonesia / Malaysia 1Melolonthinae Or.10 (1)
Scybalocanthon rugosus (Blanchard,1843)   Peru 1Scarabaeinae N.Am.1 (1)
Sebakue cf. urunguensis Decelle   Zambia 1Melolonthinae Afr.2 (1)
Sebakue coriacea Peringuey,1904   Zambia 2Melolonthinae Afr.2 (2)
Sebasteos spp.    Botswana 1Scarabaeinae Afr.52 (1)
South Africa 1Scarabaeinae Afr.52 (1)
Unknown 2Scarabaeinae Afr.52 (2)
Selaserica padukkana Ahrens in litt.,   Sri Lanka 1Sericini (1) 1 paratype Or.2
Semanopterus angustatus Blackburn,1888   Australia 2Dynastinae Pac.2 (2)
Semanopterus tricostatus Blackburn,1896   Australia 2Dynastinae Pac.2 (2)
Serica alternata LeConte,1856   United States 4Arizona
Sericini N.Am.1 (4)
Serica anthracina LeConte,1856   United States 1Nevada
Sericini N.Am.1 (1)
Serica atracapilla (Kirby,1837)   United States 4West Virginia
Sericini (4) N.Am.1 (as Serica cucullata Dawson)

Serica boops Waterhouse,1875   Japan 1Sericini Pal.2 (1)
Serica brevitarsis rectipes Nomura,1972   Japan 3Sericini Pal.1 (3)
Serica brunnea (Linnaeus,1758)   Estonia 1Sericini Pal.1 (1)
Finland 1Sericini (1)
Germany 1Sericini Pal.2 (1)
Norway 12Sericini (12)
Spain 1Sericini Pal.1 (1)
Sweden 436  Sk Bl Ha Sm Öl Ög Vg Bo Ds Sö Up Vs Vr Dr Gä Hs Hr Ån
Alan Dufberg 48 (10)
Sericini Main (426)
Switzerland 1Sericini Pal.1 (1)
United Kingdom 8Scotland
Sericini Pal.1 (8)
Serica campestris Dawson,1919   United States 3California
Sericini N.Am.1 (3)
Serica fimbriata LeConte,1856   United States 4California
Sericini N.Am.1 (4)
Serica fusa Brenske,1898   Sri Lanka 2Sericini Or.1 (2)
Serica iricolor (Say,1824)   United States 5Pennsylvania
Sericini N.Am.1 (5)
America borealis, United States/Canada 5Sericini N.Am.1 (5)
Serica lurida Brenske,1898   Sri Lanka 1Sericini Or.2 (1)
Serica mixta LeConte,1856   United States 4California
Sericini N.Am.1 (4)
Serica nepalensis (Frey,1969)   Sri Lanka 1Sericini Or.2 (1)
Serica nigroguttata Brenske,1897   China 1Sericini Pal.1 (1)
Serica sericea Illiger,1802   United States 1Connecticut
Sericini N.Am.1 (1)
United States 2New York
Sericini N.Am.1 (2)
United States 1Sericini N.Am.1 (1)
United States 1Washington
Sericini N.Am.1 (1)
Unknown 2Sericini N.Am.1 (2)
Serica setifera (Gyllenhal,1817)   Indonesia 1Java
Sericini Or.2 (1)
Unknown 1Sericini Or.2 (1)
Serica spp.    China 2Sericini Pal.2 (2)
Japan 1Sericini Pal.2 (1)
United States 2New York
Sericini N.Am.1 (2)
United States 1Virginia
Sericini N.Am.1 (1)
United States 1Pennsylvania
Sericini N.Am.1 (1)
United States 1Massachusetts
Sericini N.Am.1 (1)
United States 7California
Sericini N.Am.1 (7)
United States 3Washington
Sericini N.Am.1 (3)
United States 18Maryland
Sericini N.Am.1 (18)
United States 1Oregon
Sericini N.Am.1 (1)
United States 1South Carolina
Sericini N.Am.1 (1)
Unknown 2Sericini N.Am.1 (3)
Serica thibetana Brenske,1897   Nepal 1Sericini Or.2 (1)
Serica vespertina (Gyllenhal,1817)   United States 2New York
Sericini N.Am.1 (2)
Sericania fuscolineata Motschulsky,1860   Korea, Republic of 1Sericini Pal.1 (1)
Sericesthis pruinosa Dalman   Australia 1Melolonthinae Pac.1 (1)
Sericesthis spp.    Australia 1Western Australia
Melolonthinae Pac.1 (1)
Sericoides lucida (Curtis,1844)   Chile 1Sericini S.Am.1 (1)
Sericoides nitida Philippi & Philippi,1864   Chile 1Sericini S.Am.1 (1)
Sericoides viridis (Solier,1851)   Chile 5Sericini S.Am.1 (5)
Simogonius beccarii Harold,1871   Kenya 1Aphodiinae (1) Afr.13
Simogonius janssensi Paulian,1954   Congo, the democratic republic of the 1Aphodiinae (1) Afr.16 (incl. 1 paratype)

Sinapisoma minutum Boucomont,   Peru 1Scarabaeinae S.Am.2 (1)
Singhala tenella Blanchard,1850   Sri Lanka 76Rutelinae Or.6 (76)
Sisyphus bowringi White,1844   Unknown 3Scarabaeinae Or.2 (3)
Sisyphus crispatus Gory,1833   Gambia 7Scarabaeinae Afr.5 (7)
Lesotho 1Scarabaeinae Afr.31 (1)
Nigeria 1Scarabaeinae Afr.23 (1)
South Africa 10Scarabaeinae (10) Afr.23+31
Tanzania 3Scarabaeinae Afr.23 (3)
Sisyphus fasciculatus Boheman,1857   Swaziland 7Scarabaeinae Afr.5 (7)
Sisyphus goryi Harold,1859   Guinea-Bissau 1Scarabaeinae Afr.6 (1)
Swaziland 8Scarabaeinae Afr.6 (8)
Sisyphus hirtus Wiedemann,1823   Nepal 1Scarabaeinae Or.2 (1)
Sisyphus impressipennis Lansberge,1886   Lesotho 3Scarabaeinae Afr.32 (3) as S. transvaalensis Peringuey
1908
South Africa 24Scarabaeinae Afr.32 (24) as S. transvaalensis Peringuey
1908
Sisyphus indicus Hope,1831   Unknown 1Scarabaeinae Or.2 (1)
Sisyphus longipes (Olivier,1789)   Indonesia 1Java
Scarabaeinae Or.2 (1)
Sisyphus mexicanus Harold,1863   Guatemala 1Scarabaeinae S.Am.2 (1)
Sisyphus muricatus (Olivier,1789)   Unknown 1Scarabaeinae Afr.23 (1)
Sisyphus nanniscus Peringuey,1901   Cameroon 2Scarabaeinae Afr.23 (2) as S. seminulum Gerstaecker
1871
Guinea-Bissau 1Scarabaeinae Afr.5 (1) as S. seminulum Gerstaecker
1871
South Africa 15Scarabaeinae Afr.32 (15) as S. ocellatus nanniscus Per.
Sisyphus schaefferi (Linnaeus,1758)   Bulgaria 40Scarabaeinae (40) Pal.14+31
Cyprus 2Scarabaeinae Pal.14 (2)
Germany 4Scarabaeinae Pal.33 (4)
Greece 6Scarabaeinae Pal.14 (6)
Iran 1Scarabaeinae Pal.45 (1)
Israel 1Scarabaeinae Pal.33 (1)
Italy 24Scarabaeinae (24) Pal.14+24+33
Morocco 2Scarabaeinae Pal.33 (2)
Romania 13Scarabaeinae Pal.33 (13)
Unknown 7Scarabaeinae (7) Pal.33+34
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 10Scarabaeinae Pal.14 (10)
Sisyphus schaefferi (Linnaeus,1758) (Sisyphus)   Switzerland 1Scarabaeinae Pal.33 (1)
Turkey 29Scarabaeinae (7) Pal.14+27+33
Pal.45 (3)
Pal.47 (17)
Pal.53 (2)
Sisyphus schaefferi boschniaki Fischer von Waldheim,1823   Cyprus 3Scarabaeinae Pal.50 (3)
Sisyphus spp.    Gambia 9Scarabaeinae (9) Afr.21
Kenya 14Scarabaeinae (14) Afr.21
Malawi 1Scarabaeinae (1) Afr.20
South Africa 5Scarabaeinae (5) Afr.20
Swaziland 1Scarabaeinae Afr.5 (1)
Tanzania 1Scarabaeinae (1) Afr.20
Sisyraphora tomentosa (Gory-Percheron,1833)   South Africa 14Cetoniinae (14) Afr.15
Unknown 1Cetoniinae (1) Afr.26
Smaragdesthes africana (Drury,1773)   Congo, the democratic republic of the 1Cetoniinae (1) Afr.18
Unknown 1East Africa
Cetoniinae (1) Afr.22
Smaragdesthes spp.    Unknown 1East Africa
Cetoniinae (1) Afr.22
Sophrops abscessa     Laos 1Melolonthinae Or.10 (1)
Sophrops bituberculata    Laos 1Melolonthinae Or.10 (1)
Sophrops costatus (Walker,1859)   Sri Lanka 9Melolonthinae Or.6 (9)
Sophrops cotesi (Brenske,1892)   India 1Melolonthinae Or.10 (1)
Sophrops eurystomus (Burmeister,1855)   Sri Lanka 90Melolonthinae (90) Or.7+8
Sophrops planicollis (Burmeister,1855)   Laos 2Melolonthinae Or.8 (2)
Sophrops singhalensis (Brenske,1892)   Sri Lanka 1Melolonthinae Or.8 (1)
Sparrmannia flava Arrow,1917   South Africa 1Melolonthinae Afr.4 (1)
Unknown 1Melolonthinae Afr.4 (1)
Stalagmosoma albellum (Pallas,1781)   Afghanistan 1Cetoniinae (1) Pal.10 (as Stalagmopygus a.)

Stegopterus septus Burmeister,1840   South Africa 1Cetoniinae Afr.20 (1)
Stegopterus vittatus Fabricius,1781   South Africa 1Cetoniinae Afr.31 (1)
Unknown 4Cetoniinae Afr.31 (4)
Stenocrates bicarinatus Robinson,1947   Nicaragua 1Dynastinae (1) S.Am.22
Stenocrates carbunculus Perell   South Africa 1Dynastinae Afr.9 (1)
Stenocrates difficilis Endrödi,1966   Colombia 1Dynastinae S.Am.20 (1)
Peru 1Dynastinae (1) S.Am.31
Stenocrates laevicollis Kirsch,1870   Unknown 2Dynastinae S.Am.20 (2)
Stenocrates rufipennis (Fabricius,1801)   Brazil 1Dynastinae S.Am.20 (1)
Stephanopholis singhalensis Brenske,1896   Sri Lanka 7Melolonthinae Or.4 (7)
Stephanorrhina guttata (Olivier,1789)   Cameroon 1Cetoniinae Afr.32 (1)
Congo 1Cetoniinae Afr.18 (1)
Ghana 1Cetoniinae Afr.32 (1)
Unknown 2Cetoniinae Afr.22 (1)
Afr.32 (1)
Stethaspis suturalis (Fabricius,1775)   New Zealand 1Rutelinae Pac.3 (1)
Stichothyrea densata Kolbe,1914   Tanzania 4Cetoniinae Afr.6 (4)
Strandius obliquus (Olivier,1789)   Guinea-Bissau 1Scarabaeinae Afr.3 (1)
Strategus ajax (Olivier,1789)   Unknown 3Dynastinae S.Am.14 (3)
Strategus aloeus (Linnaeus,1758)   Ecuador 1Dynastinae (1) S.Am.11+33
Mexico 3Dynastinae S.Am.11 (3)
Peru 2Dynastinae (2) S.Am.11+31
United States 2Arizona
Dynastinae N.Am.6 (2)
Unknown 10Dynastinae (10) S.Am.10+11
Venezuela 1Dynastinae (1) S.Am.24
Strategus aloeus julianus (Linnaeus,1758)   Guatemala 1Dynastinae S.Am.11 (1)
Strategus jugurtha Burmeister,1847   United States 1Florida
Dynastinae N.Am.1 (1)
Strategus sarpedon (Burmeister,1847)   Cuba 1Dynastinae S.Am.11 (1)
Strategus simson (Linnaeus,1758)   Cuba 1Dynastinae S.Am.11 (1)
Unknown 1Dynastinae S.Am.11 (1)
Strategus spp.    Guatemala 1Dynastinae S.Am.30 (1)
Peru 1Dynastinae S.Am.36 (1)
Strategus surinamensis Burmeister,1847   Ecuador 1Dynastinae S.Am.11 (1)
Trinidad and Tobago 1Trinidad
Dynastinae S.Am.11 (1)
Strategus talpa (Fabricius,1792)   Unknown 1Dynastinae S.Am.1 (1)
Strategus validus (Fabricius,1775)   Brazil 26Dynastinae (26) S.Am.12+13+14
Cuba 1Dynastinae S.Am.12 (1)
Mexico 4Dynastinae N.Am.1 (4)
United States 1Texas
Dynastinae N.Am.1 (1)
Unknown 3Dynastinae (3) S.Am.12+13+14
Strigoderma arboricola (Fabricius,1792)   United States 1Texas
Rutelinae N.Am.2 (1)
Strigoderma maculata (Olivier, 1789)   Nicaragua 4Rutelinae S.Am.1 (4)
Strigoderma marginata (Olivier,1789)   Mexico 1Rutelinae S.Am.8 (1)
Strigoderma mexicana Blanchard,1851   Mexico 2Rutelinae S.Am.8 (2)
Unknown 1Rutelinae S.Am.8 (1)
Strigoderma pimalis Casey,1884   Colombia 1Rutelinae S.Am.8 (1)
United States 1Arizona
Rutelinae N.Am.2 (1)
Strigoderma spp.    Colombia 2Rutelinae S.Am.8 (2)
Strigoderma sulcipennis Burmeister,1844   Guatemala 1Rutelinae S.Am.1 (1)
Mexico 3Rutelinae N.Am.1 (3)
Stripsipher zebra Gory & Percheron,1833   South Africa 1Cetoniinae Afr.29 (1)
Subrinus sturmi Harold,1870   Afghanistan 1Aphodiinae Pal.14 (1)
Portugal 12Aphodiinae Pal.28 (12)
Sulcophanaeus chryseicollis (Harold,1863)   Honduras 1Scarabaeinae S.Am.2 (1)
Sulcophanaeus imperator (Chevrolat,1844)   Argentina 1Scarabaeinae S.Am.1 (1)
Sulcophanaeus menelas (Castelnau,1840)   Brazil 1Scarabaeinae S.Am.14 (1) Without proper locality
Sybax distortus Schmidt,1920   Kenya 1Aphodiinae Afr.16 (1)
Sybax impressicollis Boheman,1857   Uganda 1Aphodiinae Afr.30 (1)
Sybax sulcicollis Boheman,1857   Unknown 1Aphodiinae Afr.27 (1)
Synapsis tmolus (Fischer,1821)   Uzbekistan 3Scarabaeinae Pal.18 (1)
Pal.43 (2)
Synchilus gisleni Britton,1956   Australia 4Western Australia
Type Coleoptera (4) Holotype Paratype 01710:1-4 of Synchilus gisleni Britton 1955
  Synchilus gisleni Britton 1955

Syrichthodontus cribratus (Boheman,1857)   South Africa 1Dynastinae Afr.15 (1)
Syrichthodontus spurius (Burmeister,1847)   South Africa 1Dynastinae (1) Afr.15
Syrictes syrichthus (Fabricius,1775)   South Africa 13Dynastinae (13) Afr.9+14
Unknown 4Dynastinae Afr.9 (4)
Zambia 1Dynastinae (1) Afr.14
Taeniesthes specularis (Gerstaecker,1867)   Tanzania 1Cetoniinae (1) Afr.
Taeniodera anthracina (Gory & Percheron,1833)   Indonesia 3Java
Cetoniinae Or.3 (3)
Taeniodera haematica (Perty,1831)   Indonesia 7Java
Cetoniinae Or.3 (7)
Taeniodera quadrilineata (Hope,1831)   Sri Lanka 1Cetoniinae Or.2 (1)
Taurrhina longiceps Kolbe,1892   Central African Republic 1Cetoniinae (1) Afr.18
Temnorrhynchus baal Reiche & Saulcy,1856   Egypt 2Dynastinae Pal.3 (1)
Pal.6 (1)
Greece 1Dynastinae Pal.9 (1)
Spain 1Dynastinae Pal.5 (1)
Turkey 1Dynastinae Pal.5 (1)
Temnorrhynchus clypeatus Klug,1855   Kenya 1Dynastinae Afr.7 (1)
Zimbabwe 1Dynastinae Afr.7 (1)
Temnorrhynchus coronatus (Fabricius,1781)   Kenya 2Dynastinae Afr.19 (2)
South Africa 1Dynastinae Afr.15 (1)
Tanzania 2Dynastinae Afr.7 (2)
Unknown 1Dynastinae Afr.7 (1)
Zambia 1Dynastinae Afr.7 (1)
Temnorrhynchus coronatus natalensis (Fabricius,1781)   Namibia 1Dynastinae Afr.13 (1)
South Africa 19Dynastinae Afr.12+13+15 (19)
Tanzania 1Dynastinae Afr.7 (1)
Temnorrhynchus luna Fairmaire,1868   Sierra Leone 1Dynastinae (1) Afr.19
Temnorrhynchus retusus (Fabricius,1781)   South Africa 5Dynastinae (5) Afr.12
Temnorrhynchus sjöstedti Kolbe,1910   Tanzania 2Dynastinae (2) Afr.19
Temnorrhynchus spp.    Namibia 2
Namibia 2Dynastinae Afr.19 (2)
South Africa 1Dynastinae (1) Afr.15
Temnorrhynchus tridentatus Landsberge,1886   Gabon 1Dynastinae Afr.5 (1)
Tephraea dichroa Schaum,1844   Ethiopia 1Cetoniinae (1) Afr.26
South Africa 7Cetoniinae (7) Afr.14
Tephraea morosa Schaum,1848   Central African Republic 1Cetoniinae (1) Afr.26
South Africa 1Cetoniinae (1) Afr.14
Termitodius besucheti (Paulian,1983)   Borneo, Indonesia / Malaysia 1Aphodiinae Or.6 (1)
Termitodius charki Reyes-Castillo & Martinez,1979   Mexico 3Aphodiinae N.Am.3 (3)
Tesarius caelatus (LeConte,1857)   United Kingdom 4England
Aphodiinae (4) Pal.3+15
Tesarius oregonensis (Cartwright,1955)   United States 3Washington
Aphodiinae (3) N.Am.2 (as Psammodius o.)

Tesserodon novaehollandiae (Fabricius,1775)   Unknown 2Scarabaeinae Pac.1 (2)
Theodosia magnifica Rotsch., & Jorden,1893   Borneo, Indonesia / Malaysia 1Cetoniinae Or.2 (1)
Thyridium buchwaldi Ohaus,1912   Ecuador 1Rutelinae (1) S.Am 4
Thyridium flavipenne Burmeister,1844   Unknown 1Rutelinae (1) S.Am 4
Thyridium semicinctum Bates,1888   Ecuador 2Rutelinae (2) S.Am 4
Tibiodrepanus sinicus (Harold,1868)   Malaysia 8Scarabaeinae Or.13 (8)
Thailand 22Scarabaeinae Or.13 (22)
Tibiodrepanus sulcicollis (Castelnau,1840)   Gambia 1Scarabaeinae Afr.58 (1)
Zimbabwe 1Scarabaeinae Afr.58 (1)
Tiniocellus spinipes (Roth,1851)   Ethiopia 2Scarabaeinae Afr.1 (2)
Gambia 1Scarabaeinae Afr.1 (1)
Guinea-Bissau 1Scarabaeinae Afr.3 (1)
Kenya 1Scarabaeinae Afr.1 (1)
South Africa 1Scarabaeinae Afr.31 (1)
Tmesorrhina alpestris ssp.bafutensis Darge,1988   Cameroon 1Cetoniinae (1) Afr.18
Torynorrhina apicalis (Westwood,1842)   India 1Cetoniinae Or.1 (1)
Torynorrhina fulvopilosa (Moser,1911)   China 1Cetoniinae (1) Pal.12
Torynorrhina hyacinthina (Hope,1841)   India 4Cetoniinae Or.1 (4)
Torynorrhina opalina (Hope,1831)   Japan 2Cetoniinae (2) Pal.12
Toxospathius auriventris Bates,1891   China 5Melolonthinae Pal.12 (5)
Toxospathius inconstans Fairmaire,1878   China 1Melolonthinae Pal.12 (1)
Trematodes tenebrioides (Pallas,1781)   China 2Melolonthinae Pal.18 (2)
Unknown 1Melolonthinae Pal.3 (1)
Trichaphodioides bellus Endrödi,1961   Tanzania 3Aphodiinae Afr.20 (3) 3 paratypes
Trichaphodioides calcaratus Boheman,1857   Ghana 8Aphodiinae Afr.18 (8)
Somalia 2Aphodiinae Afr.26 (2) paratypes of A. pulchellus Müller
South Africa 1Aphodiinae Afr.16 (1)
Tanzania 1Aphodiinae Afr.16 (1)
Trichaphodioides cinerascens Klug,1855   Mozambique 1Aphodiinae Afr.20 (1)
Tanzania 3Aphodiinae Afr.8 (2)
Afr.30 (1)
Trichaphodioides spp.    Tanzania 3Aphodiinae Afr.8 (2)
Afr.30 (1)
Trichaphodius badius Boheman,1857   Kenya 1Aphodiinae Afr.8 (1)
Trichaphodius divisus Schmidt,1908   Namibia 51Aphodiinae (44) Afr.22+23
Afr.30 (2)
Afr.32 (4)
Afr.35 (1)
Trichaphodius flavus Endrödi,1955   Guinea-Bissau 1Aphodiinae Afr.15 (1)
Trichaphodius spp.    Ghana 1Aphodiinae Afr.30 (1)
Kenya 6Aphodiinae Afr.30 (6)
Malawi 1Aphodiinae Afr.30 (1)
Rwanda 2Aphodiinae Afr.30 (2)
Tanzania 2Aphodiinae Afr.30 (2)
Trichillum martinezi Vaz-de-mello & Louzada,   Brazil 1Scarabaeinae (1) 1 paratype S.Am.2
Trichiorhyssemus asperulus (Waterhouse,1875)   Japan 1Aphodiinae (1) Pal.18
Trichiorhyssemus congolanus (Clouet,1901)   Gambia 54Aphodiinae Afr.12 (54)
Senegal 6Aphodiinae (6) Afr.12
Sierra Leone 60Aphodiinae (60) Afr.15
Trichiorhyssemus dalmatinus Petrovitz,1967   Italy 1Aphodiinae (1) Pal.18
Trichiorhyssemus decorsei Benard,1914   Congo 2Aphodiinae (2 ) Afr.26
Trichiorhyssemus expansicollis Benard,1930   Sri Lanka 1Aphodiinae Or.1 (1)
Trichiorhyssemus hirsutus Clouet,1901   Borneo, Indonesia / Malaysia 30Aphodiinae Or.2 (30)
Trichiorhyssemus longetarsalis Ben.   Sierra Leone 2Aphodiinae (2) Afr.14
Trichiorhyssemus occidentalis (Endrödi,1976)   Gambia 1Aphodiinae (1) Afr.12
Senegal 2Aphodiinae (2) Afr.12
Trichiorhyssemus riparius (Horn,1871)   United States 1Arizona
Aphodiinae N.Am.2 (1)
Trichiorhyssemus spp.    Sierra Leone 1Aphodiinae (1) Afr.14
Trichiotinus affinis (Gory & Percheron,1833)   United States 1New York
Cetoniinae N.Am.4 (1)
United States 7Pennsylvania
Cetoniinae N.Am.1 (6)
N.Am.3 (1)
Unknown 2Cetoniinae N.Am.3 (2)
Trichiotinus assimilis (Kirby,1837)   Canada 1Quebec
Cetoniinae N.Am.1 (1)
Canada 3Ontario
Cetoniinae N.Am.1 (3)
Canada 5British Columbia
Cetoniinae N.Am.1 (5)
Trichiotinus bibens Fabricius,1775   United States 7Connecticut
Cetoniinae N.Am.4 (7)
United States 2New York
Cetoniinae N.Am.4 (2)
Unknown 13Cetoniinae N.Am.4 (13)
America borealis, United States/Canada 1Cetoniinae N.Am.4 (1)
Trichiotinus piger (Fabricius,1775)   United States 1South Carolina
Cetoniinae N.Am.4 (1)
United States 2Ney York
Cetoniinae N.Am.4 (2)
United States 1Connecticut
Cetoniinae N.Am.4 (1)
Unknown 10Cetoniinae N.Am.4 (10)
Trichiotinus piger drummondi G. & P.   Unknown 1Cetoniinae N.Am.3 (1)
Trichiotinus spp.    Brazil 1Cetoniinae S.Am.2 (1)
Canada 1British Columbia
Cetoniinae N.Am.4 (1)
United States 1Minnesota
Cetoniinae N.Am.4 (1)
Unknown 1Cetoniinae N.Am.4 (1)
Trichiotinus viridulus Fabricius,1775   Unknown 1Cetoniinae N.Am.4 (1)
Trichius fasciatus (Linnaeus,1758)   Austria 8Cetoniinae (8) Pal.21+26
Finland 6Cetoniinae (6)
France 1Cetoniinae Pal.26 (1)
Germany 4Cetoniinae (4) Pal.21+26
Italy 1Cetoniinae Pal.21 (1)
Norway 4Cetoniinae (4)
Romania 6Cetoniinae Pal.26 (5)
Pal.35 (1)
Russian Federation 7Cetoniinae Pal.26 (5)
Pal.32 (2)
Sweden 887   Sk Bl Ha Sm Öl Go Ög Vg Bo Ds Nä Sö Up Vs Vr Dr Gä Hs Me Hr Jä Ån Vb Nb Ås Ly Pi Lu To
Alan Dufberg 49 (12)
Cetoniinae Main (875)
Switzerland 6Cetoniinae Pal.26 (6)
Unknown 9Cetoniinae Pal.26 (9)
Trichius gallicus Dejean,1821   Czech Republic 1Cetoniinae Pal.26 (1) as Trichius rosaseus zonatus Germar
1794
France 2Cetoniinae Pal.26 (1) as Trichius rosaseus zonatus Germar
1794
Pal.35 (1)
Greece 1Cetoniinae Pal.26 (1) as Trichius rosaseus zonatus Germar
1794
Italy 5Cetoniinae Pal.22 (5) as Trichius rosaceus rosaceus (Voet
1769)
Russian Federation 1Cetoniinae Pal.32 (1) as Trichius zonatus
Spain 5Cetoniinae Pal.21 (5) as Trichius rosaseus zonatus Germar
1794
Trichius sexualis Bedel,1906   Russian Federation 1Sibiria
Cetoniinae Pal.26 (1)
Turkey 4Cetoniinae Pal.28 (4)
Unknown 2Cetoniinae Pal.26 (2)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Cetoniinae Pal.21 (2)
Trichius spp.    Georgia 5Cetoniinae Pal.32 (5)
Greece 1Cetoniinae Pal.35 (1)
Turkey 1Cetoniinae Pal.35 (1)
Trichocephala brincki Schein,1960   South Africa 1Type Schein (1) Holotype 00001:1 of Heteroclita brincki Schein 1960
  Heteroclita brincki Schein 1960

Trichogomphus bronchus (Jablonsky,1785)   Unknown 5Dynastinae Or.2 (5)
Trichogomphus excavatus fairmairei Arrow,1911   Solomon Islands 1Dynastinae Pac.1 (1)
Garden Island 1Dynastinae Pac.1 (1)
Unknown 1Dynastinae Pac.1 (1)
Trichogomphus lunicollis Burmeister,1847   Borneo, Indonesia / Malaysia 1Dynastinae Or.9 (1)
Trichogomphus martabani (Guerin-Meneville,1834)   India 1Dynastinae Or.2 (1)
Malaysia 1Dynastinae Or.11 (1)
Myanmar 1Dynastinae Or.12 (1)
Trichoplus cordicollis Waterhouse,1881   South Africa 3Cetoniinae (3) Afr.17
Trichostetha capensis (Linnaeus,1767)   South Africa 45Cetoniinae (45) Afr.11+12+26
Unknown 14Cetoniinae (14) Afr.26
Trichostetha capensis fuscorubra (Linnaeus,1767)   South Africa 35Cetoniinae (35) Afr.12
Trichostetha fascicularis (Linnaeus,1767)   South Africa 46Cetoniinae (46) Afr.13+14+25
Unknown 9Cetoniinae (9) Afr.25
Trichostetha signata (Fabricius,1775)   South Africa 17Cetoniinae (17) Afr.14
Unknown 4Cetoniinae (4) Afr.24
Trigonopeltastes delta (Forster,1771)   United States 2Cetoniinae N.Am.3 (2)
United States 1Missouri
Cetoniinae N.Am.1 (1)
Trigonophorus delesserti (Guerin,1839)   India 2Cetoniinae Or.2 (2)
Trigonophorus rothschildii Fairmaire,1891   Taiwan 1Cetoniinae Pal.20 (1)
Trigonostomus ursus Arrow,1911   Sri Lanka 1Rutelinae Or.5 (1)
Triodonta tenella Fåhraeus,1857   South Africa 5Melolonthinae Afr.3 (5)
Triodontella demelti (Petrovitz,1962)   Turkey 3Melolonthinae Pal.14 (3) as Triodonta d.
Triodontella dispar (Fairmaire,1892)   Turkey 2Melolonthinae Pal.15 (1) as Triodonta d.
Pal.18 (1) as Triodonta d.
Triodontella flavimana Burmeister,1855   Turkey 5Melolonthinae Pal.14 (2)
Pal.15 (2) as Triodonta f.
Pal.18 (1) as Triodonta f.
Triodontella lateristria (Reitter,1889)   Turkey 11Melolonthinae Pal.15 (11) as Triodonta l.
Turkey 2Melolonthinae Pal.15 (2) as Triodonta l.
Triodontella nitidula (Rossi,1790)   Italy 4Sericini Pal.2 (4)
Triodontella raymondi (Perris,1869)   Italy 1Sericini Pal.2 (1)
Trioplus cylindricus (Mannerheim,1829)   Brazil 3Dynastinae (3) S.Am.21
Tropinota hirta (Poda,1761)   Austria 1Cetoniinae Pal.19 (1)
Bulgaria 2Cetoniinae Pal.7 (2)
Cyprus 7Cetoniinae Pal.7 (4)
Pal.30 (1)
Pal.35 (2)
Greece 77Cetoniinae (71) Pal.4+5+6+11+17+19
Pal.30 (2)
Pal.34 (3)
Pal.35 (1)
Greece 8Crete
Cetoniinae Pal.27 (3)
Pal.32 (3)
Pal.35 (2)
Greece 1Rhodos
Cetoniinae Pal.35 (1)
Hungary 7Cetoniinae Pal.20 (5)
Pal.32 (2)
Italy 6Cetoniinae (6) Pal.10+19
Romania 4Cetoniinae Pal.7 (4)
Spain 2Cetoniinae Pal.7 (1)
Pal.19 (1)
Spain 1Mallorca
Cetoniinae Pal.27 (1)
Sweden 1 Östergötland
Cetoniinae Main (1)
Tunisia 2Cetoniinae Pal.11 (2)
Turkey 46Cetoniinae (36) Pal.7+18+19
Pal.27 (8)
Pal.35 (2)
Tropinota hirta suturalis Reitter,1913   Armenia 1Cetoniinae Pal.6 (1)
Tropinota senicula Ménétriés,1832   Armenia 1Cetoniinae Pal.30 (1)
Tropinota spp.    Spain 3Mallorca
Cetoniinae Pal.27 (3)
Tropinota squalida Scopoli,1763   Greece 12Cetoniinae Pal.5 (12)
Israel 1Cetoniinae Pal.6 (1)
Italy 12Cetoniinae (12) Pal.8+10
Portugal 7Cetoniinae (7) Pal.7+8
Spain 11Cetoniinae Pal.8 (11)
Tunisia 15Cetoniinae Pal. (4)
Pal.8 (6)
Pal.34 (5)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 2Cetoniinae Pal.8 (2)
Unknown 6Cetoniinae Pal.19 (1)
Pal.22 (5)
Tropinota squalida canariensis H.Lindberg,1950   Spain 30Canary Islands
Cetoniinae Pal.18 (21)
Pal.27 (8)
Pal.35 (1)
Tropinota squalida vittula Reiche & Saulcy,1856   Israel 3Cetoniinae Pal.30 (3)
Tunisia 1Cetoniinae Pal.30 (1)
Tropinota turanica Reitter,1889   Afghanistan 1Cetoniinae Pal.10 (1)
Trymodera aterrima Gerstaecker,1867   Malawi 1Cetoniinae Afr.5 (1)
Unidentified Aphodiinae    Australia 3Aphodiinae Indet.1 (3)
Austria 21Aphodiinae Indet.1 (21)
Botswana 1Aphodiinae Indet.3 (1)
Brazil 4Aphodiinae Indet.2 (4)
Bulgaria 1Aphodiinae Indet.1 (1)
Burundi 2Aphodiinae Indet.3 (2)
Canada 0
Congo, the democratic republic of the 1Aphodiinae Indet.3 (1)
Costa Rica 0
Denmark 5Aphodiinae Indet.1 (5)
Dominican Republic 13Aphodiinae Indet.4 (13)
Ethiopia 0
France 2Aphodiinae Indet.1 (2)
Gambia 325Aphodiinae Indet.12 (162)
Indet.21 (163)
Germany 0
Ghana 18Aphodiinae Indet.3 (18)
Greece 3Aphodiinae Indet.1 (3)
Greece 1Crete
Aphodiinae Indet.1 (1)
Greece 8Rhodos
Aphodiinae Indet.1 (8)
Guinea-Bissau 11Aphodiinae Indet.3 (11)
Honduras 0
Iceland 1Aphodiinae Indet.1 (1)
India 0
Indonesia 0Java

Israel 0
Italy 7Aphodiinae Indet.1 (7)
Japan 0
Kazakhstan 0
Kenya 11Aphodiinae Indet.20 (11)
Lesotho 8Aphodiinae Indet.10 (2)
Indet.20 (6)
Madagascar 0
Malawi 0
Malaysia 6Borneo
Aphodiinae Indet.11 (6)
Mexico 3Aphodiinae Indet.20 (3)
Namibia 114Aphodiinae Indet.10 (101)
Indet.20 (13)
Nepal 0
Norway 0
Peru 1Aphodiinae Indet.2 (1)
Portugal 0Madeira

Portugal 0Azores

Russian Federation 2Aphodiinae Indet.20 (2)
Rwanda 2Aphodiinae Indet.3 (2)
South Africa 216Aphodiinae Indet.7 (113)
Indet.14 (103)
Spain 0Canary Islands

Spain 0
Sudan 0
Swaziland 1Aphodiinae Indet.3 (1)
Sweden 696Aphodiinae Indet.2 (47)
Indet.15 (122)
Indet.16 (109)
Indet.17 (153)
Indet.18 (148)
Indet.19 (117)
Switzerland 0
Tanzania 113Aphodiinae Indet.9 (113)
Tunisia 0
Turkey 10Aphodiinae Indet.4 (10)
Uganda 2Aphodiinae Indet.3 (2)
United States 149Aphodiinae Indet. (149)
Caucasus, Ryssland / Georgien / Azerbajdzjan / Armenien / Iran / Turkiet 1Aphodiinae Indet.1 (1)
Unknown 7Aphodiinae Indet.13 (7)
Zimbabwe 11Aphodiinae Indet.3 (11)
Unidentified Cetoniinae    Algeria 15Cetoniinae Indet. (14)
Indet.3 (1)
Australia 1Cetoniinae Indet. (1)
Austria 5Cetoniinae Indet. (5)
Bolivia 1Cetoniinae Indet. (1)
Brazil 1Cetoniinae Indet.1 (1)
Bulgaria 64Cetoniinae Indet. (64)
Cameroon 4Cetoniinae Indet. (2)
Indet.3 (2)
China 6Cetoniinae Indet. (1)
Indet.3 (5)
Colombia 3Cetoniinae Indet. (2)
Indet.1 (1)
Costa Rica 2Cetoniinae S.Am.2 (2)
Croatia 6Cetoniinae Indet. (3)
Indet.5 (3)
Cyprus 16Cetoniinae Indet. (15)
Indet.5 (1)
Ecuador 2Cetoniinae Indet. (2)
Egypt 10Cetoniinae Indet. (10)
Ethiopia 46Cetoniinae Indet. (10)
Indet.5 (15)
Indet.7 (21)
France 23Cetoniinae Indet. (19)
Indet.1 (4)
France 6Corsica
Cetoniinae Indet. (6)
Gambia 31Cetoniinae Indet. (31)
Georgia 4Cetoniinae Indet.5 (4)
Germany 6Cetoniinae Indet. (4)
Indet.2 (2)
Ghana 1Cetoniinae Indet. (1)
Ghana 1Cetoniinae Indet. (1)
Greece 2Rhodos
Cetoniinae Indet.9 (2)
Greece 2Crete
Cetoniinae Indet.9 (2)
Greece 202Cetoniinae Indet. (191)
Indet.5 (5)
Indet.9 (6)
Guinea-Bissau 8Cetoniinae Indet. (8)
Honduras 1Cetoniinae Indet.5 (1)
Hungary 2Cetoniinae Indet. (2)
India 2Cetoniinae Indet. (2)
Indonesia 14Java
Cetoniinae Indet. (14)
Indonesia 14Cetoniinae Indet. (2)
Indet.6 (12)
Iran 4Cetoniinae Indet. (4)
Israel 2Cetoniinae Indet. (2)
Italy 155Cetoniinae Indet. (127)
Indet.2 (28)
Italy 1Sardinia
Cetoniinae Indet.3 (1)
Japan 6Cetoniinae Indet. (6)
Jordan 4Cetoniinae Indet. (4)
Kazakhstan 1Cetoniinae Indet.3 (1)
Kenya 16Cetoniinae Indet. (12)
Indet.1 (2)
Indet.4 (2)
Libya 2Cetoniinae Indet. (2)
Madagascar 76Cetoniinae Indet. (74)
Indet.6 (2)
Malawi 1Cetoniinae Indet. (1)
Malta 4Cetoniinae Indet. (4)
Mexico 31Cetoniinae Indet. (30)
Indet.3 (1)
Morocco 6Cetoniinae Indet. (5)
Indet.8 (1)
Mozambique 4Cetoniinae Indet. (3)
Indet.1 (1)
Namibia 1Cetoniinae Indet.1 (1)
Nepal 6Cetoniinae Indet. (4)
Indet.3 (2)
Oman 17Cetoniinae Indet. (2)
Indet.2 (15)
Peru 5Cetoniinae Indet. (5)
Portugal 11Cetoniinae Indet. (11)
Romania 3Cetoniinae Indet. (3)
Russian Federation 16Cetoniinae Indet. (16)
Senegal 3Cetoniinae Indet. (3)
Serbia 2Cetoniinae Indet. (2)
Seychelles 1Cetoniinae Indet. (1)
Sierra Leone 17Cetoniinae Indet. (17)
South Africa 23Cetoniinae Indet. (16)
Indet.1 (7)
Spain 96Cetoniinae Indet. (81)
Indet.2 (15)
Spain 22Canary Islands
Cetoniinae Indet. (22)
Spain 1Mallorca
Cetoniinae Indet.5 (1)
Swaziland 21Cetoniinae Indet. (21)
Sweden 85Cetoniinae Indet. (76)
Indet.6 (9)
Switzerland 20Cetoniinae Indet. (20)
Syria 4Cetoniinae Indet. (2)
Indet.3 (2)
Tanzania 23Cetoniinae Indet. (10)
Indet.4 (13)
Thailand 1Cetoniinae Indet.6 (1)
Tunisia 116Cetoniinae Indet. (109)
Indet.5 (7)
Turkey 62Cetoniinae Indet. (29)
Indet. (8)
Indet.3 (11)
Indet.8 (14)
Turkmenistan 1Cetoniinae Indet.3 (1)
Uganda 9Cetoniinae Indet. (1)
Indet.4 (8)
United States 1Illinois
Cetoniinae Indet. (1)
United States 1New Hampshire
Cetoniinae Indet. (1)
United States 1Connecticut
Cetoniinae Indet. (1)
United States 4New York
Cetoniinae Indet. (4)
United States 1New Jersey
Cetoniinae Indet.1 (1)
United States 2Florida
Cetoniinae Indet.1 (2)
United States 25Cetoniinae Indet. (25)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 96Cetoniinae Indet. (96)
Unknown 41Cetoniinae Indet. (36)
Indet.1 (5)
Uzbekistan 6Cetoniinae Indet. (5)
Indet.3 (1)
Viet Nam 8Cetoniinae Indet.5 (8)
Zambia 2Cetoniinae Indet. (2)
Zimbabwe 1Cetoniinae Indet. (1)
Unidentified Cremastocheilini    Ecuador 1Cetoniinae S.Am.2 (1)
Peru 4Cetoniinae S.Am.2 (4)
South Africa 1Cetoniinae Afr.17 (1)
Tanzania 2Cetoniinae Afr.20 (2)
United States 2South Dakota, Arizona
Cetoniinae N.Am.1 (1)
N.Am.3 (1)
Unidentified Dynamopodinae    Sudan 1Dynamopodinae Indet.1 (1)
Unidentified Dynastinae    Brazil 14 Dynastinae Indet.1 (14)
Burundi 2 Dynastinae Indet.3 (2)
Cameroon 1 Dynastinae Indet.8 (1)
Colombia 3 Dynastinae Indet.8 (1)
Indet.10 (2)
Dominican Republic 21 Dynastinae Indet.2 (21)
Ecuador 12    Dynastinae Indet.4 (3)
Indet.5 (5)
Indet.6 (3)
Indet.7 (1)
Gambia 12 Dynastinae Indet.9 (12)
Greece 1Crete
Dynastinae Indet.10 (1)
Greece 12 Dynastinae Indet.3 (12)
Guatemala 1 Dynastinae Indet.2 (1)
Kenya 6 Dynastinae Indet.1 (6)
Libya 3 Dynastinae Indet.9 (3)
Malaysia 1 Borneo
Dynastinae Indet.8 (1)
Malaysia 1 Dynastinae Indet.3 (1)
Papua New Guinea 14 Dynastinae Indet.8 (14)
Peru 1 Dynastinae Indet.8 (1)
Seychelles 3 Dynastinae Indet.8 (3)
Spain 2 Canary Islands
Dynastinae Indet.9 (2)
Tunisia 12 Dynastinae Indet.7 (12)
Turkey 12Dynastinae Indet.10 (12)
United States 3 Dynastinae Indet.8 (3)
Venezuela 1 Dynastinae Indet.7 (1)
Viet Nam 1 Dynastinae Indet.3 (1)
Unidentified Hopliini    Austria 7Hopliini Indet. (3)
Indet.1 (4)
Bosnia and Herzegovina 5Hopliini Indet. (5)
Brazil 1Hopliini Indet. (1)
Canada 2Hopliini Indet.1 (2)
Costa Rica 1Hopliini Indet.1 (1)
Germany 2Hopliini Indet. (2)
Greece 2Hopliini Indet.1 (2)
Italy 2Sardinia
Hopliini Indet.1 (2)
Italy 1Hopliini Indet. (1)
Japan 1Hopliini Indet. (1)
Malaysia 1Hopliini Indet. (1)
Nepal 23Hopliini Indet. (3)
Indet.1 (20)
Russian Federation 1Hopliini Indet. (1)
South Africa 4Hopliini Indet. (1)
Indet.1 (3)
Spain 6Hopliini Indet. (4)
Indet.1 (2)
Switzerland 1Hopliini Indet. (1)
Tanzania 1Hopliini Indet. (1)
United States 3Hopliini Indet.1 (3)
Unknown 1Hopliini Indet.1 (1)
Unidentified Macrodactylini    United States 1Connecticut
Melolonthinae N.Am.1 (1)
Unidentified Melolonthinae    Afghanistan 12Melolonthinae Indet. (12)
Algeria 4Melolonthinae Indet. (4)
Angola 34Melolonthinae Indet. (34)
Argentina 19Melolonthinae Indet. (19)
Australia 66Melolonthinae Indet. (66)
Azerbaijan 2Melolonthinae Indet. (2)
Botswana 8Melolonthinae Indet. (3)
Indet.3 (5)
Brazil 15Melolonthinae Indet. (9)
Indet.1 (5)
Indet.3 (1)
Bulgaria 1Melolonthinae Indet.3 (1)
Canada 5Melolonthinae Indet. (5)
China 6Melolonthinae Indet. (6)
Colombia 49Melolonthinae Indet. (49)
Costa Rica 43Melolonthinae Indet. (43)
Croatia 24Melolonthinae Indet. (18)
Indet.4 (6)
Cyprus 4Melolonthinae Indet.3 (4)
Dominican Republic 1Melolonthinae Indet.4 (1)
Ecuador 20Melolonthinae Indet. (12)
Indet.4 (8)
Ethiopia 45Melolonthinae Indet.5 (45)
France 2Melolonthinae Indet. (2)
France 9Corsica
Melolonthinae Indet. (9)
Gambia 21Melolonthinae Indet. (21)
Germany 6Melolonthinae Indet. (5)
Indet.1 (1)
Greece 14Melolonthinae Indet. (10)
Indet.1 (4)
Greece 2Rhodos
Melolonthinae Indet.1 (2)
Guatemala 26Melolonthinae Indet. (25)
Indet.4 (1)
Guinea-Bissau 90Melolonthinae Indet. (90)
Honduras 41Melolonthinae Indet. (40)
Indet.4 (1)
Hungary 1Melolonthinae Indet.3 (1)
Indonesia 2Sumatra
Melolonthinae Indet. (2)
Indonesia 8Java
Melolonthinae Indet. (7 )
Indet.4 (1)
Indonesia 3Melolonthinae Indet. (3)
Indonesia 1Bali
Melolonthinae Indet. (1)
Italy 5Melolonthinae Indet. (3)
Indet.1 (2)
Japan 4Melolonthinae Indet. (4)
Kazakhstan 3Melolonthinae Indet. (3)
Kenya 64Melolonthinae Indet. (61)
Indet.3 (3)
Madagascar 2Melolonthinae Indet.3 (2)
Malawi 41Melolonthinae Indet. (41)
Malaysia 1Melolonthinae Indet. (1)
Malaysia 3Borneo
Melolonthinae Indet.1 (3)
Mexico 12Melolonthinae Indet. (9)
Indet.2 (3)
Morocco 2Melolonthinae Indet. (2)
Mozambique 2Melolonthinae Indet. (2)
Namibia 82Melolonthinae Indet. (80)
Indet.2 (2)
Nepal 2Melolonthinae Indet. (1)
Indet.1 (1)
Panama 8Melolonthinae Indet. (8)
Peru 20Melolonthinae Indet. (20)
Russian Federation 1Melolonthinae Indet.4 (1)
Rwanda 1Melolonthinae Indet. (1)
Senegal 24Melolonthinae Indet. (24)
Serbia 1Melolonthinae Indet. (1)
Seychelles 1Melolonthinae Indet. (1)
Sierra Leone 7Melolonthinae Indet. (7)
South Africa 468Melolonthinae Indet. (464)
Indet.5 (4)
Spain 3Melolonthinae Indet. (3)
Spain 1Canary Islands
Melolonthinae Indet. (1)
Sri Lanka 1Melolonthinae Indet. (1)
Sudan 7Melolonthinae Indet. (7)
Sweden 38Melolonthinae Indet. (34)
Indet.1 (4)
Syria 1Melolonthinae Indet. (1)
Tanzania 52Melolonthinae Indet. (43)
Indet.3 (9)
Thailand 1Melolonthinae Indet.4 (1)
Tunisia 14Melolonthinae Indet. (14)
Turkey 31Melolonthinae Indet. (11)
Indet.4 (20)
Uganda 5Melolonthinae Indet.3 (5)
United States 79Melolonthinae Indet. (77)
Indet.2 (2)
United States 14New York
Melolonthinae Indet. (6)
Indet.2 (8)
United States 1Idaho
Melolonthinae Indet.2 (1)
United States 3California
Melolonthinae Indet.2 (3)
United States 1Missouri
Melolonthinae Indet.2 (1)
United States 1Kansas
Melolonthinae Indet.2 (1)
United States 2Pennsylvania
Melolonthinae Indet.2 (2)
United States 1Montana
Melolonthinae Indet.2 (1)
United States 5Washington
Melolonthinae Indet.2 (5)
United States 26Connecticut
Melolonthinae Indet. (5)
Indet.2 (20)
Indet.3 (1)
United States 3Florida
Melolonthinae Indet. (1)
Indet.2 (2)
United States 2Massachusetts
Melolonthinae Indet. (2)
United States 1Virginia
Melolonthinae Indet. (1)
United States 1West Virginia
Melolonthinae Indet. (1)
United States 5Illinois
Melolonthinae Indet. (5)
S.America  var. old. 40Melolonthinae Indet. (40)
Borneo, Indonesia / Malaysia 16Melolonthinae Indet. (16)
Rhodesia, Zimbabwe / Zambia 2Melolonthinae Indet. (2)
Weitrea 1Melolonthinae Indet. (1)
Africa div. old. 14Melolonthinae Indet. (14)
Unknown 9Melolonthinae Indet. (9)
Uzbekistan 4Melolonthinae Indet. (4)
Zambia 5Melolonthinae Indet. (5)
Unidentified Orphninae    Kenya 3Orphninae Indet.1 (3)
Sri Lanka 91Orphninae Indet.1 (91)
Tanzania 2Orphninae Afr.1 (2)
Zambia 1Orphninae Afr.1 (1)
Unidentified Psammodiini    Cyprus 1Aphodiinae Pal.26 (1)
Spain 1Mallorca
Aphodiinae Pal.26 (1)
Turkey 80Aphodiinae Pal.26 (80)
Unidentified Rutelinae    Algeria 1Rutelinae Indet. (1)
Angola 48Rutelinae Indet. (48)
Australia 19Rutelinae Indet. (17)
Indet.2 (2)
Austria 5Rutelinae Indet. (5)
Botswana 4Rutelinae Indet. (4)
Brazil 71Rutelinae Indet. (68)
Indet.1 (3)
Bulgaria 2Rutelinae Indet. (2)
Cameroon 245Rutelinae Indet. (245)
Canada 2Rutelinae Indet. (2)
China 26Rutelinae Indet. (26)
Colombia 8Rutelinae Indet. (8)
Colombia 2Rutelinae Indet. (2)
Congo 7Rutelinae Indet. (7)
Costa Rica 48Rutelinae Indet. (48)
Croatia 2Rutelinae Indet. (1)
Indet.2 (1)
Ecuador 50Rutelinae Indet. (50)
Ethiopia 13Rutelinae Indet. (13)
France 13Rutelinae Indet. (13)
France 1Corsica
Rutelinae Indet. (1)
Gabon 1Rutelinae Indet. (1)
Gambia 33Rutelinae Indet. (33)
Ghana 8Rutelinae Indet. (8)
Ghana 1Rutelinae Indet. (1)
Greece 8Rutelinae Indet. (4)
Indet.4 (4)
Greece 1Corfu
Rutelinae Indet.1 (1)
Guatemala 34Rutelinae Indet. (11)
Indet.3 (22)
Indet.4 (1)
Guinea-Bissau 57Rutelinae Indet. (57)
Honduras 27Rutelinae Indet. (27)
Hungary 1Rutelinae Indet.2 (1)
India 10Rutelinae Indet. (10)
India 5Rutelinae Indet. (4)
Indet.2 (1)
Indonesia 4Rutelinae Indet. (3)
Indet.2 (1)
Indonesia 31Java
Rutelinae Indet. (31)
Iran 2Rutelinae Indet.1 (2)
Italy 22Rutelinae Indet. (22)
Jamaica 1Rutelinae Indet. (1)
Japan 39Rutelinae Indet. (39)
Kenya 58Rutelinae Indet. (57)
Indet.4 (1)
Korea, Democratic People's Republic of 1Rutelinae Indet. (1)
Madagascar 4Rutelinae Indet. (4)
Malawi 20Rutelinae Indet. (20)
Malaysia 4Rutelinae Indet. (3)
Indet.2 (1)
Mexico 24Rutelinae Indet. (24)
Morocco 1Rutelinae Indet. (1)
Mozambique 2Rutelinae Indet. (2)
Namibia 19Rutelinae Indet. (19)
Nepal 42Rutelinae Indet. (37)
Indet.2 (5)
Oman 3Rutelinae Indet. (3)
Panama 6Rutelinae Indet. (6)
Paraguay 2Rutelinae Indet.2 (2)
Peru 34Rutelinae Indet. (33)
Indet.4 (1)
Romania 3Rutelinae Indet. (1)
Indet.4 (2)
Russian Federation 4Rutelinae Indet. (4)
Rwanda 2Rutelinae Indet. (2)
Senegal 57Rutelinae Indet. (57)
Serbia 1Rutelinae Indet. (1)
Seychelles 1Rutelinae Indet. (1)
South Africa 45Rutelinae Indet. (45)
Spain 1Rutelinae Indet. (1)
Sri Lanka 1Rutelinae Indet. (1)
Sudan 1Rutelinae Indet. (1)
Sweden 35Rutelinae Indet. (35)
Switzerland 5Rutelinae Indet. (5)
Tanzania 143Rutelinae Indet. (143)
Thailand 1Rutelinae Indet. (1)
Tunisia 7Rutelinae Indet. (7)
Turkey 41Rutelinae Indet. (14)
Indet.1 (27)
Uganda 1Rutelinae Indet.1 (1)
United States 2Rhode Island
Rutelinae Indet. (2)
United States 75Rutelinae Indet. (75)
United States 1Florida
Rutelinae Indet.2 (1)
Kashmir: Pakistan / India / China 6Rutelinae Indet. (6)
Borneo, Indonesia / Malaysia 48Rutelinae Indet. (48)
Unknown 12Rutelinae Indet. (11)
Indet.1 (1)
Venezuela 1Rutelinae Indet. (1)
Zambia 2Rutelinae Indet. (2)
Zimbabwe 1Rutelinae Indet. (1)
Unidentified Scarabaeidae    Australia 2Scarabaeidae Indet.1 (2)
France 2Scarabaeidae Indet.1 (2)
Malaysia 1Scarabaeidae Indet.1 (1)
Mozambique 1Scarabaeidae Indet.1 (1)
Unidentified Scarabaeinae    Afghanistan 0
Algeria 2 Scarabaeinae Indet.1 (2)
Brazil 2 Scarabaeinae Indet.7 (2)
Burundi 7 Scarabaeinae Indet.13 (7)
Colombia 8 Scarabaeinae Indet.8 (8)
Croatia 2 Scarabaeinae Indet.4 (2)
Dominican Republic 18 Scarabaeinae Indet.7 (18)
Ecuador 0
Ethiopia 71  Scarabaeinae Indet.5 (17)
Indet.12 (54)
France 2 Scarabaeinae Indet.4 (2)
Gambia 27   Scarabaeinae Indet.2 (9)
Indet.10 (9)
Indet.11 (9)
Greece 25 Scarabaeinae Indet.3 (25)
Greece 3 Rhodos
Scarabaeinae Indet.3 (3)
Greece 1 Crete
Scarabaeinae Indet.3 (1)
Guatemala 19 Scarabaeinae Indet.8 (19)
Guinea-Bissau 17 Scarabaeinae Indet.13 (17)
Honduras 0
India 1 Scarabaeinae Indet.14 (1)
Indonesia 7 Java
Scarabaeinae Indet.14 (7)
Indonesia 1 Bali
Scarabaeinae Indet.14 (1)
Iran 1 Scarabaeinae Indet.12 (1)
Italy 16 Scarabaeinae Indet.3 (16)
Japan 10 Scarabaeinae Indet.14 (10)
Kenya 17 Scarabaeinae Indet.6 (17)
Libya 0
Madagascar 1 Scarabaeinae Indet.1 (1)
Malawi 4 Scarabaeinae Indet.13 (4)
Malaysia 1 Scarabaeinae Indet.14 (1)
Mozambique 3 Scarabaeinae Indet.13 (3)
Namibia 21 Scarabaeinae Indet.1 (21)
Nepal 2 Scarabaeinae Indet.14 (2)
Oman 2 Scarabaeinae Indet.14 (2)
Paraguay 0
Peru 13 Scarabaeinae Indet.7 (13)
Portugal 2 Azores
Scarabaeinae Indet.3 (2)
Romania 1 Scarabaeinae Indet.4 (1)
South Africa 6 Scarabaeinae Indet.1 (6)
Spain 1 Canary Islands
Scarabaeinae Indet.3 (1)
Spain 3 Scarabaeinae Indet.3 (3)
Sri Lanka 80  Scarabaeinae Indet.15 (24)
Indet.16 (56)
Sudan 1 Scarabaeinae Indet.1 (1)
Syrian Arab Republic 1 Scarabaeinae Indet.1 (1)
Tanzania 15 Scarabaeinae Indet.4 (15)
Thailand 1 Scarabaeinae Indet.12 (1)
Tunisia 1 Scarabaeinae Indet.1 (1)
Turkey 22Scarabaeinae Indet.17 (22)
Uganda 9 Scarabaeinae Indet.13 (9)
United States 2 Hawaii
Scarabaeinae Indet.14 (2)
United States 1 Connecticut
Scarabaeinae Indet.14 (1)
United States 1 Wisconsin
Scarabaeinae Indet.14 (1)
United States 1 Florida
Scarabaeinae Indet.14 (1)
Zambia 15 Scarabaeinae Indet.9 (15)
Unidentified Sericini    Angola 110Sericini Indet. (110)
Argentina 1Sericini Indet. (1)
Australia 5Sericini Indet. (5)
Bulgaria 2Sericini Indet. (2)
Cameroon 20Sericini Indet. (20)
Canada 8Sericini Indet. (8)
Congo 3Sericini Indet. (3)
Cyprus 2Sericini Indet. (2)
Ethiopia 127Sericini Indet. (126)
Indet.1 (1)
Gambia 54Sericini Indet. (54)
Germany 0Sericini Indet. (0)
Greece 3Sericini Indet. (3)
Guinea-Bissau 29Sericini Indet. (27)
Indet.1 (2)
Indonesia 40Java
Sericini Indet. (39)
Indet.1 (1)
Japan 7Sericini Indet. (7)
Kenya 28Sericini Indet. (28)
Madagascar 2Sericini Indet. (2)
Malawi 14Sericini Indet. (14)
Malaysia 1Sericini Indet. (1)
Namibia 2Sericini Indet. (2)
Nepal 8Sericini Indet.1 (8)
Norway 0Sericini Indet. (0)
Peru 1Sericini Indet. (1)
Rwanda 1Sericini Indet. (1)
Senegal 34Sericini Indet. (34)
Sierra Leone 10Sericini Indet. (10)
South Africa 45Sericini Indet. (45)
Spain 1Sericini Indet. (1)
Sri Lanka 11Sericini Indet. (11)
Sweden 15Sericini Indet. (13)
Indet.1 (2)
Tanzania 23Sericini Indet. (23)
Tunisia 1Sericini Indet. (1)
United States 2Massachusetts
Sericini Indet. (2)
United States 33Sericini Indet. (33)
Borneo, Indonesia / Malaysia 2Sericini Indet. (2)
Zambia 5Sericini Indet. (5)
Uroxys spp.    Guatemala 2Scarabaeinae S.Am.2 (2)
Honduras 1Scarabaeinae S.Am.2 (1)
Valgus canaliculatus (Olivier,1789)   United States 5District of Columbia
Cetoniinae N.Am.1 (5)
United States 2Maryland
Cetoniinae N.Am.1 (2)
United States 2Pennsylvania
Cetoniinae N.Am.4 (2)
Unknown 2Cetoniinae N.Am.4 (2)
America borealis, United States/Canada 3Cetoniinae N.Am.4 (3)
Valgus hemipterus (Linnaeus,1758)   Austria 5Cetoniinae Pal.26 (5)
Bulgaria 3Cetoniinae Pal.23 (2)
Pal.30 (1)
Denmark 2Cetoniinae Main (2)
France 11Cetoniinae Pal.23 (6)
Pal.26 (4)
Pal.30 (1)
Germany 5Cetoniinae Pal.23 (1)
Pal.26 (3)
Pal.28 (1)
Greece 11Cetoniinae Pal.23 (11)
Greece 9Crete
Cetoniinae Pal.28 (3)
Pal.30 (4)
Pal.35 (2)
Hungary 1Cetoniinae Pal.26 (1)
Italy 29Cetoniinae Pal.23 (29)
Romania 10Cetoniinae Pal.23 (10)
Spain 2Cetoniinae Pal.23 (2)
Sweden 1 Öland
Cetoniinae Main (1)
Turkey 6Cetoniinae Pal.23 (1)
Pal.26 (5)
Jugoslavia: Croatia / Slovenia / Macedonia / Bosnia & Herzegovina 8Cetoniinae Pal.23 (8)
Unknown 2Cetoniinae Pal.22 (1)
Pal.26 (1)
Valgus spp.    Japan 8Cetoniinae Pal.23 (8)
Turkey 3Cetoniinae Pal.32 (3)
Xeloma atra (Thunberg,1818)   Namibia 2Cetoniinae (2) Afr.15
South Africa 5Cetoniinae (5) Afr.15
Xeloma maura (Boheman,1857)   Namibia 2Cetoniinae Afr.5 (2)
Xeloma pilicollis (Kraatz,1982)   Unknown 1Cetoniinae (1) Afr.14
Xeloma spp.    Central African Republic 1Cetoniinae (1) Afr.26
Xenoreoderus humilis Gestro,1891   Australia 1Cetoniinae Pac.1 (1)
Xinidium dewitzi Harold,1878   South Africa 3Scarabaeinae (3) Afr.33
Xyloryctes corniger Bates,1888   Mexico 3Dynastinae S.Am.1 (3)
Xyloryctes ensifer Bates,1888   Guatemala 1Dynastinae (1) S.Am.32
Mexico 6Dynastinae N.Am.5 (6)
Xyloryctes jamaicensis (Drury,1773)   United States 5Connecticut
Dynastinae (5) N.Am.1+5
United States 4Arizona
Dynastinae N.Am.5 (3)
N.Am.6 (1)
United States 1New Mexico
Dynastinae N.Am.5 (1)
United States 1Pennsylvania
Dynastinae N.Am.1 (1)
Unknown 4Dynastinae S.Am.1 (4)
Xyloryctes thestalus borealis Endrödi,1975   Guatemala 2Dynastinae S.Am.1 (2)
Xylotrupes australicus Thomson,1859   Australia 8Dynastinae (8) Or.6+7
Xylotrupes gideon (Linnaeus,1767)   Solomon Islands 1Dynastinae Pac.3 (1)
Thailand 1Dynastinae Or.12 (1)
Unknown 23Dynastinae (23) Or.3+4+5
Xylotrupes gideon borneensis Minck,1920   Borneo, Indonesia / Malaysia 1Dynastinae Or.6 (1)
Xylotrupes gideon gideon (Linnaeus,1767)   Indonesia 7Java
Dynastinae (7) Or.7+10
Xylotrupes lorquini Schaufuss,1885   Unknown 2Dynastinae Or.8 (2)
Xylotrupes meridionalis Prell,1914   Sri Lanka 1Dynastinae Or.3 (1)
Xylotrupes socrates socrates Schaufuss,1864   Hong Kong 1Dynastinae Or.6 (1)
India 8Dynastinae (8) Or.5+6
Xylotrupes socrates tonkinensis Minck,1920   India 5Dynastinae Or.5 (5)
Thailand 1Dynastinae Or.10 (1)
Xylotrupes spp.    Papua New Guinea 1Dynastinae Or.11 (1)
Xylotrupes toprobanes Prell,1914   Sri Lanka 1Dynastinae Or.9 (1)
Youngaphodius agullasi (Endrödi,1983)   South Africa 1Aphodiinae Afr.31 (1)